| Name: | Rhypholophus varius (Meigen and Wiedemann, 1818) | | Publication: | In: Meigen, 1818, Syst. Beschr. 1: 115 (as Erioptera) | | Status: | Recognized taxon. | | Classification: | Family Limoniidae Subfamily Chioneinae
| | Keys: | Boardman, 2016 (craneflies Shropshire); Stubbs and Kramer, 2016j; Stubbs, 2021 (British craneflies); Quindroit, 2026a (Rhypholophus France, in French and English) | | Variant spelling(s): | varia |  |  |  |  |  |
habitus male | habitus male | habitus male | habitus male | habitus copula |
plus 25 more images of body part(s), body part(s), body part(s), body part(s), body part(s), body part(s), hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, ovipositor, ovipositor, ovipositor, ovipositor, wing, wing, wing, wing, wing, wing, habitat and habitat. | | | | Distribution: | Austria, Belgium, Czech Rep., Denmark, Finland, France, Germany, Great Britain, Ireland, Lithuania, Netherlands, Norway, Poland, Romania, Sweden, Switzerland, Ukraine; Russia: RUN, RUW. Note: see the manual for abbreviations: present-day Russia. | | Region(s): | Westpalaearctic | | General note: | For Finland check Salmela, 2012b, 2012c, and citations below. For Norway check Olsen et al., 2018, and citations below. For Great Britain and Ireland check Stubbs, 2021, and citations below. Turkey in Koc et al., 2016 refers to bifurcatus (teste J. Stary and P. Oosterbroek). Hangary in Ujvarosi, 2004 refers to bifurcatus (see Kolcsar et al., 2021b). | | Map: | Podenas et al., 2006 (Switzerland); Ashe et al., 2007b (Ireland); Boardman, 2007 (Shropshire); Quindroit, 2026a (France) | Figure references: | Habitus: Quindroit, 2026a (male) Head: Quindroit, 2026a Wing: Podenas et al., 2006; Dek and Oosterbroek, 2013; Quindroit, 2026a Hypopygium: Mendl, 1973e (the figures in Podenas et al., 2006 refer to bifurcatus); Dek and Oosterbroek, 2013; Kramer and Morris, 2022a; Quindroit, 2026a Ovipositor: Quindroit, 2026a Other figures: Quindroit, 2026a (thorax)
| | | | Citations | | | Country not relevant Habitat. Quindroit, 2026a: 30 (review general distribution and habitats, citing various authors) (in French). | | Czech Rep. Habitat. Stary and Vonicka, 2018: 61 (on Limoniidae and Pediciidae of two localities in northern Bohemia, with list of localities, altitudes and habitats). | | Finland Habitat. Salmela, 2001a: 144, 146, 151 (terrestrial species of the dryest locations, citing Brinkmann, 1991). Habitat. Penttinen et al., 2010: 487 (red list status incl. criteria, habitats, causes of threat, threat factors). Habitat. Salmela et al., 2019: 528 (red list status, habitat types, threat factors). Habitat. Rasimus, 2022: 5-6 (on the craneflies fauna of Tavastia australis biogeographical province) (in Finnish). | | France Biology. Quindroit, 2020a: 29, 45-48 (annotated list with information on abundancy and habitat of the Tipuloidea of the Pays de la Loire regoin, distr) (in French). | | Germany Habitat. Ruckert, 2005: 47, 53 (found in a study on the association with the type of source: limnokrene, rheokrene, or helokrene in Schlesig-Holstein; see paper for details, as well as Anhang A2-A10). Habitat. Wittrock, 2005: 85 (summer emergence; rheokrene and limnokrene). Habitat. Ruckert et al., 2007: 56 (Limnophil). Habitat. Lehmann and Reusch, 2009: 122 (from around lowland springs). | | Great Britain Habitat. Godfrey, 2003a: 44 (associated with woody debris). Habitat. Roper, 2005: 213 (emerging from a standing, dead Quercus tree-trunk). Habitat. Ashe et al., 2007b: 390 (in Britain a common species of moist and wet woods on clay soils). Habitat. Boardman, 2007: 58 (prefers more acid substrates such as carr and seepages). Habitat. Drake, 2011a: 25 (among the Diptera from a wet Alder [Alnus] wood in Devon). Habitat. Kramer, 2011j: 25 (a woodland species). Habitat. Brighton, 2017c: 26 (overall regional checklist Lancashire and Cheshire (VC58, 59 and 60) with notes on habitats and/or regional occurrence and/or status; see paper for details). Habitat. Cunningham, 2022: 30 (loc(s) Bovey Valley woodlands, Devon; see paper for habitat details). Habitat. Kramer and Morris, 2022a: 22 (info on the species, on the distr and map of loc(s) Leicestershire and Rutland (VC55); all VC55 records came [from] damp habitats especially in woodlands). | | Lithuania Habitat. Podeniene, 2001a: 83, 84 (larvae were found in the intertidal zones of small rivers in silt and leaf fallings). Habitat. Podeniene, 2003b: 11, 23, 24, 29 (larval habitat: sandy loam under grassy vegetation or under fallings not completely covered by forest, at the uppermost layer of soil under mosses). Habitat. Podeniene, 2009: 137 (develops in mud nearby waters and under leaf litter in soil). Biology. Gorban and Podeniene, 2022: 5, 8 (loc(s); reared from Ash [Fraxinus excelsior], species with a typical development habitat in moist soil or leaf litter; see paper for details). | | Romania Habitat. Ujvarosi et al., 2011b: 107 (loc(s) Dupa Lanca marshy area near Voslobeni) (in Hungarian, species list with habitat information in English). | | Sweden Habitat. Lindstrom and Fritz, 2015: 44, 48 (distributin and ecology in five fens in Hallands County) (in Schwedish). | | | | Country not relevant Phylogeny. Michelsen, 1996: 101 (ingroup taxon in a study titled: Neodiptera: New insights into the adult morphology and higher level phylogeny of Diptera (Insecta)). Key. Quindroit, 2026a: 28, 31-39 (belongs to the haemorrhoidalis group with characters of the group; fig(s); key [in French and English]). | | Great Britain Characters. Boardman, 2007: 58 (characters). | | Netherlands Characters. Dek and Oosterbroek, 2013: 88-89 (chars, figs, key species Netherlands). | | | | Country not relevant Distribution. Quindroit, 2026a: 30 (review general distribution and habitats, citing various authors) (in French). | | Czech Rep. Distribution. Stary, 2004c: 156 (loc(s) Bohemia, distr). Distribution. Stary, 2009i: webpage (checklist). Distribution. Stary and Vonicka, 2018: 61 (on Limoniidae and Pediciidae of two localities in northern Bohemia, with list of localities, altitudes and habitats). | | Denmark Distribution. Petersen and Jong, 2001b: 151 (checklist). | | Finland Distribution. Salmela, 2001a: 144, 146, 151 (loc(s)). Distribution. Salmela, 2012b: 239 (annotated list of Finnish crane flies). Distribution. Salmela and Petrasiunas, 2014: 26 (checklist Finnish Tipulomorpha). Distribution. Salmela et al., 2019: 528 (red list status, habitat types, threat factors). Distribution. Rasimus, 2022: 5-6 (on the craneflies fauna of Tavastia australis biogeographical province) (in Finnish). | | France Distribution. Quindroit, 2020a: 29, 45-48 (annotated list with information on abundancy and habitat of the Tipuloidea of the Pays de la Loire regoin, distr) (in French). Distribution. Quindroit, 2022a: 25-28 (additions to the annotated list of the Tipuloidea of Pays de la Loire as given in Quindroit, 2020a). Distribution. Quindroit and Racine, 2025: 358 (table specifying the presence in the five departments of the Pays de la Loire region). Distribution. Quindroit, 2026a: 30, 37 (altitudes in France below 400 m, in Switzerland at about 600 m, citing various authors, map) (in French). | | Germany Distribution. Reusch and Oosterbroek, 2000: 160 (checklist German Bundeslander). Distribution. Reusch and Bellstedt, 2001b: 74 (checklist Thuringen). Distribution. Ruckert, 2005: 47, 53 (found in a study on the association with the type of source: limnokrene, rheokrene, or helokrene in Schlesig-Holstein; see paper for details, as well as Anhang A2-A10). Distribution. Wittrock, 2005: 85 (loc(s) Schleswig-Holstein). Distribution. Ruckert et al., 2007: 56 (loc(s) Schleswig-Holstein). Distribution. Lehmann and Reusch, 2009: 122 (loc(s) Niedersachsen). Distribution. Stuke, 2019: 146 (checklist Niedersachsen and Bremen with reference to original sources). | | Great Britain Distribution. Godfrey, 2003a: 44 (distr). Distribution. Roper, 2005: 213 (loc(s) Sussex). Distribution. Boardman, 2007: 58 (loc(s), map). Distribution. Skidmore, 2009: 127 (review of occurrence on the Western Isles of Scotland, distr). Distribution. Drake, 2011a: 25 (loc(s) Devon). Distribution. Kramer, 2011j: 17 (loc(s) Leicestershire and Rutland (VC55)). Distribution. Hewitt, 2014: (provisional checklist Cumbrian Diptera). Distribution. Kramer, 2016a: 1 (loc(s) Sheffield). Distribution. Brighton, 2017c: 26 (overall regional checklist Lancashire and Cheshire (VC58, 59 and 60) with notes on habitats and/or regional occurrence and/or status; see paper for details). Distribution. Cunningham, 2022: 30 (loc(s) Bovey Valley woodlands, Devon; see paper for habitat details). Distribution. Kramer and Morris, 2022a: 22 (info on the species, on the distr and map of loc(s) Leicestershire and Rutland (VC55); all VC55 records came [from] damp habitats especially in woodlands). | | Hungary Misinterpretation. Ujvarosi, 2004: 119 (removed for Hungary in Kolcsar et al., 2021b, see under R. bifurcatus). | | Ireland Distribution. Ashe et al., 2007b: 363, 390, 406 (review Irish records, map). Distribution. Chandler et al., 2008: 10 (checklist). | | Lithuania Distribution. Podeniene, 2001a: 83, 84 (loc(s)). Distribution. Pakalniskis et al., 2006: 24 (checklist). Distribution. Gorban and Podeniene, 2022: 5, 8 (loc(s); reared from Ash [Fraxinus excelsior], species with a typical development habitat in moist soil or leaf litter; see paper for details). | | Netherlands Distribution. Jong and Oosterbroek, 2002b: 30 (checklist). Distribution. Oosterbroek, 2013: 10 (Dutch provinces). Distribution. Stary, 2020a: 45 (Limoniidae Kaaistoep, Noord-Brabant). | | Norway Distribution. Olsen et al., 2018: 156-157 (annotated checklist Nordic countries and for Norway according to the Strand regions). | | Poland Distribution. Wiedenska, 2007a: 76 (checklist). Distribution. Wiedenska, 2008: 45 (loc(s) Parowy Janinowski nature reserve near Lodz) (in Polish). Distribution. Wiedenska, 2010: 135 (loc(s) Parku Krajobrazowego Wzniesien Lodzkich, Lodz). | | Romania First record. Ujvarosi, 2007: 228-231 (checklist Limoniidae, no further details). Distribution. Ujvarosi et al., 2011b: 107 (loc(s) Dupa Lanca marshy area near Voslobeni) (in Hungarian, species list with habitat information in English). | | Russia Distribution. Jakovlev et al., 2014: 309 (loc(s) RUN: Kareliya). | | Sweden Distribution. Lindstrom and Fritz, 2015: 44, 48 (distributin and ecology in five fens in Hallands County) (in Schwedish). | | Switzerland Distribution. Podenas et al., 2006: 128, 339 (map, vertical distr per thermic level). | | Turkey Misinterpretation. Koc et al., 2016: 4-7 (first record for Turkey refers to bifurcatus) (teste J. Stary and P. Oosterbroek). | | | | Czech Rep. Stary, 2004c: month(s): 9. Stary and Vonicka, 2018: month(s): 9-10. | | Finland Salmela, 2001a: month(s): 8. Rasimus, 2022: month(s): 9. | | France Quindroit, 2020a: month(s): 9. Quindroit, 2026a: month(s): 9-10. | | Great Britain Boardman, 2007: month(s): 8-10. Kramer, 2011j: month(s): 9. Kramer, 2016a: month(s): 9. | | Ireland Ashe et al., 2007b: month(s): 9-10. | | Netherlands Oosterbroek, 2013: month(s): 8-10. | | Poland Wiedenska, 2008: month(s): 9. | | Romania Ujvarosi et al., 2011b: month(s): 10. | | Switzerland Podenas et al., 2006: month(s): 9. |
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