Details for Metalimnobia (Metalimnobia) bifasciata Name: | Metalimnobia (Metalimnobia) bifasciata (Schrank, 1781) | Publication: | Enum. ins. Austr. indig.: 429 (as Tipula) | Status: | Recognized taxon, Synonym(s): xanthoptera (Meigen, 1804); ?vittata Matsumura, 1911; avis (Alexander, 1918); flavoabdominalis (Alexander, 1918). | Classification: | Family Limoniidae Subfamily Limoniinae
| Descriptive note: | Details on colouration in Alexander, 1924n (as avis Alexander). Characters and synonymy in Sasakawa, 2002 (in Japanese). | Keys: | Podenas and Gelhaus, 2007 (Limoniinae Mongolia and nearby); Krivosheina and Krivosheina, 2010 (Metalimnobia larvae (5 species) and pupae (3 species)); Mao and Yang, 2010a (Metalimnobia China); Peeters en Oosterbroek, 2014a (craneflies Netherlands, Belgium and Luxembourg) (in Dutch); Boardman, 2016 (craneflies Shropshire); Podenas and Byun, 2016 (Metalimnobia Korea); Stubbs and Kramer, 2016h; Stubbs, 2021 (British craneflies); Jiang and Zhang, 2020 (Metalimnobia China) |  |  |  |  |  |
habitus male | habitus male | habitus male | habitus male | habitus female |
plus 28 more images of body part(s), body part(s), body part(s), body part(s), body part(s), hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, ovipositor, ovipositor, wing, wing, wing, wing, wing, wing, habitat, habitat, habitat, habitat, habitat, habitat, habitat, habitat and habitat. | | | Distribution: | Albania, Austria, Belarus, Belgium, Bulgaria, Croatia, Czech Rep., Denmark, Estonia, Finland, France, Germany, Great Britain, Hungary, Ireland, Italy, Latvia, Lithuania, Luxembourg, Netherlands, Norway, Poland, Romania, Serbia, Slovakia, Slovenia, Spain, Sweden, Switzerland, Turkey (European part: widespread), Ukraine; Russia: RUN, RUW, RUC, RUE, RUS, North Caucasus; Georgia, Turkey (Asiatic part: Karabuk, Zonguldak);; Russia: WS (north), ES (Siberia north and eastward to Yakutiya), FE (Kamchatka, Amurskaya oblast, Khabarovskiy kray, Primorskiy kray, Sakhalin (incl. Kuril Is)); Kazakhstan (east), Tajikistan; Mongolia, North Korea, South Korea, Japan (Hokkaido, Honshu, Shikoku), China (see citations for provinces);; China (Guizhou). Note: see the manual for abbreviations: present-day Russia. | Region(s): | Westpalaearctic;; Eastpalaearctic;; Oriental | General note: | Added: Croatia (Langhoffer, 1917), Slovenia (Lackschewitz, 1928), Serbia (Simova-Tosic and Vukovic, 1996), RUN (Kareliya) (Yakovlev and Polevoi, 1997). For Italy check Stary and Oosterbroek, 1996, and citations below. For Finland check Salmela, 2012b, 2012c, and citations below. For Norway check Olsen et al., 2018, and citations below. For Great Britain and Ireland check Stubbs, 2021, and citations below. | Map: | Koc et al., 2005b (NW Turkey); Podenas et al., 2006 (Switzerland); Boardman, 2007 (Shropshire); Podenas and Byun, 2016 (Korea) | | | Biology: | Alexander, 1919e (as xanthoptera Meigen) (habitat). Yakovlev and Polevoi, 1997 (list of Fungi). | Figure references: | Habitus: Nakamura, 2006b (male); Salmela, 2024 (male) Wing: Podenas et al., 2006; Boardman, 2007; Podenas and Gelhaus, 2007; Mao and Yang, 2010a; Podenas and Byun, 2016; Salmela, 2024 Hypopygium: Podenas et al., 2006; Podenas and Gelhaus, 2007; Mao and Yang, 2010a; Podenas and Byan, 2016; Salmela, 2024 Ovipositor: Podenas and Byun, 2016 Other figures: Salmela, 2024 (thorax)
Immature stages Larva: Podeniene, 2003a; Krivosheina and Krivosheina, 2010; Krivosheina and Krivosheina, 2011c Pupa: Krivosheina and Krivosheina, 2010
| | | Citations | | | Albania Habitat. Kolcsar et al., 2021b: 151 (Beech [Fagus] forest, next to stream). | | Belgium Habitat. Tack et al., 2021: 141 (loc(s) Antwerpen, alle taxa biodiversiteit inventarisatie Bos t Ename, op paddestoelen op/in hout (in Dutch), see paper for description). | | Bulgaria Habitat. Hubenov, 2025b: suppl: 7 (annotated list Diptera high Bulgarian mts, local distr, habitats and zoogeography). | | Country not relevant Habitat. Krivosheina and Krivosheina, 2011c: 1-294 (larva: loc(s), descr, figs, biology, key) (in Russian, key also in English). | | Czech Rep. Habitat. Sevcik, 2006: 8 (rearing records; a common species with fungicolous and lignicolous larvae, known from many species of fungi, apparently preferring, in contrast to M. quadrimaculata, terrestrial agarics and boletes). | | Denmark Habitat. Nielsen and Nielsen, 2009: 121, 132 (emerging from a forest floor in a Beech [Fagus] stand) (in Danish). | | Finland Habitat. Kahanpaa and Salmela, 2007: 43-45, 48 (found at a small stream with a width of three to five meters, a riverbed of soft muddy sediments, and surrounded by fully grown but not particularly old Spruce [Picea abies] forest, small stands of old Birch [Betula] and Aspen [Populus tremula] are present but there is little dead wood available in the forest) (this is a summary of the available habitat information in Finnish). Habitat. Salmela, 2012c: 4 (strictly fungivorous). | | France Habitat. Kramer and Langlois, 2019a: 83 (on larval fungal habitats in the Ravin de Valbois; see paper for details). Habitat. Quindroit, 2020b: 139 (loc(s) Nord (59) and Pas-de-Calais (62) with notes on habitats and distr) (in French). Habitat. Quindroit, 2021a: 36 (loc(s), resultats d une campagne de piegeage dans le Pas-de-Calais (62); see paper for details on habitat). Habitat. Quindroit and Lemoine, 2022a: 24 (loc(s) Aisne (02) and Somme (80) with notes on habitats and distr) (in French). Habitat. Quindroit and Racine, 2025: 336 (annotated list with information on abundancy and habitat of the Tipuloidea of the Pays de la Loire region, distr) (in French). | | Great Britain Habitat. Alexander, 2002: 90 (develops in a wide range of fungi, terrestrial as well as wood-decay). Habitat. Boardman, 2007: 86 (woodland; breeds on fungi). Habitat. Chandler, 2010c: 431 (overview of known fungus associations). Habitat. Kramer, 2011j: 23, 25 (a woodland species; larvae in fungi). Habitat. Boardman, 2012b: 2 (found at the edge of the moss by a shelter belt of wet Birch [Betula], Alder [Alnus] and Sallow [Salix] woodland). Habitat. Kramer, 2012h: 3 (Summer field meeting Scottish Highlands: a fungivorous woodland species). Habitat. Bloxham, 2014: 82 (larvae in fungi in open woodland). Habitat. Brighton, 2017c: 29, 40 (overall regional checklist Lancashire and Cheshire (VC58, 59 and 60) with notes on habitats and/or regional occurrence and/or status; see paper for details). Habitat. Kramer and Morris, 2022b: 25 (info on the species, on the distr and map of loc(s) Leicestershire and Rutland (VC55); inhabits woodlands where the larvae feed on fungi). | | Japan Habitat. Sasakawa, 2002: 30 (cultivated shiitake mushrooms). Biology. Sueyoshi et al., 2007: 247-257 (loc(s) Honshu; on host abundance of limoniid flies of Agaricales (gill mushrooms) and Aphyllophorales (non-gilled fungi) and the importance of these flies as phoronts of mites inhabiting these fungal orders; see paper for details on fungal species and forest types). | | Lithuania Habitat. Podeniene, 2003a: 1-295 (larva, descr, figs, habitat) (in Lithuanian). Habitat. Podeniene, 2003b: 11, 23, 24 (larval habitat: uppermost layer of soil in old deciduous forest rich in epiphytes, mushrooms). Habitat. Podeniene et al., 2010: 234, 237 (list of fungi in which larvae develop; the commonest species of crane flies developing in fungi in Lithuania; habitats: broadleaved deciduous forests (24% of samples with larvae), Spruce [Picea] forests (20%), Oak-Hornbeam [Quercus-Carpinus] forests (20%), swamp and bog woodland (16%), Pine [Pinus] forests (12%) and mixed forests (8%)). | | Luxembourg Habitat. Vogtenhuber, 2007: 355 (Larvenentwicklung in Basisiomyceten). | | Mongolia Habitat. Gelhaus and Podenas, 2006: 290, 296 (habitat preference: 100% lotic). Habitat. Yadamsuren et al., 2015: 475 (fungi and soil). | | Norway Habitat. Olsen and Andersen, 2021: 215 (loc(s) Innlandet, in former Hedmark, with habitats specified separately, for habitats see also Jonassen and Andersen, 2020). | | Poland Habitat. Wiedenska, 2007a: 75 (mycophagous, feeding on fungi). | | Romania Habitat. Ujvarosi, 2005a: 246 (fleshy fungi, citing various sources). Habitat. Ujvarosi and Poti, 2006: 256 (habitat in Dupa Lunca marshland: shrubs along rivers and streams). Habitat. Ujvarosi et al., 2011b: 109 (loc(s) Dupa Lanca marshy area near Voslobeni) (in Hungarian, species list with habitat information in English). Habitat. Kolcsar et al., 2013: 72 (loc(s) Cluj-Napoca and surroundings; list of habitats with altitudes). | | Russia Habitat. Krivosheina, N.P., 2008: 781 (the larvae have been reared from fruit bodies of ascomycetes (morels and saddle fungi) as well as polypores, agarics, boletes, and russules (cting various authors); at the same time these larvae typically occur in the old strongly decomposed carpophores of polypores and in the dust at their bases). Habitat. Krivosheina, N.P., 2009a: 125-133 (review Russian xylophilous Limoniidae, mainly inhabiting bark and wood of decomposing trees, with new original data on larval biology) (in Russian). Habitat. Krivosheina and Krivosheina, 2010: 773, 777 (larvae from fruit bodies of various agaric fungi and from Boletus luteus; the paper present further details on larval and pupal development, literature review). Habitat. Lantsov, 2014d: 109-110 (on coniferous forest communities of the Teberda River valley: found in fir-tree reed grass and light coniferous sparse pine-forb communities) (in Russian) (information kindly provided by Vladimir Lantsov). Habitat. Polevoi et al., 2018: supplementary species list: (emerged from fallen Aspens [Populus tremula] in Kivach Nature Reserve, Kareliya). Habitat. Dvorak et al., 2020: 6 (on Diptera collected in beer traps in the RUC: Mordoviya Rep.; collected in Pine [Pinus], floodplain and mixed forests). Habitat. Dvorak et al., 2022a: 134 (on Diptera collected in beer traps in the Volga Region, M. bifasciata being collected in Pine [Pinus], decidous and mixed forests). | | Slovakia Habitat. Sevcik, 2004b: 294-295 (larvae fungicolous and lignicolous, known from many species of fungi, Russula nigricans being a new host record). | | South Korea Habitat. Podenas and Byun, 2016: 334 (adults are flying in different types ofdamp forests, in parks, shrubs and gardens; larvaedevelop in different species of mushrooms and could be found in large quantities). | | Ukraine Habitat. Dvorakova et al., 2020: 14-15 (on Diptera collected in beer traps in the western Ukraine; collected in a deciduous forest). | | | | China Revision. Mao and Yang, 2010a: 2-5 (descr, key Chinese Metalimnobia, figs). Description. Zhang, X. et al., 2017b: 72-74 (short descr, figs) (in Chinese). Characters. Jiang and Zhang, 2020: 99, 103 (diagnosis, key, comparison, figs). | | Country not relevant Phylogeny. Boldgiv, 2006: 96-154 (one of the 84 morphological taxa from northern Mongolia in a consistency test of morphology based species identification of crane flies against DNA-barcoding identification using 536-base-pair sequence profiles of the COI gene, also known as the DNA barcode). Phylogeny. Ahonen, 2007: 1-28 (included in phylogenetic analysis of Tipuloidea subfamilies relationships based on molecular data). Immatures. Krivosheina and Krivosheina, 2011c: 1-294 (larva: loc(s), descr, figs, biology, key) (in Russian, key also in English). Description. Podenas and Byun, 2016: 334-339, 343, 345 (short descr, key, comparison, figs). | | Great Britain Characters. Boardman, 2007: 86 (characters, figs). Characters. Kramer and Morris, 2022b: 25 (diagnosis). | | Japan Characters. Sasakawa, 2002: 30 (characters, new synonymy of Japanese taxa). | | Lithuania Immatures. Podeniene, 2003a: 1-295 (larva, descr, figs, habitat) (in Lithuanian). Immatures. Podeniene et al., 2010: 237-240 (key to crane fly larvae developing in mushrooms, figs). | | Mongolia Key. Podenas and Gelhaus, 2007: 73-74 (key, figs). | | Russia Immatures. Krivosheina and Krivosheina, 2010: 773-777 (descr, key, figs, larva, pupa; pupae from Kunahir show some differences and are considered to belong to the subsp. bifasciata avis [according to Sasakawa, 2002, a synonym of bifasciata]). | | | | Albania First record. Kolcsar et al., 2021b: 151 (loc(s)). | | Austria Distribution. Vogtenhuber and Kofler, 2017: 140 (loc(s) Osttirol and Karnten). | | Belarus First record. Kolcsar et al., 2021b: 151 (loc(s) Minsk region). | | Belgium Distribution. Tack et al., 2021: 141 (loc(s) Antwerpen, alle taxa biodiversiteit inventarisatie Bos t Ename, op paddestoelen op/in hout (in Dutch), see paper for description). | | Bulgaria Distribution. Hubenov, 2021a: 38 (listed for Bulgaria, without data, references). Distribution. Hubenov, 2025b: suppl: 7 (annotated list Diptera high Bulgarian mts, local distr, habitats and zoogeography). | | China First record. Mao and Yang, 2010a: 2-5 (loc(s) in 10 Chinese provinces incl. Oriental Guizhou). Distribution. Zhang, X. et al., 2017b: 74 (loc(s) Shaanxi, distr Chinese provinces) (in Chinese). Distribution. Jiang and Zhang, 2020: 99-100 (loc(s) Sichuan, distr). Distribution. Provinces Eastpalaearctic part, 2024: (known from Beijing, Hebei, Heilongjiang, Hubei, Jilin, Liaoning, Ningxia, Shaanxi, Shanxi, Sichuan). | | Country not relevant Distribution. Krivosheina and Krivosheina, 2011c: 1-294 (larva: loc(s), descr, figs, biology, key) (in Russian, key also in English). | | Croatia Distribution. Oosterbroek and Simova-Tosic, 2004: 448 (review literature). | | Czech Rep. Distribution. Stary et al., 2005a: 28 (loc(s)). Distribution. Sevcik, 2006: 8 (loc(s)). Distribution. Stary, 2009i: webpage (checklist). | | Denmark Distribution. Petersen and Jong, 2001b: 150 (checklist). Distribution. Nielsen and Nielsen, 2009: 121 (loc(s)). | | Estonia Distribution. Stary, 2004f: (Fauna Europaea: added Estonia). | | Finland Distribution. Polevoi, 2001: 191 (loc(s)). Distribution. Salmela, 2001a: 144, 151 (loc(s)). Distribution. Mukkala et al., 2005: 16 (among list of insects collected in South Ostrobothnia, Ilmajoki during 2001-2004). Distribution. Kahanpaa and Salmela, 2007: 48 (loc(s)). Distribution. Salmela et al., 2007b: 46 (loc(s)). Distribution. Salmela, 2010a: 12 (survey fauna Tahmela spring in city of Tampere) (in Finnish). Distribution. Salmela, 2012b: 238 (annotated list of Finnish crane flies). Distribution. Salmela and Petrasiunas, 2014: 31 (checklist Finnish Tipulomorpha). Distribution. Salmela et al., 2015: 103 (list of localities in extensive report on malaise-trapping of insects in conservation areas in Lapland in 2012–2014) (in Finnish). | | France Distribution. Kramer and Langlois, 2019a: 76 (record(s) Ravin de Valbois, Doubs (25)). Distribution. Quindroit, 2020b: 139 (loc(s) Nord (59) and Pas-de-Calais (62) with notes on habitats and distr) (in French). Distribution. Quindroit, 2021a: 36 (loc(s), resultats d une campagne de piegeage dans le Pas-de-Calais (62); see paper for details on habitat). Distribution. Tissot et al., 2021: 209 (the Diptera of the Reserves Naturelles Nationales du Doubs (25), results of Malaise trapping since 2009) (in French). Distribution. Quindroit and Lemoine, 2022a: 24 (loc(s) Aisne (02) and Somme (80) with notes on habitats and distr) (in French). Distribution. Quindroit and Lemoine, 2022a: 37 (table specifying the presence in the five departments of the Hauts-de-France region). Distribution. Quindroit and Racine, 2025: 336 (annotated list with information on abundancy and habitat of the Tipuloidea of the Pays de la Loire region, distr) (in French). Distribution. Quindroit and Racine, 2025: 360 (table specifying the presence in the five departments of the Pays de la Loire region). | | Germany Distribution. Schacht, 1999: 135 (checklist Bayern). Distribution. Reusch and Oosterbroek, 2000: 162 (checklist German Bundeslander). Distribution. Reusch and Bellstedt, 2001a: 268 (red list status Thuringen). Distribution. Reusch and Bellstedt, 2001b: 75 (checklist Thuringen). Distribution. Schacht, 2005: 5 (checklist Bayern). Distribution. Hable et al., 2010b: 91 (loc(s) Bayern). Distribution. Schacht, 2010: 17 (checklist Diptera Bayern). Distribution. Dunk and Kraus, 2014: 169 (loc(s) Bayern). Distribution. Stuke, 2019: 144 (checklist Niedersachsen and Bremen with reference to original sources). | | Great Britain Distribution. Boardman, 2007: 86 (loc(s), map). Distribution. Skidmore, 2009: 129 (review of occurrence on the Western Isles of Scotland, distr). Distribution. Kramer, 2011j: 23 (loc(s) Leicestershire and Rutland (VC55)). Distribution. Boardman, 2012b: 2 (loc(s) Shropshire). Distribution. Kramer, 2012h: 3 (loc(s) Summer field meeting Scottish Highlands). Distribution. Kramer, 2013i: 8 (checklist of species from Leigh Woods NNR, Bristol district). Distribution. Bloxham, 2014: 82 (loc(s) West Bromwich). Distribution. Hewitt, 2014: (provisional checklist Cumbrian Diptera). Distribution. Wolton et al., 2014: 27 (among Diptera associated with a hedge in Devon). Distribution. Brighton, 2017c: 29, 40 (overall regional checklist Lancashire and Cheshire (VC58, 59 and 60) with notes on habitats and/or regional occurrence and/or status; see paper for details). Distribution. Cunningham, 2020: 24 (loc(s) Stover Country PArk, Devon). Distribution. Kramer and Morris, 2022b: 25 (info on the species, on the distr and map of loc(s) Leicestershire and Rutland (VC55); inhabits woodlands where the larvae feed on fungi). | | Ireland Distribution. Chandler et al., 2008: 14 (checklist). | | Italy Distribution. Podenas and Podeniene, 2008: 351-352 (annotated checklist South Tyrol [Prov. Bolzano]). | | Japan First record. Sasakawa, 2002: 30 (new Japanese synonyms). Distribution. Nakamura, 2002: 1, 169 (loc(s) Honshu). Distribution. Sueyoshi et al., 2007: 247-257 (loc(s) Honshu; on host abundance of limoniid flies of Agaricales (gill mushrooms) and Aphyllophorales (non-gilled fungi) and the importance of these flies as phoronts of mites inhabiting these fungal orders; see paper for details on fungal species and forest types). Distribution. Kato, 2023a: 30 (loc(s) Honshu) (in Japanese). Distribution. Kato and Yamauchi, 2024: 28 (loc(s) Toyama pref., Honshu). | | Kazakhstan First record. Devyatkov, 2020a: 53 (loc(s) East Kazakhstan, distr) (in Russian). | | Latvia First record. Stary, 2004f: (Fauna Europaea: added ?Latvia). Distribution. Kolcsar et al., 2021b: 151-152 (loc(s); confirmation for Latvia). | | Lithuania Distribution. Pakalniskis et al., 2006: 22 (checklist). Distribution. Podeniene et al., 2010: 234 (loc(s), distr). | | Luxembourg First record. Vogtenhuber, 2007: 354 (loc(s)). | | Mongolia Distribution. Boldgiv, 2006: 153 (loc(s)). Distribution. Gelhaus and Podenas, 2006: 290, 296 (loc(s)). Distribution. Yadamsuren et al., 2015: 475 (loc(s) Mongolia). | | Netherlands Distribution. Jong and Oosterbroek, 2002b: 34 (checklist). | | North Korea First record. Podenas and Byun, 2016: 334-335, 344 (loc(s), map). | | Norway Distribution. Olsen et al., 2018: 162-163 (annotated checklist Nordic countries and for Norway according to the Strand regions). Distribution. Olsen and Andersen, 2021: 215 (loc(s) Innlandet, in former Hedmark, with habitats specified separately, for habitats see also Jonassen and Andersen, 2020). | | Poland Distribution. Wiedenska, 2007a: 75 (checklist). Distribution. Wiedenska, 2010: 134 (loc(s) Parku Krajobrazowego Wzniesien Lodzkich, Lodz). | | Romania Distribution. Ujvarosi and Poti, 2006: 256 (loc(s) eastern Carpathians). Distribution. Ujvarosi, 2007: 228-231 (checklist Limoniidae). Distribution. Ujvarosi et al., 2011b: 109 (loc(s) Dupa Lanca marshy area near Voslobeni) (in Hungarian, species list with habitat information in English). Distribution. Kolcsar et al., 2013: 72 (loc(s) Cluj-Napoca and surroundings; list of habitats with altitudes). | | Russia Distribution. Lantsov, 2003c: 104 (loc(s) Caucasus region, distr). Distribution. Lantsov, 2003e: 85 (loc(s), distr Caucasus region). Distribution. Polevoi and Humala, 2005: 182 (RUN: list of Diptera of Vepsskaya Volost, Kareliya) (in Russian). Distribution. Paramonov, 2006a: 152 (loc(s) RUC: Ulyanovskaya oblast, distr). Distribution. Pilipenko and Sidorenko, 2006b: 150 (checklist Kedrovaya Pad, distr) (in Russian). Distribution. Humala and Polevoi, 2008: 133 (loc(s) RUN: Kareliya). Distribution. Pilipenko, 2009c: 335 (loc(s) Lazovsky Nature Reserve, Primorskiy kray). Distribution. Krivosheina and Krivosheina, 2010: 773 (loc(s) RUC: Moscow prov., Kunashir). Distribution. Gavryushin in litt., 2011e: (loc(s) RUN: Murmanskaya oblast). Distribution. Gavryushin in litt., 2011f: (loc(s) RUN: Arkhangelskaya oblast). Distribution. Gavryushin in litt., 2012e: (loc(s) RUE: Bashkortostan Rep.). Distribution. Humala and Polevoi, 2012: 143 (loc(s) RUN: Kareliya) (in Russian). Distribution. Gavryushin in litt., 2013b: (loc(s) Shikotan, Kuril Is). Distribution. Jakovlev et al., 2014: 310 (loc(s) RUN: Kareliya). Distribution. Paramonov, 2014: 39-43 (revision coll. Eversmann, loc(s) RUE: Orenburgskaya oblast) (in Russian). Distribution. Gavryushin in litt., 2015a: (loc(s) RUE: Bashkortostan Rep.). Distribution. Paramonov, 2015: 69 (loc(s) RUS: Saratovskaya oblast) (in Russian). Distribution. Ruchin and Pilipenko, 2015: 58 (loc(s), checklist RUC: Mordoviya Rep.). Distribution. Lantsov, 2016: 165-170 (on craneflies from Arkhyz, Karachay-Cherkessia Rep., North Caucasus) (in Russian). Distribution. Paramonov and Pilipenko, 2016: 100 (loc(s) RUC: Tverskaya oblast) (in Russian). Distribution. Paramonov, 2016: 130 (loc(s) RUC: Chuvash Rep.) (in Russian). Distribution. Paramonov, 2017a: 172 (loc(s) RUC: Chuvash Rep.) (in Russian). Distribution. Paramonov, 2018a: 184 (loc(s) RUC: Chuvash Rep., distr) (in Russian). Distribution. Polevoi et al., 2018: supplementary species list: (emerged from fallen Aspens [Populus tremula] in Kivach Nature Reserve, Kareliya). Distribution. Dvorak et al., 2020: 6 (on Diptera collected in beer traps in the RUC: Mordoviya Rep.; collected in Pine [Pinus], floodplain and mixed forests). Distribution. Pilipenko et al., 2020: 361 (loc(s) RUC: Mordoviya Rep., distr). Distribution. Dvorak et al., 2022a: 134 (on Diptera collected in beer traps in the Volga Region, M. bifasciata being collected in Pine [Pinus], decidous and mixed forests). Distribution. Humala and Polevoi, 2022: 33 (loc(s) RUN: Arkhangelskaya oblast). Distribution. Solodkiy and Sergeev, 2023: 171-172 (loc(s) Primorskiy kray) (in Russian). Distribution. Devyatkov and Dubatolov, 2025: 35-36 (loc(s) Khabarovskiy kray, Bolshekhekhtsirsky Nature Reserve, distr) (in Russian). | | Serbia Distribution. Oosterbroek and Simova-Tosic, 2004: 448 (review literature). | | Slovakia Distribution. Sevcik, 2004b: 294-295 (loc(s), a common species). Distribution. Sevcik, 2006: 8 (loc(s)). Distribution. Stary, 2009d: 37 (loc(s) Polana area). Distribution. Stary, 2009i: webpage (checklist). | | Slovenia Distribution. Oosterbroek and Simova-Tosic, 2004: 448 (review literature). | | South Korea First record. Podenas and Byun, 2016: 335, 344 (loc(s), map). | | Spain First record. Kolcsar et al., 2021b: 152 (loc(s) Lugo). | | Sweden Distribution. Fritz and Lindstrom, 2013: 62 (loc(s) Hallands County) (in Schwedish). | | Switzerland Distribution. Podenas et al., 2006: 175, 284 (map, vertical distr per thermic level). Distribution. Dufour and Merz, 2012: 355, 356 (annotated checklist canton Geneva, with note on specimen). | | Turkey First record. Koc et al., 2005b: 616-618 (loc(s), map, checklist). Distribution. Koc et al., 2016: 12 (provinces Marmara region). | | Ukraine Distribution. Dvorakova et al., 2020: 14-15 (on Diptera collected in beer traps in the western Ukraine; collected in a deciduous forest). | | | | Albania Kolcsar et al., 2021b: month(s): 7. | | Austria Vogtenhuber and Kofler, 2017: month(s): 8-9. | | Belarus Kolcsar et al., 2021b: month(s): 7. | | China Mao and Yang, 2010a: month(s): 5-8. Zhang, X. et al., 2017b: month(s): 8. Jiang and Zhang, 2020: month(s): 7. | | Czech Rep. Stary et al., 2005a: month(s): 6-10. Sevcik, 2006: month(s): 7-10. | | Finland Salmela, 2001a: month(s): 7-9. | | France Quindroit, 2020b: month(s): 9, 11. Quindroit and Lemoine, 2022a: month(s): 7, 10. | | Great Britain Kramer, 2011j: month(s): 5, 7. Boardman, 2012b: month(s): 6. Kramer, 2012h: month(s): 7. Cunningham, 2020: month(s): 9. Kramer and Morris, 2022b: month(s): 7-10. | | Japan Nakamura, 2002: month(s): 8-9. Kato, 2023a: month(s): 7, 9. Kato and Yamauchi, 2024: month(s): 7-9. | | Kazakhstan Devyatkov, 2020a: month(s): 6-8. | | Latvia Kolcsar et al., 2021b: month(s): 7. | | Lithuania Podeniene et al., 2010: month(s): 5-9. | | Mongolia Gelhaus and Podenas, 2006: month(s): 7. | | North Korea Podenas and Byun, 2016: month(s): 6-8. | | Norway Olsen and Andersen, 2021: month(s): 6-9. | | Romania Ujvarosi et al., 2011b: month(s): 6. Kolcsar et al., 2013: month(s): 6. | | Russia Lantsov, 2003c: month(s): 7, 9. Lantsov, 2003e: month(s): 7, 9. Paramonov, 2006a: month(s): 7. Pilipenko, 2009c: month(s): 7-9. Gavryushin in litt., 2011e: month(s): 7. Gavryushin in litt., 2011f: month(s): 8. Gavryushin in litt., 2012e: month(s): 8. Gavryushin in litt., 2013b: month(s): 8-9. Gavryushin in litt., 2015a: month(s): 7. Paramonov, 2015: month(s): 8. Ruchin and Pilipenko, 2015: month(s): 5, 8-9. Paramonov and Pilipenko, 2016: month(s): 6, 8-10. Paramonov, 2016: month(s): 7-9. Paramonov, 2017a: month(s): 7. Paramonov, 2018a: month(s): 7-8. Dvorak et al., 2020: month(s): 8-9. Pilipenko et al., 2020: month(s): 6-9. Dvorak et al., 2022a: month(s): 6-9(10). Solodkiy and Sergeev, 2023: month(s): 6, 8. Devyatkov and Dubatolov, 2025: month(s): 6-9. | | Slovakia Sevcik, 2006: month(s): 7-10. Stary, 2009d: month(s): 9. | | South Korea Podenas and Byun, 2016: month(s): 7-10. | | Spain Kolcsar et al., 2021b: month(s): 9. | | Switzerland Podenas et al., 2006: month(s): 5-10. | | Turkey Koc et al., 2005b: month(s): 6. | | Ukraine Dvorakova et al., 2020: month(s): 7. | | | | Albania Kolcsar et al., 2021b: altitude: 690 m. | | Austria Vogtenhuber and Kofler, 2017: altitude: 610-1050 m. | | Belarus Kolcsar et al., 2021b: altitude: 690 m. | | China Mao and Yang, 2010a: altitude: 300-2020 m. Jiang and Zhang, 2020: altitude: 2673 m. | | Japan Kato and Yamauchi, 2024: altitude: 1120 m. | | Kazakhstan Devyatkov, 2020a: altitude: 280-1468 m. | | Latvia Kolcsar et al., 2021b: altitude: 690 m. | | Mongolia Gelhaus and Podenas, 2006: altitude: 1790 m. | | North Korea Podenas and Byun, 2016: altitude: 183-1981 m. | | Romania Ujvarosi and Poti, 2006: altitude: 670 m. | | Russia Lantsov, 2003c: altitude: 750-1600 m. Lantsov, 2003e: altitude: 750-1600 m. Gavryushin in litt., 2011f: altitude: 19 m. Gavryushin in litt., 2012e: altitude: 552-607 m. Lantsov, 2014d: altitude: 1586-1883 m. Gavryushin in litt., 2015a: altitude: 607 m. Paramonov, 2015: altitude: 302 m. Devyatkov and Dubatolov, 2025: altitude: 70-100 m. | | South Korea Podenas and Byun, 2016: altitude: 89-885 m. | | Spain Kolcsar et al., 2021b: altitude: 386 m. | | Turkey Koc et al., 2005b: altitude: 100-670 m. |
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