Details for Tipula (Beringotipula) unca
Name:Tipula (Beringotipula) unca Wiedemann, 1817
Publication:Zool. Mag., Kiel 1: 64 (as Tipula)
Status:Recognized taxon, Synonym(s): hortensis Meigen, 1818; longicornis Schummel, 1833.
Classification:Family Tipulidae
Subfamily Tipulinae
Keys:Peeters en Oosterbroek, 2016a (craneflies Netherlands, Belgium and Luxembourg) (in Dutch); Boardman, 2016 (craneflies Shropshire); Stubbs and Kramer, 2016d; Stubbs, 2021 (British craneflies)
Variant spelling(s):unga
Tipula (Beringotipula) unca : habitus - maleTipula (Beringotipula) unca : habitus - maleTipula (Beringotipula) unca : habitus - maleTipula (Beringotipula) unca : habitus - maleTipula (Beringotipula) unca : habitus - male
habitus
male
habitus
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plus 34 more images of habitus, habitus, habitus, habitus, habitus, habitus, habitus, habitus, body part(s), body part(s), body part(s), body part(s), body part(s), body part(s), body part(s), hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, ovipositor, ovipositor, wing, wing, wing, habitat, habitat, habitat, habitat, habitat, habitat, habitat and habitat.
Distribution:Albania, Andorra, Austria, Belarus, Belgium, Bosnia-Herzegovina, Bulgaria, Czech Rep., Denmark, Estonia, Finland, France, Germany, Great Britain, Greece, Hungary, Ireland, Italy, Latvia, Lithuania, Luxembourg, Montenegro, Netherlands, Norway, Poland, Romania, Serbia, Slovakia, Spain (north), Sweden, Switzerland, Ukraine; Russia: RUN, RUW, RUC, RUE, North Caucasus; Georgia, Armenia, Azerbaijan, Turkey (Asiatic part: Erzurum);; Russia: WS (west); Kazakhstan (east).
Note: see the manual for abbreviations: present-day Russia.
Region(s):Westpalaearctic;; Eastpalaearctic
General note:For Finland check Salmela, 2012b, 2012c, and citations below. For Norway check Hofsvang et al., 2019, and citations below. For Great Britain and Ireland check Stubbs, 2021, and citations below.
Map:Ashe et al., 2007a (Ireland); Boardman, 2007 (Shropshire)
Figure
references:
Habitus: Boumans and Aarnes, 2010 (of copula coloured blue because of Iridovirus infection); Quindroit and Lemoine, 2022a (female)

Immature stages
Larva: Podeniene, 2001a; Podeniene, 2003a; Podeniene et al., 2019 (first instar larva)
Citationson biology (mainly from 2000 onward):
Austria
Habitat. Vogtenhuber and Kofler, 2017: 124 (found in the montane zone).
Belarus
Habitat. Paramonov, 2019a: 96-97 (loc(s) Berezinsky Biosphere Reserve, distr, habitats, citing various authors) (in Russian).
Belgium
Biology. Peeters, 2023: 233 (loc(s) Botanic Garden Jean Massart, Brussels-Capital Region, rather common spring species in its habitat of marshes and wet woods).
Country not relevant
Biology. Wohltmann, 1996: 203-205, 207 (on the mite Johnstonia errans, exclusively parasitizing on adult Tipula spp.) (also as Tipula longicorns).
Habitat. Byers and Arnaud, 2001: 15 (immature forms have been found in or beneath mats of mosses on moist, organically rich soil at or near the margins of small streams).
Czech Rep.
Habitat. Stary et al., 2013a: 370 (on Tipulidae from Vraz near Pisek, with general habitat description).
Denmark
Habitat. Byriel and Rojas, 2017: 118 (among the 101 species in a study on cranefly hotspots in unmanaged and managed forests) (in Danish).
Finland
Habitat. Autio and Salmela, 2010: 49 (habitats Aland Is).
France
Habitat. Tillier and Dehalleux, 2019: 156-157 (loc(s) with habitat info, summary habitat) (in French).
Habitat. Quindroit, 2020a: 19, 45-48 (annotated list with information on abundancy and habitat of the Tipuloidea of the Pays de la Loire regoin, distr) (in French).
Habitat. Quindroit, 2020b: 127-128 (loc(s) Nord (59) and Pas-de-Calais (62) with notes on habitats and distr) (in French).
Habitat. Quindroit and Lemoine, 2021a: 26 (loc(s), resultats de deux campagnes de piegeage dans le Pas-de-Calais (62); see paper for details on habitat).
Habitat. Quindroit, 2021a: 36 (loc(s), resultats d une campagne de piegeage dans le Pas-de-Calais (62); see paper for details on habitat).
Habitat. Quindroit and Lemoine, 2022a: 16 (loc(s) Aisne (02), Oise (60) and Somme (80) with notes on habitats and distr) (in French).
Georgia
Biology. Carreno et al., 2018: 136-137 (on the thelastomatid nematode parasite Cephalobellus brevicaudatus in larvae of T. unca, citing Leibersperger, 1960).
Germany
Habitat. Schacht et al., 2001: 426 (abbreviated habitat information).
Habitat. Heiss and Merkel-Wallner, 2013: 21-24 (loc(s) Bayern, table with Malaise trap habitats and species; see paper for details).
Habitat. Heiss et al., 2017: 179 (overview of collecting methods and habitats).
Habitat. Dunk, 2018: 79, 95 (loc(s) Bayern with list of habtats).
Great Britain
Habitat. Godfrey, 2000a: 19, 21.
Habitat. Boardman, 2007: 21 (wet grassland, wet woodland or other damp habitats are suitable).
Habitat. Stubbs, 2018b: 2 (prefers meadow seepages and wet scrubb carr).
Habitat. Kramer and Morris, 2021b: 26 (info on the species, on the distr and map of loc(s) Leicestershire and Rutland (VC55); tends to be found in marshy places and damp woods [with] the larvae living in the damp humus, sometimes under moss pads).
Habitat. Devlin et al., 2022: 35 (list of Tipulidae observations across Wales, 1976-2019, in paper on the impact of climate change; observations: 320; habitat: Marshy grasslands).
Lithuania
Habitat. Podeniene, 2001a: 81-84 (larvae were found in silt and leaf fallings on the banks of small rivers and in leaf litter in Alnus swamp and Salix shrubs).
Habitat. Podeniene and Gelhaus, 2002: 70 (larvae in the riparian zone of river, between sand and leaf litter of Populus tremula).
Habitat. Podeniene, 2002: 306 (larvae were found in mud of low-laying marsh (Alnetalia glutinosae) on boggy lakeshore).
Habitat. Podeniene, 2003a: 1-295 (larva, descr, figs, habitat) (in Lithuanian).
Habitat. Podeniene, 2003b: 12, 21-24, 29 (larval habitat: sand and silt in riparian zone of small rivulets, green mosses on the banks of rivulets and land-improvement channels, mud in bogs of low lying woody swamps, fallings in swampy deciduous forests, fallings and uppermost clay soil layer in Salix groves, sandy loam under grassy vegetation or under fallings not completely covered by forest, uppermost layer of soil under mosses).
Biology. Podeniene et al., 2019: 92, 97-98, 105-109 (descr first instar larva, rearing, figs).
Biology. Gorban and Podeniene, 2022: 8 (loc(s); reared from Oak [Quercus robur], species with a typical development habitat in moist soil or leaf litter, reared from dead wood for the first time; see paper for details).
Netherlands
Biology. Peeters, 2023: 233 (rather common spring species in its habitat of marshes and wet woods).
Norway
Biology. Boumans and Aarnes, 2010: 120-122 (documentation with colourphoto of copula coloured blue because of Iridovirus infection).
Poland
Biology. Kehlmaier and Floren, 2010: 174 (three larvae of the pipunculid Nephrocerus flavicornis were extracted from the abdomen of females of T. (B.) unca collected on Carpinus betulus and Picea abies).
Romania
Habitat. Ujvarosi and Poti, 2006: 257 (habitat in Dupa Lunca marshland: shrubs along rivers and streams).
Habitat. Ujvarosi et al., 2011b: 111 (loc(s) Dupa Lanca marshy area near Voslobeni) (in Hungarian, species list with habitat information in English).
Russia
Habitat. Lantsov, 2002c: 113 (damp deciduous and mixed forests, near bodies of water) (in Russian) (information on the Teberdinsky State Nature Biosphere Reserve kindly provided by Vladimir Lantsov).
on characters and taxonomy (mainly from 2000 onward):
Country not relevant
Description. Sivell et al., 2023: 1-15 (presenting a genome assembly from an individual female; the genome sequence is 692.2 megabases in span; most of the assembly is scaffolded into four chromosomal pseudomolecules; the mitochondrial genome has also been assembled and is 16.57 kilobases in length).
Great Britain
Characters. Kramer and Morris, 2021b: 26 (diagnosis, figs).
Lithuania
Immatures. Podeniene, 2001a: 81-84 (descr larva, figs).
Immatures. Podeniene, 2003a: 1-295 (larva, descr, figs, habitat) (in Lithuanian).
Immatures. Podeniene et al., 2019: 92, 97-98, 105-109 (descr first instar larva, rearing, figs).
Poland
Figures. Kehlmaier and Floren, 2010: 174 (figs infected females).
on distribution (mainly from 2000 onward):
Andorra
Distribution. Eiroa and Baez, 2002b: 79 (checklist).
Armenia
Distribution. Hakobyan and Jenderedjian, 2023: 1519 (annotated checklist of the craneflies of Armenia with refs, distr and flight period in Caucasus Ecoregion with refs).
Austria
Distribution. Vogtenhuber, 2011: 46 (checklist Austria, list of provinces).
Distribution. Heiss et al., 2016: 70, 81 (loc(s) Karnten).
Distribution. Vogtenhuber and Kofler, 2017: 124 (loc(s) Osttirol, distr in Austria).
Azerbaijan
Distribution. Snegovaya, 2021: 1151 (checklist with details on distr in Azerbaijan).
Belarus
Distribution. Paramonov, 2019a: 96-97 (loc(s) Berezinsky Biosphere Reserve, distr, habitats, citing various authors) (in Russian).
Belgium
Distribution. Peeters, 2023: 233 (loc(s) Botanic Garden Jean Massart, Brussels-Capital Region, rather common spring species in its habitat of marshes and wet woods).
Bosnia-Herzegowina
First record. Simova-Tosic and Oosterbroek, 2003: 47 (loc(s)).
Bulgaria
Distribution. Hubenov, 2021a: 27 (listed for Bulgaria, without data, references).
Country not relevant
Distribution. Byers and Arnaud, 2001: 17 (distr).
Czech Rep.
Distribution. Stary et al., 2005d: 37 (loc(s)).
Distribution. Stary, 2009k: webpage (checklist).
Distribution. Stary et al., 2013a: 370 (on Tipulidae from Vraz near Pisek, with general habitat description).
Denmark
Distribution. Petersen et al., 2001b: 163 (checklist).
Finland
Distribution. Salmela, 2001a: 142, 149 (loc(s)).
Distribution. Salmela, 2002: 30-32 (loc(s) lake Iidesjarvi) (in Finnish).
Distribution. Salmela et al., 2007b: 46 (loc(s)).
Distribution. Autio and Salmela, 2010: 49 (loc(s) Aland Is).
Distribution. Salmela, 2010a: 12 (survey fauna Tahmela spring in city of Tampere) (in Finnish).
Distribution. Salmela, 2012b: 240 (annotated list of Finnish crane flies).
Distribution. Salmela and Petrasiunas, 2014: 33 (checklist Finnish Tipulomorpha).
France
Distribution. Labat, 2013: 74 (on a sampling campaign on the Dordogne river sources, five Upper-Dordogne tributaries and a travertine, Auvergne; see paper for details) (in French).
Distribution. Tillier and Dehalleux, 2019: 156 (new and published records Ile-de-France, loc(s) Seine-et-Marne (77), Yvelines (78), Hauts-de-Seine (92) and Val-d’Oise (95)).
Distribution. Quindroit, 2020a: 19, 45-48 (annotated list with information on abundancy and habitat of the Tipuloidea of the Pays de la Loire regoin, distr) (in French).
Distribution. Quindroit, 2020b: 127-128 (loc(s) Nord (59) and Pas-de-Calais (62) with notes on habitats and distr) (in French).
Distribution. Labat, 2021: 93 (on Diptera taxa recorded along the over-all length of the Dordogne River and some of its tributaries; see paper for the locality details) (in French).
Distribution. Quindroit and Lemoine, 2021a: 26 (loc(s), resultats de deux campagnes de piegeage dans le Pas-de-Calais (62); see paper for details on habitat).
Distribution. Quindroit, 2021a: 36 (loc(s), resultats d une campagne de piegeage dans le Pas-de-Calais (62); see paper for details on habitat).
Distribution. Quindroit and Lemoine, 2022a: 16 (loc(s) Aisne (02), Oise (60) and Somme (80) with notes on habitats and distr) (in French).
Distribution. Quindroit and Lemoine, 2022a: 34 (table specifying the presence in the five departments of the Hauts-de-France region).
Distribution. Lemoine and Quindroit, 2023: 7 (loc(s) Jura (39)).
Distribution. Tillier et al., 2023: 173 (loc(s) Doubs (25), with list of altitude(s) and habitat(s)) (in French).
Distribution. Quindroit and Racine, 2025: 356 (table specifying the presence in the five departments of the Pays de la Loire region).
Germany
Distribution. Schacht et al., 2001: 426 (loc(s) Bayern).
Distribution. Heiss, 2003: 13 (checklist Thuringen).
Distribution. Schacht, 2005: 3 (checklist Bayern).
Distribution. Hable et al., 2010b: 90 (loc(s) Bayern).
Distribution. Schacht, 2010: 38 (checklist Diptera Bayern).
Distribution. Merkel-Wallner et al., 2011: 214 (loc(s) Bayern).
Distribution. Heiss and Merkel-Wallner, 2013: 21-24 (loc(s) Bayern, table with Malaise trap habitats and species; see paper for details).
Distribution. Merkel-Wallner, 2014: 49 (loc(s) Bayern).
Distribution. Heiss et al., 2017: 164 (loc(s) Thuringen).
Distribution. Heiss, 2017b: 22 (checklist Tipulidae Thuringen).
Distribution. Dunk, 2018: 79, 95 (loc(s) Bayern with list of habtats).
Distribution. Heiss et al., 2019: 250 (loc(s); annotated checklist of the Tipulidae of Niedersachsen and Bremen, including the presence in the surrounding Bundeslandern and the Netherlands) (in German).
Distribution. Heiss, 2019: 36, 44 (loc(s) and checklist Mecklenburg-Vorpommern, see paper for further details).
Distribution. Stuke, 2019: 241 (checklist Niedersachsen and Bremen with reference to original sources).
Distribution. Stuke et al., 2020: 291 (loc(s) Niedersachsen).
Distribution. Heiss et al., 2021: 239 (found in caves in Thuringen, Hessen and Rheinland-Pfalz, see paper on details of which part of the caves) (in German).
Distribution. Heiss and Malec, 2023: 316 (loc(s) Hessen).
Distribution. Heiss and Malec, 2023: 329 (annotated checklist Hessen).
Distribution. Kehlmaier et al., 2023: 219 (loc(s) Saarland).
Great Britain
Distribution. Godfrey, 2000a: 19, 21 (see paper for details of sites surveyed in Shropshire and into Wales).
Distribution. Hewitt et al., 2005: 37 (loc(s) Cumbria).
Distribution. Boardman, 2007: 21 (loc(s), map).
Distribution. Skidmore, 2009: 124 (review of occurrence on the Western Isles of Scotland, distr).
Distribution. Kramer, 2011g: 2 (loc(s) Devon).
Distribution. Kramer, 2011j: 9 (loc(s) Leicestershire and Rutland (VC55)).
Distribution. Hewitt, 2014: (provisional checklist Cumbrian Diptera).
Distribution. Kramer, 2014c: 4 (in table of cranefly species recorded at Shapwick Heath NNR, as given in H. Audcent, 1949 and/or collected by J. Kramer in 2013).
Distribution. Wolton et al., 2014: 26 (among Diptera associated with a hedge in Devon).
Distribution. Drake, 2016: 22 (study on the relative importance to Diptera of pasture and ditch margins on Catcott Reserve, Somerset).
Distribution. Brighton, 2017c: 30 (overall regional checklist Lancashire and Cheshire (VC58, 59 and 60) with notes on habitats and/or regional occurrence and/or status; see paper for details).
Distribution. Stubbs, 2018b: 2 (loc(s) Staffordshire).
Distribution. Kramer and Morris, 2021b: 26 (info on the species, on the distr and map of loc(s) Leicestershire and Rutland (VC55); tends to be found in marshy places and damp woods [with] the larvae living in the damp humus, sometimes under moss pads).
Distribution. Sivell et al., 2023: 4 (loc(s) Oxfordshire).
Ireland
Distribution. Ashe et al., 2007a: 301, 317-319, 342, 352 (review Irish records, map).
Distribution. Chandler et al., 2008: 5 (checklist).
Italy
Distribution. Oosterbroek, 2008c: 340 (annotated checklist South Tyrol [Prov. Bolzano]).
Distribution. Terzani et al., 2012: 306 (known from Riserva naturale Camaldoli (Arezzo), citing Oosterbroek, 2005).
Kazakhstan
First record. Devyatkov, 2008: 140 (loc(s) East Kazakhstan, distr) (in Russian).
Distribution. Devyatkov, 2022b: 26 (loc(s) East Kazakhstan, distr) (in Russian).
Lithuania
Distribution. Podeniene, 2001a: 81-84 (loc(s)).
Distribution. Podeniene and Gelhaus, 2002: 70 (loc(s)).
Distribution. Podeniene, 2002: 306 (loc(s)).
Distribution. Pakalniskis et al., 2006: 27 (checklist).
Distribution. Gorban and Podeniene, 2022: 8 (loc(s); reared from Oak [Quercus robur], species with a typical development habitat in moist soil or leaf litter, reared from dead wood for the first time; see paper for details).
Montenegro
Distribution. Simova-Tosic and Oosterbroek, 2003: 47 (review literature).
Netherlands
Distribution. Jong and Oosterbroek, 2002d: 41 (checklist).
Distribution. Oosterbroek, 2014: 258 (loc(s) Friesland).
Distribution. Oosterbroek and Jong, 2020: 46 (Tipulidae Kaaistoep, Noord-Brabant).
Norway
Distribution. Boumans and Aarnes, 2010: 120 (loc(s) Finmark).
Distribution. Hofsvang et al., 2019: 106-107, 142-143 (review Norwegian records, annotated checklist Nordic countries and for Norway according to the Strand regions).
Poland
Distribution. Palaczyk, 2001: 270 (loc(s)).
Distribution. Skibinska and Chudzicke, 2007b: 77 (checklist).
Distribution. Kehlmaier and Floren, 2010: 174 (loc(s)).
Romania
Distribution. Ujvarosi and Poti, 2006: 257 (loc(s) eastern Carpathians).
Distribution. Ujvarosi, 2007: 232-233 (checklist Tipulidae).
Distribution. Ujvarosi et al., 2011b: 111 (loc(s) Dupa Lanca marshy area near Voslobeni) (in Hungarian, species list with habitat information in English).
Distribution. Torok et al., 2014: 18 (checklist Romania).
Russia
Distribution. Humala and Polevoi, 1999: 111 (RUN: list of Diptera of the Karelian shore and islands of the White Sea) (in Russian).
Distribution. Lantsov, 2002c: 113 (on the faunal structure, biotope(s) and chorology of crane-flies of the Teberdinsky State Nature Biosphere Reserve (North Caucasus: Karachay-Cherkess Rep.) (in Russian) [first recorded for North Caucasus by Savchenko, 1961a: 478].
Distribution. Humala and Polevoi, 2008: 132 (loc(s) RUN: Kareliya).
Distribution. Humala and Polevoi, 2009: 66 (loc(s) RUN: Kareliya) (in Russian).
Distribution. Pilipenko, 2009a: 210 (annotated checklist Tipulidae Central European Territory, loc(s) RUC, distr).
Distribution. Paramonov, 2011: 41-43 (loc(s)) RUE: Mariy El Rep.).
Distribution. Paramonov and Klepikov, 2014: 37 (loc(s) Yaroslavskaya oblast).
Distribution. Paramonov, 2014: 39-43 (revision coll. Eversmann, loc(s) RUE: Orenburgskaya oblast) (in Russian).
Distribution. Paramonov and Pilipenko, 2016: 101 (loc(s) RUC: Tverskaya oblast) (in Russian).
Distribution. Paramonov, 2018a: 185 (loc(s) RUC: Chuvash Rep., distr) (in Russian).
Distribution. Paramonov and Korobkov, 2019: 100 (loc(s) RUC: Tverskaya oblast, distr, citing various authors) (in Russian).
Serbia
Distribution. Simova-Tosic and Oosterbroek, 2003: 47 (review literature).
Slovakia
Distribution. Stary, 2009g: 43 (loc(s) Polana area).
Distribution. Stary, 2009k: webpage (checklist).
Spain
Distribution. Eiroa and Baez, 2002b: 79 (checklist).
Sweden
Distribution. Salmela, 2010b: 134 (loc(s) Malmo).
Distribution. Fritz and Lindstrom, 2013: 63 (loc(s) Hallands County) (in Schwedish).
Switzerland
Distribution. Dufour and Merz, 2012: 356 (annotated checklist canton Geneva).
Turkey
First record. Oosterbroek, 2009c: 182 (loc(s) Erzurum).
on flight period (mainly from 2000 onward):
Austria
Heiss et al., 2016: month(s): 6-7.
Vogtenhuber and Kofler, 2017: month(s): 6-7.
Belarus
Paramonov, 2019a: month(s): 6.
Belgium
Peeters, 2023: month(s): 5-7.
Czech Rep.
Stary et al., 2005d: month(s): 5-7.
Stary et al., 2013a: month(s): 6.
Finland
Salmela, 2001a: month(s): 6-7.
France
Tillier and Dehalleux, 2019: month(s): 5-7.
Quindroit, 2020a: month(s): 6.
Quindroit, 2020b: month(s): 4-7.
Quindroit and Lemoine, 2022a: month(s): 5-7.
Lemoine and Quindroit, 2023: month(s): 6.
Tillier et al., 2023: month(s): 6.
Germany
Schacht et al., 2001: month(s): 5-8.
Heiss and Merkel-Wallner, 2013: month(s): 6-7.
Heiss et al., 2017: month(s): 5-7.
Dunk, 2018: month(s): 6, 8.
Heiss et al., 2019: month(s): 6.
Heiss, 2019: month(s): 5-7.
Heiss et al., 2021: month(s): 2, 6-7.
Heiss and Malec, 2023: month(s): 6.
Kehlmaier et al., 2023: month(s): 6.
Great Britain
Godfrey, 2000a: month(s): 6.
Boardman, 2007: month(s): 5-7.
Kramer, 2011j: month(s): 5-7.
Kramer, 2014c: month(s): 6.
Stubbs, 2018b: month(s): 6.
Kramer and Morris, 2021b: month(s): 5-7.
Ireland
Ashe et al., 2007a: month(s): 6-7, 9-10.
Kazakhstan
Devyatkov, 2008: month(s): 7.
Devyatkov, 2022b: month(s): 6-7.
Lithuania
Podeniene, 2001a: month(s): 5-6.
Norway
Boumans and Aarnes, 2010: month(s): 7.
Hofsvang et al., 2019: month(s): 6-9.
Poland
Kehlmaier and Floren, 2010: month(s): 6-7.
Romania
Ujvarosi et al., 2011b: month(s): 5-6.
Russia
Lantsov, 2002c: month(s): 5-7.
Pilipenko, 2009a: month(s): 6-7.
Paramonov, 2011: month(s): 6.
Paramonov, 2014: month(s): 6.
Paramonov, 2018a: month(s): 7.
Paramonov and Korobkov, 2019: month(s): 6.
Slovakia
Stary, 2009g: month(s): 7.
Turkey
Oosterbroek, 2009c: month(s): 6.
on altitude (mainly from 2000 onward):
Austria
Vogtenhuber and Kofler, 2017: altitude: 1410-1550 m.
France
Labat, 2021: altitude: 1101-1358 m.
Germany
Merkel-Wallner et al., 2011: altitude: <700 m.
Heiss et al., 2021: altitude: 250-312 m.
Kazakhstan
Devyatkov, 2022b: altitude: 300-465 m.
Romania
Ujvarosi and Poti, 2006: altitude: 670 m.
Russia
Lantsov, 2002c: altitude: 1200-1400 m.
Turkey
Oosterbroek, 2009c: altitude: 2100-2850 m.

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