Details for Tipula (Vestiplex) nubeculosa

Name: Tipula (Vestiplex) nubeculosa Meigen, 1804 Publication: Klass. Beschr. 1: 70 (as Tipula) Status: Recognized taxon, Synonym(s): rubripes Schummel, 1833; guttulifera Zetterstedt, 1838; pseudoscripta Pierre, 1921. Classification: Family Tipulidae Subfamily Tipulinae Revision: Starkevich et al., 2020b. Keys: Peeters en Oosterbroek, 2016a (craneflies Netherlands, Belgium and Luxembourg) (in Dutch); Stubbs and Kramer, 2016d; Stubbs, 2021 (British craneflies); Starkevich et al., 2020b (Vestiplex Mongolia) habitus male habitus male habitus male habitus female habitus female plus 32 more images of habitus, habitus, body part(s), body part(s), body part(s), body part(s), body part(s), body part(s), body part(s), body part(s), hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, ovipositor, ovipositor, ovipositor, ovipositor, ovipositor, ovipositor, ovipositor, wing, wing, habitat, habitat, habitat and habitat. Distribution: Austria, Belarus, Belgium, Bosnia-Herzegovina, Bulgaria, Croatia, Czech Rep., Denmark, Estonia, Finland, France, Germany, Great Britain, Greece (north), Hungary, Ireland, Italy (north, Toscana), Latvia, Lithuania, Luxembourg, Montenegro, Netherlands, North Macedonia, Norway, Poland, Romania, Serbia, Slovakia, Slovenia, Sweden, Switzerland, Ukraine; Russia: RUN, RUW, RUC, RUE;; Russia: ES (Buryatia), Mongolia. Note: see the manual for abbreviations: present-day Russia. Region(s): Westpalaearctic;; Eastpalaearctic General note: Added: Latvia (Spuris, 1995). For Finland check Salmela, 2012b, 2012c, and citations below. For Norway check Hofsvang et al., 2019, and citations below. For Great Britain and Ireland check Stubbs, 2021, and citations below. Map: Ashe et al., 2007a (Ireland); Starkevich et al., 2020b (Mongolia) Figure references: Hypopygium: Starkevich et al., 2020b Ovipositor: Starkevich et al., 2020b Immature stages Larva: Podeniene, 2003a Miscellaneous Heiss and Malec, 2023 (habitat) Citations on biology (mainly from 2000 onward): Belarus Habitat. Paramonov, 2019a: 97 (loc(s) Berezinsky Biosphere Reserve, distr, habitats, citing various authors) (in Russian). Bulgaria Habitat. Hubenov, 2025b: suppl: 2 (annotated list Diptera high Bulgarian mts, local distr, habitats and zoogeography). Country not relevant Habitat. Vogtenhuber and Kofler, 2017: 136 (a species of forest from the woodland to the upper montane zone). Biology. Gudin et al., 2022: 399 (paper includes an updated catalogue of dipteran hosts of Tachinidae with list of the tachinid species involved; of the host records from Diptera, most are from larvae of Tabanidae and Tipulidae, with a few occasional records in other families). Denmark Habitat. Byriel and Rojas, 2017: 118 (among the 101 species in a study on cranefly hotspots in unmanaged and managed forests) (in Danish). Finland Habitat. Salmela and Autio, 2007a: 58 (in the North boreal ecoregion in a strip of old-growth forest dominated by Norway Spruce [Picea abies] surrounding a forest brook and with the ground layer vegetation characterized by tall herbs). Habitat. Salmela, 2008: 12, 24, 33, 53 (49) (ecology or habitat: eurytopic, terrestrial). Habitat. Salmela, 2011a: 1-28 (study on three wetland habitats: aapa mires, springs and headwater streams; found in headwater streams only). France Habitat. Dufour, 2003b: 91 (loc(s) with information on habitat(s)) (in French). Habitat. Tillier and Oosterbroek, 2019: 238 (loc(s) and distr Mercantour; habitat: prairies et foret de Melezes [Larix] et feuillus). Germany Habitat. Schacht et al., 2001: 429 (abbreviated habitat information). Habitat. Heiss and Merkel-Wallner, 2013: 21-24 (loc(s) Bayern, table with Malaise trap habitats and species; see paper for details). Habitat. Heiss et al., 2017: 179-182 (overview of collecting methods and habitats). Habitat. Heiss, 2017a: 563, 568, 571 (loc(s) Dinkelberg, Baden-Wurttemberg and table with details on presence in the various biotop-groups). Habitat. Dunk, 2018: 79, 96 (loc(s) Bayern with list of habtats). Great Britain Habitat. Brighton, 2017c: 31, 45 (overall regional checklist Lancashire and Cheshire (VC58, 59 and 60) with notes on habitats and/or regional occurrence and/or status; see paper for details). Biology. Tschorsnig, 2017: 58 (host of Palaearctic Tachinidae, for species of Tachinidae, localities and references see paper). Lithuania Habitat. Podeniene, 2000: 6 (larvae were found in sandy soil in Pinus forest). Habitat. Podeniene, 2003a: 1-295 (larva, descr, figs, habitat) (in Lithuanian). Habitat. Podeniene, 2003b: 12, 23, 28 (larval habitat: fallings in mixed and deciduous forests, uppermost layer of soil (of western taiga type forest, of old deciduous forest rich in epiphytes)). Luxembourg Biology. Vogtenhuber, 2007: 353 (habitat: Larven entwickeln sich in den oberen Bodenschichten von Nadelwaldern insbesondere unter Fichten [Picea]; flight: Weibchen sind offenbar wesentlich weniger flugaktif als die Mannchen). Netherlands Habitat. Oosterbroek and Jong, 2001: 110 (a species of forests, either coniferous or broad-leaved, especially Fagus; larvae develop in the soil, under leaf-litter, under mosses, under the bark of trees, and have also been bred from wood-mould of Fagus, citing various authors). Norway Habitat. Olsen and Andersen, 2022: 40 (loc(s) Innlandet, in former Hedmark, with habitats specified separately, for habitats see also Jonassen and Andersen, 2020). Romania Habitat. Kolcsar et al., 2013: 74 (loc(s) Cluj-Napoca and surroundings; list of habitats with altitudes). on characters and taxonomy (mainly from 2000 onward): Country not relevant Phylogeny. Ahonen, 2007: 1-28 (included in phylogenetic analysis of Tipuloidea subfamilies relationships based on molecular data). Phylogeny. Starkevich, 2012a: 1-219 (included in a taxonomic and phylogentic review of the subgenus Vestiplex based on morphological data, key, figs) (in Lithuanian, see Starkevich, 2012b for detailed English summary). Characters. Starkevich et al., 2021: 44-45 (comparison larvae). Lithuania Immatures. Podeniene, 2003a: 1-295 (larva, descr, figs, habitat) (in Lithuanian). Mongolia Description. Starkevich et al., 2020b: 6-23, 59-62 (descr, key, figs). on distribution (mainly from 2000 onward): Austria Distribution. Vogtenhuber, 2011: 53 (checklist Austria, list of provinces). Distribution. Reusch and Heiss, 2012: 343-351 (loc(s) Nat. Park Gesause, see PDF for months and altitudes). Distribution. Heiss et al., 2016: 78, 81 (loc(s) Karnten and Niederosterreich). Distribution. Vogtenhuber and Kofler, 2017: 136 (loc(s) Osttirol, distr in Austria). Belarus Distribution. Palaczyk, 2001: 270 (loc(s)). Distribution. Paramonov, 2019a: 97 (loc(s) Berezinsky Biosphere Reserve, distr, habitats, citing various authors) (in Russian). Bosnia-Herzegowina Distribution. Simova-Tosic and Oosterbroek, 2003: 56 (review literature). Bulgaria Distribution. Hubenov, 2016: 47 (annotated checklist Diptera of the Rila Mountains based on data from the literature). Distribution. Hubenov, 2017: 67 (on vertical distribution in the Pirin and Rila Mts; see paper for details on forests and subalpine-alpine zones, distr). Distribution. Hubenov, 2018b: 123 (review loc(s) Vrachanska Planina Mts). Distribution. Hubenov, 2021a: 29 (checklist with summary of distr in Bulgaria, references). Distribution. Hubenov, 2025b: suppl: 2 (annotated list Diptera high Bulgarian mts, local distr, habitats and zoogeography). Croatia Distribution. Simova-Tosic and Oosterbroek, 2003: 56 (review literature). Czech Rep. Distribution. Stary et al., 2005d: 37 (loc(s)). Distribution. Stary, 2009k: webpage (checklist). Denmark Distribution. Petersen et al., 2001b: 163 (checklist). Finland Distribution. Polevoi, 2001: 191 (loc(s)) (as rubripes). Distribution. Salmela, 2001a: 142, 149 (loc(s)). Distribution. Stary and Salmela, 2004: 47 (loc(s)). Distribution. Mukkala et al., 2005: 16 (among list of insects collected in South Ostrobothnia, Ilmajoki during 2001-2004). Distribution. Salmela and Autio, 2007a: 58 (loc(s)). Distribution. Salmela et al., 2007b: 46 (loc(s)). Distribution. Salmela, 2008: 12, 53 (49) (loc(s), known from all Finnish ecoregions). Distribution. Salmela, 2011a: 24 (loc(s) subalpine northern boreal Finland). Distribution. Salmela, 2012b: 241 (annotated list of Finnish crane flies). Distribution. Salmela and Petrasiunas, 2014: 35 (checklist Finnish Tipulomorpha). Distribution. Salmela et al., 2015: 108 (list of localities in extensive report on malaise-trapping of insects in conservation areas in Lapland in 2012–2014) (in Finnish). France Distribution. Dufour, 2003b: 91 (loc(s) with information on habitat(s)) (in French). Distribution. Tillier and Dehalleux, 2019: 171 (review records Ile-de-France, loc(s) Seine-et-Marne (77), Yvelines (78) and Essonne (91)). Distribution. Tillier and Oosterbroek, 2019: 238 (loc(s) and distr Mercantour; habitat: prairies et foret de Melezes [Larix] et feuillus). Distribution. Tissot et al., 2021: 217 (the Diptera of the Reserves Naturelles Nationales du Doubs (25), results of Malaise trapping since 2009) (in French). Distribution. Tillier et al., 2023: 178 (loc(s) Doubs (25), with list of altitude(s) and habitat(s)) (in French). Germany Distribution. Schacht et al., 2001: 429 (loc(s) Bayern). Distribution. Heiss, 2003: 14 (checklist Thuringen). Distribution. Schacht, 2005: 4 (checklist Bayern). Distribution. Barndt, 2008: 82 (loc(s) Brandenburg). Distribution. Hable et al., 2010b: 89 (loc(s) Bayern). Distribution. Schacht, 2010: 38 (checklist Diptera Bayern). Distribution. Merkel-Wallner et al., 2011: 214 (loc(s) Bayern). Distribution. Heiss and Merkel-Wallner, 2013: 21-24 (loc(s) Bayern, table with Malaise trap habitats and species; see paper for details). Distribution. Dunk and Kraus, 2014: 168 (loc(s) Bayern). Distribution. Merkel-Wallner, 2014: 49 (loc(s) Bayern). Distribution. Wolz, 2016: 3-4 (one of the 13 Tipulidae species of which remains were found in the faeces of the Greater Horshoe Bat (Rhinolophus ferrumequinum) at Hohenburg, Bayern; final report in Wolz, 2018). Distribution. Heiss et al., 2017: 171 (loc(s) Thuringen). Distribution. Heiss, 2017a: 563, 568, 571 (loc(s) Dinkelberg, Baden-Wurttemberg and table with details on presence in the various biotop-groups). Distribution. Heiss, 2017b: 23 (checklist Tipulidae Thuringen). Distribution. Dunk, 2018: 79, 96 (loc(s) Bayern with list of habtats). Distribution. Heiss et al., 2019: 253 (loc(s); annotated checklist of the Tipulidae of Niedersachsen and Bremen, including the presence in the surrounding Bundeslandern and the Netherlands) (in German). Distribution. Heiss, 2019: 37, 44 (loc(s) and checklist Mecklenburg-Vorpommern, see paper for further details). Distribution. Stuke, 2019: 240 (checklist Niedersachsen and Bremen with reference to original sources). Distribution. Heiss et al., 2021: 242 (found in caves in Hessen, Rheinland-Pfalz and Saarland, see paper on details of which part of the caves) (in German). Distribution. Heiss and Malec, 2023: 320 (loc(s) Hessen). Distribution. Heiss and Malec, 2023: 330 (annotated checklist Hessen). Great Britain Distribution. Kramer, 2008d: 1 (historical record Scotland). Distribution. Skidmore, 2009: 125 (review of occurrence on the Western Isles of Scotland, distr). Distribution. Hewitt, 2014: (provisional checklist Cumbrian Diptera). Distribution. Brighton, 2017c: 31, 45 (overall regional checklist Lancashire and Cheshire (VC58, 59 and 60) with notes on habitats and/or regional occurrence and/or status; see paper for details). Distribution. Tschorsnig, 2017: 58 (host of Palaearctic Tachinidae, for species of Tachinidae, localities and references see paper). Ireland Distribution. Ashe et al., 2007a: 302, 336-337, 343, 350, 355 (review Irish records, red data book status, map). Distribution. Chandler et al., 2008: 7 (checklist). Italy Distribution. Oosterbroek, 2008c: 339, 342 (loc(s), annotated checklist South Tyrol [Prov. Bolzano]). Lithuania Distribution. Podeniene, 2000: 6 (loc(s)). Distribution. Pakalniskis et al., 2006: 29 (checklist). Distribution. Starkevich, 2012a: 213 (loc(s)). Luxembourg Distribution. Vogtenhuber, 2007: 351, 353 (loc(s), distr). Mongolia Distribution. Starkevich, 2012a: 214 (loc(s) Tov aimag). Distribution. Starkevich et al., 2020b: 59, 68 (loc(s), distr, map). Montenegro Distribution. Simova-Tosic and Oosterbroek, 2003: 56 (review literature). Netherlands First record. Theowald, 1952b: 47 (XXX) (as rubripes). Distribution. Oosterbroek and Jong, 2001: 110 (review Dutch records). Distribution. Jong and Oosterbroek, 2002d: 42 (checklist). North Macedonia Distribution. Simova-Tosic and Oosterbroek, 2003: 56 (review literature). Norway Distribution. Hofsvang, 2010: 13 (loc(s)). Distribution. Hofsvang et al., 2019: 132-134, 144-145 (review Norwegian records, annotated checklist Nordic countries and for Norway according to the Strand regions). Distribution. Olsen and Andersen, 2022: 40 (loc(s) Innlandet, in former Hedmark, with habitats specified separately, for habitats see also Jonassen and Andersen, 2020). Poland Distribution. Skibinska and Chudzicke, 2007b: 78 (checklist). Romania Distribution. Ujvarosi, 2007: 232-233 (checklist Tipulidae). Distribution. Keresztes, 2012: 31 (loc(s) Rimetea area [Alba County]). Distribution. Kolcsar et al., 2013: 74 (loc(s) Cluj-Napoca and surroundings; list of habitats with altitudes). Distribution. Torok et al., 2014: 19 (checklist Romania). Russia Distribution. Paramonov, 2006a: 149 (loc(s) RUC: Ulyanovskaya oblast, distr). Distribution. Humala and Polevoi, 2009: 66 (loc(s) RUN: Kareliya) (in Russian). Distribution. Pilipenko, 2009a: 217 (annotated checklist Tipulidae Central European Territory, loc(s) RUC: Tverskaya oblast, Moskovskaya oblast, distr). Distribution. Jakovlev et al., 2014: 310 (loc(s) RUN: Kareliya). Distribution. Ruchin and Pilipenko, 2015: 59 (loc(s), checklist RUC: Mordoviya Rep.). Distribution. Paramonov and Pilipenko, 2016: 101 (loc(s) RUC: Tverskaya oblast) (in Russian). Distribution. Paramonov, 2017a: 173 (loc(s) RUC: Chuvash Rep.) (in Russian). Distribution. Paramonov and Korobkov, 2019: 102 (loc(s) RUC: Tverskaya oblast, distr, citing various authors) (in Russian). Distribution. Pilipenko et al., 2020: 366 (loc(s) RUC: Mordoviya Rep., distr). Distribution. Starkevich et al., 2020b: 59 (loc(s) Buryatia, first record for Eastpaearctic Russia). Distribution. Kolcsar et al., 2023a: 23 (loc(s) RUE: Tatarstan Rep.). Serbia Distribution. Simova-Tosic and Oosterbroek, 2003: 56 (review literature). Slovakia Distribution. Stary, 2009g: 45 (loc(s) Polana area). Distribution. Stary, 2009k: webpage (checklist). Slovenia Distribution. Simova-Tosic and Oosterbroek, 2003: 56 (review literature). Switzerland Distribution. Starkevich, 2012a: 213-214 (loc(s). on flight period (mainly from 2000 onward): Austria Heiss et al., 2016: month(s): 6-7. Vogtenhuber and Kofler, 2017: month(s): 6-7. Belarus Paramonov, 2019a: month(s): 5-6. Bulgaria Hubenov, 2016: month(s): 6-7. Czech Rep. Stary et al., 2005d: month(s): 4-6. Finland Salmela, 2001a: month(s): 5, 7. Salmela and Autio, 2007a: month(s): 7 or 8. France Dufour, 2003b: month(s): 7. Tillier and Oosterbroek, 2019: month(s): 6. Tillier et al., 2023: month(s): 6. Germany Schacht et al., 2001: month(s): 5-9. Heiss and Merkel-Wallner, 2013: month(s): 5-6. Heiss et al., 2017: month(s): 5-7. Heiss, 2017a: month(s): 5. Dunk, 2018: month(s): 5-6, 9. Heiss, 2019: month(s): 6. Heiss et al., 2021: month(s): 5-7. Heiss and Malec, 2023: month(s): 5-6. Ireland Ashe et al., 2007a: month(s): 5, 7-8. Italy Oosterbroek, 2008c: month(s): 5-7. Lithuania Starkevich, 2012a: month(s): 5-6. Luxembourg Vogtenhuber, 2007: month(s): 5-6. Mongolia Starkevich, 2012a: month(s): 7. Starkevich et al., 2020b: month(s): 7. Netherlands Oosterbroek and Jong, 2001: month(s): 5-9. Norway Hofsvang et al., 2019: month(s): 5-8. Olsen and Andersen, 2022: month(s): (6)7(-9). Romania Kolcsar et al., 2013: month(s): 4, 6. Russia Paramonov, 2006a: month(s): 5-6. Pilipenko, 2009a: month(s): 5-6. Ruchin and Pilipenko, 2015: month(s): 5-6. Paramonov, 2017a: month(s): 5-6. Paramonov and Korobkov, 2019: month(s): 5. Pilipenko et al., 2020: month(s): 5. Starkevich et al., 2020b: month(s): 7. Kolcsar et al., 2023a: month(s): 6. Slovakia Stary, 2009g: month(s): 5-6. Switzerland Starkevich, 2012a: month(s): 6. on altitude (mainly from 2000 onward): Austria Vogtenhuber and Kofler, 2017: altitude: 1330-1700 m. Bulgaria Hubenov, 2016: altitude: 1350 m. Hubenov, 2017: altitude: 1350 m. Hubenov, 2018b: altitude: 457 m. Hubenov, 2021a: altitude: 450-1350 m. Hubenov, 2025b: altitude: 450-1350 m. France Dufour, 2003b: altitude: 1770 m. Tillier and Oosterbroek, 2019: altitude: 1436 m. Germany Merkel-Wallner et al., 2011: altitude: <700-1099 m. Heiss et al., 2021: altitude: 226-450 m. Italy Oosterbroek, 2008c: altitude: 1630 m. Mongolia Starkevich, 2012a: altitude: 1608 m. Starkevich et al., 2020b: altitude: 1500-2400 m. Russia Kolcsar et al., 2023a: altitude: 75-100 m. Slovakia Stary, 2009g: altitude: 630-780 m. Switzerland Starkevich, 2012a: altitude: 540 m.