Details for Tipula (Savtshenkia) benesignata

Name: Tipula (Savtshenkia) benesignata Mannheims, 1954 Publication: Notul. Ent. 34: 35 (as Tipula) Status: Recognized taxon. Classification: Family Tipulidae Subfamily Tipulinae Descriptive note: Phylogeny: Jong, 1994c. Holotype [= Lectotype] designation: Mannheims and Pechlaner, 1963. Type-material: Sinclair and Dorchin, 2010. See also boreosignata and eleonorae. Keys: Peeters en Oosterbroek, 2016a (craneflies Netherlands, Belgium and Luxembourg) (in Dutch) habitus male habitus female habitus female body part(s) head body part(s) head and thorax plus 19 more images of body part(s), body part(s), body part(s), body part(s), body part(s), hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, ovipositor, ovipositor, ovipositor, wing, wing, wing and habitat. Distribution: Austria, Belgium, Bulgaria, Croatia, Czech Rep., Finland, France, Germany, Greece (incl. Euboea [Evia]), Hungary, Italy (north), Kosovo, Luxembourg, Montenegro, Norway, Poland, Romania, Slovakia, Slovenia, Sweden, Switzerland, Ukraine; Russia: RUN, RUW, RUC, RUE, North Caucasus; Turkey (Asiatic part: Bursa, Canakkale);; Kyrgyzstan (Tien Mts). Note: see the manual for abbreviations: present-day Russia. Region(s): Westpalaearctic;; Eastpalaearctic General note: Added: Euboea (Theischinger, 1983). For France check Dufour and Brunhes, 1990, and citations below. For Finland check Salmela, 2012b, 2012c, and citations below. For Norway check Hofsvang et al., 2019, and citations below. Map: Jong, 1998 (general) (Auvergne omitted); Przhiboro, 2018e (Leningrad prov.) Figure references: Habitus: Przhiboro, 2009 (female); Przhiboro, 2018e (female) Immature stages Larva: Savchenko, 1963b Miscellaneous Przhiboro, 2009 (habitat); Syratt, 2022a (habitat) Citations on biology (mainly from 2000 onward): Finland Habitat. Rasimus, 2022: 6 (on the craneflies fauna of Tavastia australis biogeographical province) (in Finnish). France Habitat. Dufour, 2003b: 89 (loc(s) with information on habitat(s)) (in French). Habitat. Kramer and Langlois, 2019b: 229 (details on habitat and biology, citing various authors). Habitat. Tillier and Oosterbroek, 2019: 236 (loc(s) and distr Mercantour; habitat: foret de Melezes [Larix] avec sous-bois de Noisetiers [Hazel]). Kosovo Habitat. Bilalli et al., 2021: 147 (collected from nearby riverside habitats within moist forests). Poland Biology. Syratt, 2022a: 6 (The site in Krakow is a shallow stream in a human-modified fragment of Alder [Alnus] riparian forest; tufts of wet moss on rocks and logs are present but overall scarce; review habitats citing various authors, period of flight). Romania Habitat. Erhan, 1986b: XXX (Fagus forest). Habitat. Ujvarosi, 2003: 59 (coniferous forest). Russia Habitat. Lantsov, 2002c: 114 (larvae under moss cushions in springs and streams) (in Russian) (information on the Teberdinsky State Nature Biosphere Reserve kindly provided by Vladimir Lantsov). Biology. Przhiboro, 2003: 362, 363 (habitat: larvae abundant in water margin zone of an oligotrophic lake, developing in microhabitats with dense cover of mosses; extensive notes on feeding and gut contents, species is possibly a poly-saprophage with elements of bryophagy and facultative predation; rearing). Habitat. Lantsov, 2009a: 75 (ecology) (in Russian). Habitat. Przhiboro, 2009: 229-232 (reared from semiaquatic substrata from a cold spring habitat; the paper includes further habitat details including photos). Biology. Przhiboro, 2018e: 11, 94-95 (Red data book Leningrad prov.; figs larva, adult female, short descr, distr, local records with map, habitats incl larval habitats, rearing, data on bionomics, proposals for protection of local populations) (in Russian). Turkey Habitat. Koc et al., 2015: 52 (collected from river side within moist forests). on characters and taxonomy (mainly from 2000 onward): Russia Description. Przhiboro, 2018e: 11, 94-95 (Red data book Leningrad prov.; figs larva, adult female, short descr, distr, local records with map, habitats incl larval habitats, rearing, data on bionomics, proposals for protection of local populations) (in Russian). on distribution (mainly from 2000 onward): Austria Distribution. Aistleitner, 2011: 112 (loc(s) Vorarlberg). Distribution. Vogtenhuber, 2011: 50 (checklist Austria, list of provinces). Distribution. Aistleitner, 2015: 97 (loc(s) Vorarlberg). Distribution. Heiss et al., 2016: 75, 81 (loc(s) Oberosterreich and Salzburg). Bulgaria First record. Kolcsar et al., 2023a: 22 (loc(s) Balkanets and Vitosha Mts). Croatia Distribution. Simova-Tosic and Oosterbroek, 2003: 54 (review literature). Distribution. Kolcsar et al., 2023b: 35 (loc(s)). Czech Rep. Distribution. Stary et al., 2005d: 36 (loc(s)). Distribution. Stary, 2009k: webpage (checklist). Finland Distribution. Salmela, 2012b: 241 (annotated list of Finnish crane flies). Distribution. Salmela and Petrasiunas, 2014: 34 (checklist Finnish Tipulomorpha). Distribution. Rasimus, 2022: 6 (on the craneflies fauna of Tavastia australis biogeographical province) (in Finnish). France Distribution. Dufour, 2003b: 89 (loc(s) with information on habitat(s)) (in French). Distribution. Kramer and Langlois, 2019b: 228-229 (list of craneflies captured 25-27 september 2019 in Ravin de Valbois, Doubs (25), loc(s), distr) (see also Tillier et al., 2023: 178). Distribution. Tillier and Oosterbroek, 2019: 236 (loc(s) and distr Mercantour; habitat: foret de Melezes [Larix] avec sous-bois de Noisetiers [Hazel]). Distribution. Tissot et al., 2021: 217 (the Diptera of the Reserves Naturelles Nationales du Doubs (25), results of Malaise trapping since 2009) (in French). Germany Distribution. Schacht et al., 2001: 428 (loc(s) Bayern). Distribution. Heiss, 2003: 14 (checklist Thuringen). Distribution. Schacht, 2005: 4 (checklist Bayern). Distribution. Schacht, 2010: 38 (checklist Diptera Bayern). Distribution. Heiss, 2017b: 23 (checklist Tipulidae Thuringen). Distribution. Heiss et al., 2021: 241, 244 (found in cave(s) in Bayern, see paper on details of which part of the cave(s)) (in German). Distribution. Heiss and Malec, 2023: 329 (annotated checklist Hessen). Italy Distribution. Oosterbroek, 2008c: 341 (annotated checklist South Tyrol [Prov. Bolzano]). Kosovo First record. Bilalli et al., 2021: 147 (loc(s) Kosovo, distr). Montenegro Distribution. Simova-Tosic and Oosterbroek, 2003: 54 (review literature). Norway Distribution. Hofsvang et al., 2019: 120, 144-145 (review Norwegian records, annotated checklist Nordic countries and for Norway according to the Strand regions). Poland First record. Syratt, 2022a: 6 (loc(s) Krakow Bridge (Pomost Krakowski), distr). Romania First record. Erhan, 1986b: XXX (loc(s)). Distribution. Ujvarosi, 2003: 59 (review Romanian records, loc(s)). Distribution. Ujvarosi, 2007: 232-233 (checklist Tipulidae). Distribution. Torok et al., 2014: 19 (checklist Romania). Russia Distribution. Lantsov, 2002c: 114 (on the faunal structure, biotope(s) and chorology of crane-flies of the Teberdinsky State Nature Biosphere Reserve (North Caucasus: Karachay-Cherkess Rep.)) (in Russian). First record. Przhiboro, 2003: 362, 363 (loc(s) RUN: Kareliya, distr). Distribution. Lantsov, 2009a: 75 (loc(s) Central Caucasus, distr). Distribution. Pilipenko, 2009a: 215 (annotated checklist Tipulidae Central European Territory, loc(s) RUC: Moskovskaya oblast, distr). Distribution. Przhiboro, 2009: 229-232 (loc(s) RUW: Leningradskaya oblast). Distribution. Przhiboro, 2018e: 11, 94-95 (Red data book Leningrad prov.; figs larva, adult female, short descr, distr, local records with map, habitats incl larval habitats, rearing, data on bionomics, proposals for protection of local populations) (in Russian). Distribution. Kolcsar et al., 2023a: 22 (loc(s) RUE: Tatarstan Rep.). Slovakia Distribution. Stary, 2009g: 44 (loc(s) Polana area). Distribution. Stary, 2009k: webpage (checklist). Slovenia Distribution. Simova-Tosic and Oosterbroek, 2003: 54 (review literature). Switzerland Distribution. Dufour and Merz, 2012: 355 (annotated checklist canton Geneva). Turkey First record. Koc et al., 2015: 50 (results of Tipulidae investigations in the 11 provinces of the Marmara region which forms the passage between European and Asiatic Turkey). on flight period (mainly from 2000 onward): Austria Aistleitner, 2011: month(s): 11. Aistleitner, 2015: month(s): 11. Heiss et al., 2016: month(s): 8, 10-11. Bulgaria Kolcsar et al., 2023a: month(s): 9-10. Croatia Kolcsar et al., 2023b: month(s): 9. Finland Rasimus, 2022: month(s): 9-10. France Dufour, 2003b: month(s): 10. Kramer and Langlois, 2019b: month(s): 9. Tillier and Oosterbroek, 2019: month(s): 9. Germany Schacht et al., 2001: month(s): 9-10. Heiss et al., 2021: month(s): 9. Kosovo Bilalli et al., 2021: month(s): 10. Norway Hofsvang et al., 2019: month(s): 8-10. Poland Syratt, 2022a: month(s): 9. Romania Ujvarosi, 2003: month(s): 10. Russia Lantsov, 2002c: month(s): 9. Przhiboro, 2003: month(s): 8. Lantsov, 2009a: month(s): 9. Pilipenko, 2009a: month(s): 8-9, 11. Przhiboro, 2009: month(s): 8-9. Kolcsar et al., 2023a: month(s): 8. Slovakia Stary, 2009g: month(s): 10. Turkey Koc et al., 2015: month(s): 10. on altitude (mainly from 2000 onward): Austria Aistleitner, 2011: altitude: 1000 m. Aistleitner, 2015: altitude: 1000 m. Bulgaria Kolcsar et al., 2023a: altitude: 723 m. Croatia Kolcsar et al., 2023b: altitude: 545 m. France Dufour, 2003b: altitude: 70-1800 m. Tillier and Oosterbroek, 2019: altitude: 1850-1900 m. Germany Schacht et al., 2001: altitude: 1650 m. Heiss et al., 2021: altitude: 775 m. Romania Ujvarosi, 2003: altitude: 1450 m. Russia Lantsov, 2002c: altitude: 1330-1520 m. Lantsov, 2009a: altitude: 1571-1737 m. Kolcsar et al., 2023a: altitude: 80 m. Turkey Koc et al., 2015: altitude: 52-970 m.