Details for Tipula (Lunatipula) fascipennis

Name: Tipula (Lunatipula) fascipennis Meigen, 1818 Publication: Syst. Beschr. 1: 187 (as Tipula) Status: Recognized taxon. Classification: Family Tipulidae Subfamily Tipulinae Keys: Podeniene, 2000 (Lunatipula larva Lithuania); Peeters en Oosterbroek, 2016b (craneflies Netherlands, Belgium and Luxembourg) (in Dutch); Boardman, 2016 (craneflies Shropshire); Stubbs and Kramer, 2016d; Stubbs, 2021 (British craneflies) Variant spelling(s): fasciapennis habitus male habitus male habitus male habitus male habitus male plus 27 more images of habitus, habitus, habitus, body part(s), hypopygium, hypopygium, hypopygium, hypopygium, ovipositor, ovipositor, ovipositor, ovipositor, ovipositor, wing, wing, habitat, habitat, habitat, habitat, habitat, habitat, habitat, habitat, habitat, habitat, habitat and habitat. Distribution: Albania, Austria, Belarus, Belgium, Bosnia-Herzegovina, Bulgaria, Croatia, Czech Rep., Denmark, Estonia, Finland, France, Germany, Great Britain, Greece (north), Hungary, Ireland, Italy (north, Giglio), Latvia, Lithuania, Luxembourg, Montenegro, Netherlands, North Macedonia, Norway, Poland, Romania, Serbia, Slovakia, Slovenia, Spain, Sweden, Switzerland, Ukraine; Russia: RUN, RUW, RUC, RUE, RUS, North Caucasus; Georgia. Note: see the manual for abbreviations: present-day Russia. Region(s): Westpalaearctic General note: Added: Italy (north) (Oosterbroek and Stary, 1995). For Finland check Salmela, 2012b, 2012c, and citations below. For Norway check Hofsvang et al., 2019, and citations below. For Spain check Eiroa and Carles-Tolra, 2019, and citations below. For Great Britain and Ireland check Stubbs, 2021, and citations below. Removed, as found in earlier CCW versions: Armenia (excluded for Armenia by Hakobyan and Jenderedjian, 2023). Map: Ashe et al., 2007a (Ireland); Boardman, 2007 (Shropshire) Figure references: Habitus: Quindroit, 2020a (male); Kramer and Morris, 2021b (female) Hypopygium: Kramer and Morris, 2021b Ovipositor: Kramer and Morris, 2021b Immature stages Larva: Podeniene, 2000; Podeniene, 2003a; Podeniene et al., 2019 (first instar larva) Miscellaneous Quindroit, 2023 (habitat); Heiss and Malec, 2023 (habitat) Citations on biology (mainly from 2000 onward): Austria Habitat. Aistleitner et al., 2024: 33 (loc(s) Vorarlberg, see habitat photo above). Belgium Biology. Peeters, 2023: 230 (loc(s) Botanic Garden Jean Massart, Brussels-Capital Region, common spring and summer species mostly from moist habitats). Bosnia-Herzegowina Habitat. Graf et al., 2023: 34 (inventory of the upper Neretva, an unimpacted riverine system embedded in dense natural and near-natural forest; see paper for details). Bulgaria Habitat. Hubenov, 2025a: suppl: 2 (annotated list Diptera Stara Planina Mts, local distr, habitats and zoogeography). Habitat. Hubenov, 2025b: suppl: 1 (annotated list Diptera high Bulgarian mts, local distr, habitats and zoogeography). Czech Rep. Habitat. Stary et al., 2013a: 370 (on Tipulidae from Vraz near Pisek, with general habitat description). Denmark Habitat. Byriel and Rojas, 2017: 118 (among the 101 species in a study on cranefly hotspots in unmanaged and managed forests) (in Danish). Finland Habitat. Autio and Salmela, 2010: 49, 50 (habitats Aland Is, a common forest species also found in wooded mire). France Habitat. Dufour, 2003b: 87 (loc(s) with information on habitat(s)) (in French). Habitat. Kramer and Withers, 2007: 160 (species with larvae feeding in decaying leaves, citingCrisp and Lloyd, 1954). Habitat. Tillier and Dehalleux, 2019: 157-158 (loc(s) with habitat info, summary habitat) (in French). Habitat. Quindroit, 2020a: 19-20, 45-48 (annotated list with information on abundancy and habitat of the Tipuloidea of the Pays de la Loire regoin, distr) (in French). Habitat. Quindroit, 2020b: 128 (loc(s) Nord (59) and Pas-de-Calais (62) with notes on habitats and distr) (in French). Habitat. Tillier et al., 2020b: 188 (dept. Doubs (25): habitat est classe comme pelouse calcicole mesophile de l Est; see paper for details). Habitat. Herbrecht et al., 2021: 58 (loc(s) Loire-Atlantique (44) based on malaise-trapping with notes on habitats) (in French). Habitat. Quindroit, 2021a: 36 (loc(s), resultats d une campagne de piegeage dans le Pas-de-Calais (62); see paper for details on habitat). Habitat. Labat, 2022: 52-67 (preliminary study on Diptera of riparian corridors at several localities in France; supplements: loc(s) Puy-de-Dome (63)) (in French). Habitat. Quindroit and Lemoine, 2022a: 16 (loc(s) Aisne (02), Oise (60) and Somme (80) with notes on habitats and distr) (in French). Habitat. Quindroit, 2023: 62 (loc(s) Hautes-Alpes (05)) [locality and habitat are not given in the text but with the habitat figure]. Germany Habitat. Schacht et al., 2001: 426 (abbreviated habitat information). Habitat. Heiss and Merkel-Wallner, 2013: 21-24 (loc(s) Bayern, table with Malaise trap habitats and species; see paper for details). Biology. Heiss and Flugel, 2015: 271, 277 (loc(s) Hessen; review local distribution and biology, citing various authors). Habitat. Heiss et al., 2017: 179-182 (overview of collecting methods and habitats). Habitat. Heiss, 2017a: 562, 568, 570 (loc(s) Dinkelberg, Baden-Wurttemberg and table with details on presence in the various biotop-groups). Habitat. Dunk, 2018: 79, 95 (loc(s) Bayern with list of habtats). Habitat. Heiss et al., 2019: 250 (collected in a Fagus-Quercus forest, at the edge of a forest and at a Hartholzauwald) (in German). Great Britain Habitat. Godfrey, 2000a: 21. Habitat. Godfrey, 2003a: 45 (associated with woody debris). Habitat. Boardman, 2007: 22 (found along woodland egdes, dingle woodland, hedgerows and generaly anywhere with lush vegetation, Shropshire habitats include among others upland wet grassland, dry lowland heath). Habitat. Kramer, 2012h: 3 (Summer field meeting Scottish Highlands: often common in marshes and on river banks). Habitat. Showers, 2015: 2 (on craneflies in moth traps at Pitsford Water Nature Reserve in 2013 and 2014, including information for the two traps and photos of their habitat). Habitat. Kramer and Morris, 2021b: 14 (info on the species, on the distr and map of loc(s) Leicestershire and Rutland (VC55); usually associated with hedgerows and woodland edges where the larvae can be found in moist soil). Habitat. Devlin et al., 2022: 35 (list of Tipulidae observations across Wales, 1976-2019, in paper on the impact of climate change; observations: 223; habitat: Damp grasslands and woodland edge). Greece Habitat. Oosterbroek, 2009c: 183 (in wet deciduous forests). Lithuania Habitat. Podeniene, 2000: 3-10 (larvae were found in sandy soil in a Pinus forest, in dunes, under moss; (various adult habitats listed after Podenas, 1995)). Habitat. Podeniene, 2003a: 1-295 (larva, descr, figs, habitat) (in Lithuanian). Habitat. Podeniene, 2003b: 12, 23, 24, 29 (larval habitat: uppermost layer of soil in western taiga type forest, uppermost layer of soil under mosses). Biology. Podeniene et al., 2019: 92, 95-97, 105-109 (descr first instar larva, rearing, figs). Netherlands Biology. Peeters, 2023: 230 (common spring and summer species mostly from moist habitats). Romania Habitat. Parvu, 2003: 234. Habitat. Ujvarosi, 2003: 57. Habitat. Ujvarosi and Poti, 2006: 257 (habitat in Dupa Lunca marshland: predominantly shrubs along rivers and streams and swamp with bushes and trees). Habitat. Ujvarosi et al., 2011b: 111 (loc(s) Dupa Lanca marshy area near Voslobeni) (in Hungarian, species list with habitat information in English). Habitat. Kolcsar et al., 2013: 74 (loc(s) Cluj-Napoca and surroundings; list of habitats with altitudes). Russia Habitat. Lantsov, 2002c: 114 (dry and moderately humid biotopes of deciduous and mixed forests) (in Russian) (information on the Teberdinsky State Nature Biosphere Reserve kindly provided by Vladimir Lantsov). on characters and taxonomy (mainly from 2000 onward): China Taxonomy. : 5 (on a collection of craneflies from Jersey, collected between 1985 and 2002 by Tony Warne). Country not relevant Phylogeny. Gelhaus, 2005: 7, 19-20, 139-141 (also as fasciapennis) (phylogeny Eremotipula: outgroup taxon). Characters. Brodo, 2017a: 8 (comparison: short female genitalia). Phylogeny. Gao, Y. et al., 2023: 233-235 (phylogeny Tipuloidea based on mitochondrial genome sequencing). Phylogeny. Kang et al., 2023: 735-741 (phylogeny Tipuloidea based on mitochondrial genome sequencing). Phylogeny. Song et al., 2023: 7-9 (phylogeny Tipuloidea based on mitochondrial genome sequencing). Phylogeny. Gao, Y. et al., 2024: 267-288 (phylogeny Tipuloidea based on mitochondrial genome sequencing). Phylogeny. Zhang, L. et al., 2024: 897-901 (phylogeny Tipuloidea based on mitochondrial genome sequencing). Phylogeny. Feng et al., 2025: 4-17 (phylogeny Tipuloidea based on mitochondrial genome sequencing). Lithuania Immatures. Podeniene, 2000: 3-10 (key larva, comparison, figs). Immatures. Podeniene, 2003a: 1-295 (larva, descr, figs, habitat) (in Lithuanian). Immatures. Podeniene et al., 2019: 92, 95-97, 105-109 (descr first instar larva, rearing, figs). on distribution (mainly from 2000 onward): Austria Distribution. Vogtenhuber, 2011: 47 (checklist Austria, list of provinces). Distribution. Aistleitner et al., 2024: 33 (loc(s) Vorarlberg, see habitat photo above). Azerbaijan Distribution not confirmed. Snegovaya, 2021: 1133 (previous records could not be confirmed and not included in the checklist of the craneflies of Azerbaijan). Belgium Distribution. Peeters, 2023: 230 (loc(s) Botanic Garden Jean Massart, Brussels-Capital Region, common spring and summer species mostly from moist habitats). Bosnia-Herzegowina Distribution. Simova-Tosic and Oosterbroek, 2003: 49 (review literature). Distribution. Graf et al., 2023: 34 (inventory of the upper Neretva, an unimpacted riverine system embedded in dense natural and near-natural forest; see paper for details). Distribution. Kolcsar et al., 2023b: 34 (loc(s)). Bulgaria Distribution. Hubenov, 2021a: 27 (checklist with summary of distr in Bulgaria, references). Distribution. Hubenov, 2025a: suppl: 2 (annotated list Diptera Stara Planina Mts, local distr, habitats and zoogeography). Distribution. Hubenov, 2025b: suppl: 1 (annotated list Diptera high Bulgarian mts, local distr, habitats and zoogeography). Channel Is Distribution. Stubbs, 2022b: 5 (on a collection of craneflies from Jersey, collected between 1985 and 2002 by Tony Warne). China Distribution. : 5 (on a collection of craneflies from Jersey, collected between 1985 and 2002 by Tony Warne). Croatia Distribution. Simova-Tosic and Oosterbroek, 2003: 49 (review literature). Czech Rep. Distribution. Stary et al., 2005d: 36 (loc(s)). Distribution. Stary, 2009k: webpage (checklist). Distribution. Stary et al., 2013a: 370 (on Tipulidae from Vraz near Pisek, with general habitat description). Distribution. Mocek, 2014: 156 (loc(s) NE Bohemia). Denmark Distribution. Petersen et al., 2001b: 162 (checklist). Finland Distribution. Salmela, 2001a: 142, 149 (loc(s)). Distribution. Salmela, 2002: 30-32 (loc(s) lake Iidesjarvi) (in Finnish). Distribution. Autio and Salmela, 2010: 49 (loc(s) Aland Is). Distribution. Salmela, 2012b: 240 (annotated list of Finnish crane flies). Distribution. Salmela and Petrasiunas, 2014: 33 (checklist Finnish Tipulomorpha). France Distribution. Dufour, 2003b: 87 (loc(s) with information on habitat(s)) (in French). Distribution. Kramer and Withers, 2007: 160, 162 (loc(s) Ain (01)). Distribution. Withers, 2008: 137 (loc(s) Ain (01)). Distribution. Labat, 2013: 74 (on a sampling campaign on the Dordogne river sources, five Upper-Dordogne tributaries and a travertine, Auvergne; see paper for details) (in French). Distribution. Tillier and Dehalleux, 2019: 157-158 (new and published records Ile-de-France, loc(s) Seine-et-Marne (77), Yvelines (78), Essonne (91), Hauts-de-Seine (92), Seine-Saint-Denis (93) and Val-d’Oise (95)). Distribution. Quindroit, 2020a: 19-20, 45-48 (annotated list with information on abundancy and habitat of the Tipuloidea of the Pays de la Loire regoin, distr) (in French). Distribution. Quindroit, 2020b: 128 (loc(s) Nord (59) and Pas-de-Calais (62) with notes on habitats and distr) (in French). Distribution. Tillier et al., 2020b: 188 (dept. Doubs (25): habitat est classe comme pelouse calcicole mesophile de l Est; see paper for details). Distribution. Herbrecht et al., 2021: 58 (loc(s) Loire-Atlantique (44) based on malaise-trapping with notes on habitats) (in French). Distribution. Labat, 2021: 93 (on Diptera taxa recorded along the over-all length of the Dordogne River and some of its tributaries; see paper for the locality details) (in French). Distribution. Quindroit, 2021a: 36 (loc(s), resultats d une campagne de piegeage dans le Pas-de-Calais (62); see paper for details on habitat). Distribution. Labat, 2022: 52-67 (preliminary study on Diptera of riparian corridors at several localities in France; supplements: loc(s) Puy-de-Dome (63)) (in French). Distribution. Quindroit and Lemoine, 2022a: 16 (loc(s) Aisne (02), Oise (60) and Somme (80) with notes on habitats and distr) (in French). Distribution. Quindroit and Lemoine, 2022a: 34 (table specifying the presence in the five departments of the Hauts-de-France region). Distribution. Lemoine and Quindroit, 2023: 6 (loc(s) Finistere (29), Loiret (45), Nievre (58), Seine-et-Marne (77)). Distribution. Quindroit, 2023: 62 (loc(s) Hautes-Alpes (05)) [locality and habitat are not given in the text but with the habitat figure]. Distribution. Tillier et al., 2023: 175 (loc(s) Doubs (25), Jura (39) and Haute-Saone (70), with list of altitude(s) and habitat(s)) (in French). Distribution. Quindroit and Racine, 2025: 356 (table specifying the presence in the five departments of the Pays de la Loire region). Germany Distribution. Schacht et al., 2001: 426 (loc(s) Bayern). Distribution. Heiss, 2003: 11, 13 (in collection Janner and in Janner, 1937 under affinis, checklist Thuringen). Distribution. Schacht, 2005: 3 (checklist Bayern). Distribution. Barndt, 2010: 289 (loc(s) Brandenburg). Distribution. Hable et al., 2010b: 89 (loc(s) Bayern). Distribution. Schacht, 2010: 38 (checklist Diptera Bayern). Distribution. Merkel-Wallner et al., 2011: 214 (loc(s) Bayern). Distribution. Heiss and Merkel-Wallner, 2013: 21-24 (loc(s) Bayern, table with Malaise trap habitats and species; see paper for details). Distribution. Dunk and Kraus, 2014: 168 (loc(s) Bayern). Distribution. Merkel-Wallner, 2014: 49 (loc(s) Bayern). Distribution. Heiss and Flugel, 2015: 271, 277 (loc(s) Hessen; review local distribution and biology, citing various authors). Distribution. Wolz, 2016: 3 (one of the 13 Tipulidae species of which remains were found in the faeces of the Greater Horshoe Bat (Rhinolophus ferrumequinum) at Hohenburg, Bayern; final report in Wolz, 2018). Distribution. Heiss et al., 2017: 165 (loc(s) Thuringen). Distribution. Heiss, 2017a: 562, 568, 570 (loc(s) Dinkelberg, Baden-Wurttemberg and table with details on presence in the various biotop-groups). Distribution. Heiss, 2017b: 22 (checklist Tipulidae Thuringen). Distribution. Dunk, 2018: 79, 95 (loc(s) Bayern with list of habtats). Distribution. Heiss et al., 2019: 250 (loc(s); annotated checklist of the Tipulidae of Niedersachsen and Bremen, including the presence in the surrounding Bundeslandern and the Netherlands) (in German). Distribution. Heiss, 2019: 36, 39, 44 (loc(s) and checklist Mecklenburg-Vorpommern, see paper for further details). Distribution. Stuke, 2019: 239 (checklist Niedersachsen and Bremen with reference to original sources). Distribution. Stuke et al., 2020: 291 (loc(s) Niedersachsen). Distribution. Heiss and Malec, 2023: 317 (loc(s) Hessen). Distribution. Heiss and Malec, 2023: 329 (annotated checklist Hessen). Distribution. Kehlmaier et al., 2023: 219 (loc(s) Saarland). Distribution. Kopetz et al., 2024: 210 (loc(s) Thuringen). Distribution. Stuke et al., 2024: 169 (loc(s) Niedersachsen). Great Britain Distribution. Godfrey, 2000a: 21 (see paper for details of sites surveyed in Shropshire and into Wales). Distribution. Godfrey, 2003a: 45 (distr). Distribution. Hewitt et al., 2005: 37 (loc(s) Cumbria). Distribution. Boardman, 2007: 22 (loc(s), map). Distribution. Kramer, 2011g: 2 (loc(s) Devon). Distribution. Kramer, 2011j: 9 (loc(s) Leicestershire and Rutland (VC55)). Distribution. Kramer, 2012h: 3 (loc(s) Summer field meeting Scottish Highlands). Distribution. Hewitt, 2014: (provisional checklist Cumbrian Diptera). Distribution. Kramer, 2014c: 4 (in table of cranefly species recorded at Shapwick Heath NNR, as given in H. Audcent, 1949 and/or collected by J. Kramer in 2013). Distribution. Wolton et al., 2014: 26 (among Diptera associated with a hedge in Devon). Distribution. Chandler, 2015: 89 (loc(s) Bushy Park, Middlesex). Distribution. Showers, 2015: 2 (on craneflies in moth traps at Pitsford Water Nature Reserve in 2013 and 2014, including information for the two traps and photos of their habitat). Distribution. Drake, 2016: 22 (study on the relative importance to Diptera of pasture and ditch margins on Catcott Reserve, Somerset). Distribution. Brighton, 2017c: 30 (overall regional checklist Lancashire and Cheshire (VC58, 59 and 60) with notes on habitats and/or regional occurrence and/or status; see paper for details). Distribution. Lovegrove et al., 2018: 6-7 (among the cranefly species found in a survey of the historical site for Idiocera sexguttat at Stony Moors in the New Forest). Distribution. Kramer and Morris, 2021b: 14 (info on the species, on the distr and map of loc(s) Leicestershire and Rutland (VC55); usually associated with hedgerows and woodland edges where the larvae can be found in moist soil). Greece First record. Oosterbroek, 2009c: 183 (loc(s) Florina). Hungary Distribution. Toth, 2001a: 378 (literature records Somogy county). Ireland Distribution. Ashe et al., 2007a: 301, 319-321, 342, 353 (review Irish records, map). Distribution. Chandler et al., 2008: 6 (checklist). Italy Distribution. Oosterbroek, 2008c: 341 (annotated checklist South Tyrol [Prov. Bolzano]). Lithuania Distribution. Podeniene, 2000: 3-10 (loc(s)). Distribution. Pakalniskis et al., 2006: 27 (checklist). Montenegro Distribution. Simova-Tosic and Oosterbroek, 2003: 49 (review literature). Netherlands Distribution. Jong and Oosterbroek, 2002d: 41 (checklist). Distribution. Oosterbroek, 2014: 258 (loc(s) Friesland). Distribution. Barendregt, 2015: 9 (loc(s) prov. Groningen). Distribution. Steenis, 2019: 18 (loc(s) Noord-Brabant, Biesbosch). Distribution. Oosterbroek and Dek, 2020: 18 (Limoniidae and Tipulidae Nationaal Park Hollandse Duinen in 2018, Zuid-Holland). Distribution. Oosterbroek and Jong, 2020: 46 (Tipulidae Kaaistoep, Noord-Brabant). North Macedonia Distribution. Simova-Tosic and Oosterbroek, 2003: 49 (review literature). Norway Distribution. Hofsvang, 2010: 13 (loc(s)). Distribution. Hofsvang et al., 2019: 109-110, 142-143 (review Norwegian records, annotated checklist Nordic countries and for Norway according to the Strand regions). Distribution. Hofsvang, 2025: 8 (loc(s) AK). Poland Distribution. Palaczyk, 2001: 270 (loc(s)). Distribution. Skibinska and Chudzicke, 2007b: 78 (checklist). Romania First record. Erhan and Theowald, 1961: XXX (loc(s)). Distribution. Parvu, 2003: 234 (loc(s)). Distribution. Ujvarosi, 2003: 57 (review Romanian records, loc(s)). Distribution. Parvu, 2004: 187 (loc(s)). Distribution. Parvu, 2005: 149 (loc(s)). Distribution. Parvu, 2006: 261 (loc(s)). Distribution. Ujvarosi and Poti, 2006: 257 (loc(s) eastern Carpathians). Distribution. Ujvarosi, 2007: 232-233 (checklist Tipulidae). Distribution. Ujvarosi et al., 2011b: 111 (loc(s) Dupa Lanca marshy area near Voslobeni) (in Hungarian, species list with habitat information in English). Distribution. Keresztes, 2012: 31 (loc(s) Rimetea area [Alba County]). Distribution. Kolcsar et al., 2013: 74 (loc(s) Cluj-Napoca and surroundings; list of habitats with altitudes). Distribution. Torok et al., 2014: 18 (checklist Romania). Russia Distribution. Lantsov, 2002c: 114 (on the faunal structure, biotope(s) and chorology of crane-flies of the Teberdinsky State Nature Biosphere Reserve (North Caucasus: Karachay-Cherkess Rep.)) (in Russian). Distribution. Pilipenko, 2009a: 211 (annotated checklist Tipulidae Central European Territory, loc(s) RUC: Tverskaya oblast, Moskovskaya oblast, distr). Distribution. Jakovlev et al., 2014: 310 (loc(s) RUN: Kareliya). Distribution. Paramonov and Klepikov, 2014: 37 (loc(s) Yaroslavskaya oblast). Distribution. Paramonov, 2014: 39-43 (revision coll. Eversmann), loc(s) RUE: Tatarstan Rep.) (in Russian. Distribution. Ruchin and Pilipenko, 2015: 59 (loc(s), checklist RUC: Mordoviya Rep.). Distribution. Lantsov, 2016: 165-170 (on craneflies from Arkhyz, Karachay-Cherkessia Rep., North Caucasus) (in Russian). Distribution. Paramonov and Pilipenko, 2016: 101 (loc(s) RUC: Tverskaya oblast) (in Russian). Distribution. Paramonov, 2016: 131 (loc(s) RUC: Chuvash Rep.) (in Russian). Distribution. Paramonov, 2017a: 173 (loc(s) RUC: Chuvash Rep.) (in Russian). Distribution. Paramonov, 2018a: 185 (loc(s) RUC: Chuvash Rep., distr) (in Russian). Distribution. Paramonov, 2018b: 62 (loc(s) RUS: Saratovskaya oblast) (in Russian). Distribution. Pilipenko et al., 2020: 364 (loc(s) RUC: Mordoviya Rep., distr). Serbia Distribution. Simova-Tosic and Oosterbroek, 2003: 49 (review literature). Slovakia Distribution. Stary, 2009g: 44 (loc(s) Polana area). Distribution. Stary, 2009k: webpage (checklist). Slovenia Distribution. Simova-Tosic and Oosterbroek, 2003: 49 (review literature). Spain Distribution. Eiroa and Baez, 2002b: 79 (checklist). Distribution. Aistleitner, 2011: 112 (loc(s) Palencia). Distribution. Eiroa and Carles-Tolra, 2019: 133 (loc(s) Barcelona, list of Spanish provinces, distr) (in Spanish). Sweden Distribution. Fritz and Lindstrom, 2013: 63 (loc(s) Hallands County) (in Schwedish). Switzerland Distribution. Dufour and Merz, 2012: 355 (annotated checklist canton Geneva). on flight period (mainly from 2000 onward): Austria Aistleitner et al., 2024: month(s): 6. Belgium Peeters, 2023: month(s): 6. Bosnia-Herzegowina Kolcsar et al., 2023b: month(s): 6. Channel Is Stubbs, 2022b: month(s): 6. Czech Rep. Stary et al., 2005d: month(s): 6-7. Stary et al., 2013a: month(s): 6. Finland Salmela, 2001a: month(s): 6-7. France Dufour, 2003b: month(s): 6. Kramer and Withers, 2007: month(s): 6-7. Tillier and Dehalleux, 2019: month(s): 5-7. Quindroit, 2020a: month(s): 5-7. Quindroit, 2020b: month(s): 5-7. Quindroit and Lemoine, 2022a: month(s): 5-7. Lemoine and Quindroit, 2023: month(s): 4-6. Quindroit, 2023: month(s): 6. Tillier et al., 2023: month(s): 6-8. Germany Schacht et al., 2001: month(s): 5-8. Heiss and Merkel-Wallner, 2013: month(s): 5-7. Heiss et al., 2017: month(s): 6-7. Heiss, 2017a: month(s): 6-7. Dunk, 2018: month(s): 5-7. Heiss et al., 2019: month(s): 6. Heiss, 2019: month(s): 6-7. Heiss and Malec, 2023: month(s): 6-7. Kehlmaier et al., 2023: month(s): 6. Stuke et al., 2024: month(s): 6. Great Britain Godfrey, 2000a: month(s): 6. Boardman, 2007: month(s): 6-7. Kramer, 2011j: month(s): 7. Kramer, 2012h: month(s): 7. Kramer, 2014c: month(s): 7. Chandler, 2015: month(s): 6-7. Kramer and Morris, 2021b: month(s): 5-7. Greece Oosterbroek, 2009c: month(s): 6. Ireland Ashe et al., 2007a: month(s): 6-8. Netherlands Barendregt, 2015: month(s): 5. Oosterbroek and Dek, 2020: month(s): 5-6. Norway Hofsvang et al., 2019: month(s): 5-8. Hofsvang, 2025: month(s): 6. Romania Parvu, 2003: month(s): 6-7. Ujvarosi, 2003: month(s): 5-8. Parvu, 2004: month(s): 6-7. Parvu, 2005: month(s): 6. Parvu, 2006: month(s): 6. Ujvarosi et al., 2011b: month(s): 5-6. Kolcsar et al., 2013: month(s): 5-7. Russia Lantsov, 2002c: month(s): 5-7. Pilipenko, 2009a: month(s): 6-7. Paramonov and Klepikov, 2014: month(s): 6-8. Ruchin and Pilipenko, 2015: month(s): 6. Paramonov and Pilipenko, 2016: month(s): 6. Paramonov, 2016: month(s): 6. Paramonov, 2017a: month(s): 6. Paramonov, 2018a: month(s): 7. Paramonov, 2018b: month(s): 6-7. Pilipenko et al., 2020: month(s): 6-7. Slovakia Stary, 2009g: month(s): 7. Spain Aistleitner, 2011: month(s): 7. Eiroa and Carles-Tolra, 2019: month(s): 7. on altitude (mainly from 2000 onward): Austria Aistleitner et al., 2024: altitude: 1000 m. Bosnia-Herzegowina Kolcsar et al., 2023b: altitude: 640 m. Bulgaria Hubenov, 2021a: altitude: 400-600 m. Hubenov, 2025a: altitude: 400-600 m. Hubenov, 2025b: altitude: 400-660 m. France Dufour, 2003b: altitude: 1080-1150 m. Labat, 2021: altitude: 580-1357 m. Quindroit, 2023: altitude: 1900 m. Germany Merkel-Wallner et al., 2011: altitude: <700-799 m. Heiss et al., 2017: altitude: 335 m. Stuke et al., 2024: altitude: 150-270 m. Greece Oosterbroek, 2009c: altitude: 850-1000 m. Romania Parvu, 2003: altitude: 800 m. Ujvarosi, 2003: altitude: 700-1300 m. Ujvarosi and Poti, 2006: altitude: 670 m. Russia Paramonov, 2018b: altitude: 302 m. Spain Aistleitner, 2011: altitude: 1000 m. Eiroa and Carles-Tolra, 2019: altitude: 1235 m.