Details for Ctenophora (Cnemoncosis) ornata

Name: Ctenophora (Cnemoncosis) ornata Meigen and Wiedemann, 1818 Publication: In: Meigen, 1818, Syst. Beschr. 1: 166 (as Ctenophora) Status: Recognized taxon. Classification: Family Tipulidae Subfamily Ctenophorinae Keys: Bygebjerg and Munk, 2005 (Danish Ctenophorinae (in Danish); Oosterbroek et al., 2006 (Westpalaearctic Ctenophorinae); Peeters en Oosterbroek, 2013e (craneflies Netherlands, Belgium and Luxembourg) (in Dutch); Men and Huang, 2014 (World species Ctenophora); Stubbs and Kramer, 2016c; Stubbs, 2021 (British craneflies) habitus male habitus male habitus male habitus male habitus male plus 23 more images of habitus, habitus, habitus, habitus, habitus, habitus, habitus, body part(s), body part(s), body part(s), body part(s), hypopygium, hypopygium, ovipositor, ovipositor, wing, wing, pupa, habitat, habitat, habitat, habitat and habitat. Distribution: Albania, Austria, Belgium, Bulgaria, Croatia, Czech Rep., Denmark, France, Germany, Great Britain, Greece (incl. Andros, Kerkyra [Corfu]), Hungary, Ireland, Italy (north, Roma), Liechtenstein, Luxembourg, Netherlands, North Macedonia, Poland, Portugal, Romania, Serbia, Slovakia, Slovenia, Spain, Sweden (south), Switzerland, Turkey (European part: Kirklareli), Ukraine; Russia: North Caucasus; Turkey (Asiatic part: Icel), Cyprus. Note: see the manual for abbreviations: present-day Russia. Region(s): Westpalaearctic General note: Added: Ireland (Speight and Nash, 1993). For Spain check Eiroa and Carles-Tolra, 2019, and citations below. For Portugal check Oosterbroek et al., 2020, and citations below. For Great Britain and Ireland check Stubbs, 2021, and citations below. Map: Sorensen, 2002 (Denmark); Ashe et al., 2007a (Ireland); Bakowski et al., 2011 (Poland); Kramer, 2013h (Great Britain); Pilipenko and Lantsov, 2017 (Krasnodarskiy kay); Oosterbroek, 2022a (Netherlands, 2012-2121); Kania et al., 2023 (Poland); Jentzsch et al., 2024 (Sachsen) Figure references: Habitus: Albertini, 2003 (male); Oosterbroek et al., 2006 (male); Camano Portela et al., 2010 (male); Bakowski et al., 2011 (male); Brock, 2011 (male); Kramer, 2012l (male); Oosterbroek, 2012 (male, female); Kramer, 2013h (male, female); Aistleitner, 2015 (male); Kowalczyk and Senn, 2017 (female); Pilipenko and Lantsov, 2017 (male); Pino Perez and Pino Perez, 2016 (male); Kohler and Niermann, 2021 (male); Boardman and Owens, 2021 (male); Wiebel and Kessler, 2021 (male); Kolcsar et al., 2023a (male); Kania et al., 2023 (male, female); ; Jentzsch et al., 2024 (male); Quindroit and Lemoine, 2022a (male); Guidetti, 2024 (male) Head: Sorensen, 2002 (antenna); Oosterbroek et al., 2006 (antenna); Malkiewicz et al., 2012 (female head and antenna) Wing: Dominiak, 2011 Hypopygium: Oosterbroek et al., 2006; Dominiak, 2011 Other figures: Oosterbroek et al., 2006 (abdomen male, female); Dominiak, 2011 (thorax, abdomen); Malkiewicz et al., 2012 (abdomen Immature stages Larva: Sorensen, 2002 Pupa: Habda, 2010 (hatching female) Miscellaneous Camano Portela et al., 2010 (habitat); Pino Perez and Pino Perez, 2016 (habitat) Citations on biology (mainly from 2000 onward): Country not relevant Biology. Dajoz, 2000: 475 (larvae live in wood invded by white rot fungi and for optimal growth require wood with a water content equal to 240% of dry weight). Cyprus Habitat. Aristophanous in litt., 2021: (collected at night in a riparian vegetation, see photos above of male and habitat, recorded as new for Cyprus in Kolcsar et al., 2023a). Denmark Biology. Sorensen, 2002: 129-135 (habitat: occurs in decaying wood and is also found under bark of fallen logs and old stems; biology; conservation status; review literature) (in Danish). France Habitat. Quindroit and Lemoine, 2022a: 11 (loc(s) Aisne (02) and Oise (60) with notes on habitats and distr) (in French). Germany Habitat. Heiss et al., 2017: 159-160, 180-182 (overview of collecting methods and habitats). Habitat. Heiss, 2017a: 562, 567, 570 (loc(s) Dinkelberg, Baden-Wurttemberg and table with details on presence in the various biotop-groups). Habitat. Kohler and Niermann, 2020: 325 (Abstract: collected in a light trap hung at 1.5 m height on the bushy margin of an orchard meadow with partly dead trees; see paper for details, habitat figs). Great Britain Habitat. Alexander, 2002: 88 (reared from porridge-like wet wood mould in standing or fallen Fagus trees). Habitat. Skidmore, 2003: 9, 22 (from hollow Fagus). Habitat. Stubbs, 2003: 68-70, 77 (associated with woodland and wood pasture where large trees with rot holes are present, especially Fagus; larvae have been found in porridge-texture wet rotten wood mould at the base of a large hollow Fagus and in similar material in a fallen Fagus trunk only c. 0.6 m thick; (adults fly at some speed through shade of old woodland during daylight hours, where its curious, tumbling flight makes it extremely difficult to capture, citing Menier, 1973); the paper reviews recent literature and includes further details on subhabitat preferences and monitoring techniques). Biology. Godfrey, 2008: 3 (males were collected using a mercury vapour lamp, females don t seem to be attracted to light). Habitat. Waring, 2013: 58 (loc(s) new Forest, Hampshire; in a Robinson trap in open heathland). Biology. Leftwich, 2018: 4 (descr New Forest habitat and on four males being attract to light between 2300 hours and midnight). Luxembourg Habitat. Vogtenhuber, 2007: 352 (Larven in Mulm hohler Buchen [Fagus]). Netherlands Biology. Oosterbroek and Jong, 2001: 101-102 (sexratio; habitat: ancient broad-leaved forest with dead wood and old or decaying trees; larvae live in soft wood-mould; the species has been reared from a variety of trees: Apple [Malus], Beech [Fagus], Horse Chestnut [Aesculus], Elm [Ulmus], Maple [Acer], Plane [Platanus] and Sweet Chestnut [Castanea], citing various sources). Poland Habitat. Hebda, 2010: 101-104 (reared from pupa found in decaying Sycamore [Acer pseudoplatanus], photo hatching female). Habitat. Malkiewicz et al., 2012: 120 (belongs to the saproxylic fauna, whose presence is connected with old forests and dying trees; all Ctenophora species are rare in Poland and most of them are listed in the Red List of Threatened Animals in Poland). Romania Habitat. Kolcsar et al., 2013: 73, 75 (loc(s) Cluj-Napoca and surroundings; list of habitats with altitudes). Russia Habitat. Pilipenko and Lantsov, 2017: 415 (on the red data status in North Caucasus: Krasnodarskiy kray, distr, map) (in Russian). Spain Habitat. Camano Portela et al., 2011: 103-105 (collected at a small open area surrounded by orocantabric bushes and the sintaxon Linario triornithophorae-Querceto pyrenaicae, ecological information) (in Spanish). Habitat. Pino Perez and Pino Perez, 2014: 167-170 (first record for Orense with habitat information) (in Spanish). Habitat. Pino Perez and Pino Perez, 2016: 1-5 (first record for Pontevedra with habitat information) (in Spanish). on characters and taxonomy (mainly from 2000 onward): Country not relevant Key. Oosterbroek et al., 2006: 139-146 (key, figs, review literature). Characters. Bakowski et al., 2011: 29-32 (characters, comparison, photo male). Characters. Malkiewicz et al., 2012: 116-120 (illustrated comparison table with festiva) (in Polish). Denmark Immatures. Sorensen, 2002: 129-135 (descr, figs, review literature) (in Danish). Key. Bygebjerg and Munk, 2005: 42 (in Danish). Great Britain Figures. Kramer, 2013h: 3-4 (male, female). Portugal Characters. Ferreira et al., 2021: 11 (barcoding based on material mentioned in Oosterbroek et al. 2020). Russia Description. Pilipenko and Lantsov, 2017: 415 (descr, fig.) (in Russian). on distribution (mainly from 2000 onward): Austria Distribution. Vogtenhuber, 2011: 44 (checklist Austria, list of provinces). Distribution. Hellrigl, 2012: 119 (loc(s) Sudtirol). Distribution. Vogtenhuber and Kofler, 2017: 120 (loc(s) Osttirol, distr in Austria). Bulgaria Distribution. Bechev, 2009: 2 (Sredna Gora, citing Nedelkov, 1912). Distribution. Hubenov, 2021a: 26 (checklist with summary of distr in Bulgaria, references). Country not relevant Distribution. Oosterbroek et al., 2006: 139-146 (distr). Croatia Distribution. Simova-Tosic and Oosterbroek, 2003: 43 (review literature). Distribution. Kolcsar et al., 2023b: 33 (loc(s)). Cyprus First record. Kolcsar et al., 2023a: 15 (loc(s) Tsakistra, see also photo above by M. Aristophanous). Czech Rep. Distribution. Martinovsky and Bartak, 2005: 245 (red list status Czech Rep.: endangered). Distribution. Stary, 2009k: webpage (checklist). Denmark First record. Petersen et al., 2001b: 162 (checklist). Distribution. Sorensen, 2002: 129-135 (loc(s), map) (in Danish). France Distribution. Tillier and Dehalleux, 2019: 145-146 (new and published records Ile-de-France, loc(s) Paris (75), Seine-et-Marne (77), Yvelines (78), Essonne (91), Hauts-de-Seine (92), Val-de-Marne (94) and Val-d’Oise (95)). Distribution. Quindroit, 2020a: 9, 12, 45-48 (annotated list of the Tipuloidea of Pays de la Loire) (in French). Distribution. Tissot et al., 2021: 217 (the Diptera of the Reserves Naturelles Nationales du Doubs (25), results of Malaise trapping since 2009) (in French). Distribution. Quindroit and Lemoine, 2022a: 11 (loc(s) Aisne (02) and Oise (60) with notes on habitats and distr) (in French). Distribution. Quindroit and Lemoine, 2022a: 34 (table specifying the presence in the five departments of the Hauts-de-France region). Distribution. Lemoine and Quindroit, 2023: 5 (loc(s) Gironde (33)). Distribution. Tillier et al., 2023: 169 (loc(s) Doubs (25), with list of altitude(s) and habitat(s)) (in French). Distribution. Oosterbroek in litt., 2024: (loc(s) Vaucluse (84)). Distribution. Quindroit and Racine, 2025: 356 (table specifying the presence in the five departments of the Pays de la Loire region). Germany Distribution. Schacht et al., 2001: 423 (loc(s) Bayern). Distribution. Heiss, 2003: 12 (checklist Thuringen). Distribution. Schacht, 2005: 3 (checklist Bayern). Distribution. Hable et al., 2010a: 80, 82 (loc(s) Bayern). Distribution. Schacht, 2010: 38 (checklist Diptera Bayern). Distribution. Dunk and Kraus, 2014: 168 (loc(s) Bayern). Distribution. Heiss et al., 2017: 159-160 (loc(s) Thuringen, review distr). Distribution. Heiss, 2017a: 562, 567, 570 (loc(s) Dinkelberg, Baden-Wurttemberg and table with details on presence in the various biotop-groups). Distribution. Heiss, 2017b: 21 (checklist Tipulidae Thuringen). Distribution. Kohler and Niermann, 2020: 328 (loc(s) Thuringen). Distribution. Heiss and Malec, 2023: 328 (annotated checklist Hessen). Distribution. Guidetti, 2024: 1-3 (on two new records from Leipzig, Sachsen). Distribution. Jentzsch et al., 2024: 2-3, 8 (loc(s) Sachsen and one older record for Sachsen-Anhalt, map). Distribution. Stuke et al., 2025a: 51 (loc(s) Niedersachsen). Great Britain Distribution. Green, 2001: 54-55 (loc(s) New Forest, Hamshire, review British records, citing Falk, 1991, with additional New Forest localities; the species is restricted to ancient broad-leaved woodland and has been reared from wet ‘porridge’ wood mould in a Beech [Fagus sylvatica]; old trees and stumps of particularly large girth are considered to be probably essential for this species; adults have been recorded from june to august and have been taken at moth traps at night and in the vicinity of old, living Oaks [Quercus] spp.). Distribution. Alexander, 2002: 88 (distr). Distribution. Albertini, 2003: 177 (a male found at Langley Park, near Slough, Bucks, VC 24, fig). Distribution. Skidmore, 2003: 9, 22 (loc(s) Windsor Park). Distribution. Stubbs, 2003: 69 (distr in GB). Distribution. Godfrey, 2008: 3 (loc(s) New Forest; see also Brock, 2011). Distribution. Whiteley, 2010: 3 (loc(s) Derbyshire and/or Nottinghamshire). Distribution. Dickson, 2011: 161 (loc(s) New Forest, South Hamsphire). Distribution. Dickson, 2011: 161 (loc(s) South Hampshire). Distribution. Kramer, 2012i: 5 (on the surprising presence of this species in the Wingate collection because the species has never been recorded north of Sherwood Forest; same, with colourphoto of male, in Kramer, 2012l). Distribution. Kramer, 2013h: 3-4, 11 (note on dispersal and increase in frequency at various loc(s), distr, map GB). Distribution. Waring, 2013: 58 (loc(s) new Forest, Hampshire; in a Robinson trap in open heathland). Distribution. Sims, 2014a: 76 (loc(s) Berkshire, distr) (same record in Sims, 2014b). Distribution. Finch, 2017: 100 (loc(s) Berkshire, distr). Distribution. Miles, 2017: 95 (loc(s) Setley Common, New Forest). Distribution. Leftwich, 2018: 4 (loc(s) New Forest). Distribution. Boardman and Owens, 2021: 4 (loc(s) South Wiltshire). Ireland Distribution. Ashe et al., 2007a: 299-303, 343, 351 (review single Irish record, red data book status, map). Distribution. Chandler et al., 2008: 4 (checklist). Italy Distribution. Oosterbroek, 2008c: 340 (annotated checklist South Tyrol [Prov. Bolzano]). Distribution. Rivosecchi and Maltzeff, 2011: 16 (loc(s) Roma). Liechtenstein First record. Aistleitner, 2015: 92, 99 (loc(s), distr). Luxembourg Distribution. Vogtenhuber, 2007: 351, 352 (loc(s), distr). Netherlands Distribution. Oosterbroek and Jong, 2001: 101-102 (review Dutch records). Distribution. Jong and Oosterbroek, 2002d: 39 (checklist). Distribution. Oosterbroek, 2012: 10-15 (distr in The Netherlands) (in Dutch). Distribution. Oosterbroek, 2022a: 6-13 (update on recent records (2012-2021) of Dutch Ctenophorinae) (in Dutch). North Macedonia Distribution. Simova-Tosic and Oosterbroek, 2003: 43 (review literature). Poland Distribution. Palaczyk et al., 2002: 40 (in red list of threatened Diptera in Poland) (in Polish). First record. Skibinska and Chudzicke, 2007b: 77 (checklist, vulnerable). Distribution. Hebda, 2010: 101-104 (loc(s), review of Polish records). Distribution. Bakowski et al., 2011: 29-32 (loc(s), review of Polish records, review red-list status). Distribution. Dominiak, 2011: 2-4 (loc(s), distr) (in Polish). Distribution. Malkiewicz and Nalewalska, 2011: 269-270 (loc(s), review Polish records) (in Polish). Distribution. Kowalczyk and Senn, 2017: 4 (loc(s) Gdynia, with references for the species from Poland) (in Polish). Distribution. Kania et al., 2023: 177-179 (loc(s), distr, review references, map) (in Polish). Portugal First record. Grosso-Silva and Soares-Vieira, 2011: 67 (loc(s) Viano do Castelo: Penede-Geres NP). Distribution. Oosterbroek et al., 2020: 337 (loc(s) Braganca and Guarda, also known from Viana do Castelo, distr). Romania Distribution. Ujvarosi, 2003: 56 (in Romania only 19th century records). Distribution. Ujvarosi, 2007: 232-233 (checklist Tipulidae). Distribution. Parvu, 2009: 448 (on some Diptera from Bucuresti and its surroundings). Distribution. Kolcsar et al., 2013: 73, 75 (loc(s) Cluj-Napoca and surroundings; list of habitats with altitudes). Distribution. Torok et al., 2014: 17 (checklist Romania). Russia First record. Pilipenko, 2016: 275-277 (loc(s) North Caucasus: Krasnodarskiy kray) (in Russian). Distribution. Pilipenko and Lantsov, 2017: 415 (on the red data status in North Caucasus: Krasnodarskiy kray, distr, map) (in Russian). Distribution. Lantsov, 2019a: 54-55 (in Caucasus region only known from Krasnodarskiy kray) (in Russian, English translation in Lantsov, 2019b). Distribution. Lantsov, 2019c: 134 (North Caucasus: on tipuloid dipterans of the protected areas of Dagestan) (in Russian). Serbia Distribution. Simova-Tosic and Oosterbroek, 2003: 43 (review literature). Slovakia Distribution. Stary, 2009k: webpage (checklist). Slovenia Distribution. Simova-Tosic and Oosterbroek, 2003: 43 (review literature). Spain Distribution. Eiroa and Baez, 2002b: 79 (checklist). Distribution. Hancock in litt., 2006: (having reared Ctenophora ornata from Cabaneros Reserve, south west of Toledo). Distribution. Carles-Tolra, 2010: 56 (loc(s) various provinces). Distribution. Camano Portela et al., 2011: 103-105 (loc(s) Lugo). Distribution. Pino Perez and Pino Perez, 2014: 167-170 (first record for Orense with habitat information) (in Spanish). Distribution. Robledo, 2014: 264 (loc(s) Sierra de Guadarrama). Distribution. Pino Perez and Pino Perez, 2016: 1-5 (first record for Pontevedra with habitat information) (in Spanish). Distribution. Eiroa and Carles-Tolra, 2019: 132 (loc(s) Leon, list of Spanish provinces, distr) (in Spanish). Sweden First record. Bartsch et al., 2005: 331 (red list status Sweden, distr). Distribution. Cederberg et al., 2010: webpage (red list status Sweden, map, distr). Switzerland Distribution. Dufour and Merz, 2012: 355 (annotated checklist canton Geneva). Distribution. Weibel and Kessler, 2021: 15-16 (loc(s) canton Schaffhausen, conservation status, distr). Turkey Distribution. Koc and Oosterbroek, 2001: 464 (checklist). Distribution. Oosterbroek, 2009c: 188 (new for European part: prov. Kirklareli). on flight period (mainly from 2000 onward): Austria Hellrigl, 2012: month(s): 7. Vogtenhuber and Kofler, 2017: month(s): 7. Croatia Kolcsar et al., 2023b: month(s): 4. Cyprus Kolcsar et al., 2023a: month(s): 6. Denmark Sorensen, 2002: month(s): 5-7. France Tillier and Dehalleux, 2019: month(s): 5-7. Quindroit, 2020a: month(s): 6-7. Quindroit and Lemoine, 2022a: month(s): 5-7. Lemoine and Quindroit, 2023: month(s): 6. Tillier et al., 2023: month(s): 7. Oosterbroek in litt., 2024: month(s): 7. Germany Schacht et al., 2001: month(s): 6. Hable et al., 2010a: month(s): 6. Dunk and Kraus, 2014: month(s): 5-6. Heiss et al., 2017: month(s): 4-7, 9. Heiss, 2017a: month(s): 7. Kohler and Niermann, 2020: month(s): 6-7. Guidetti, 2024: month(s): 5, 7. Jentzsch et al., 2024: month(s): 5-8. Stuke et al., 2025a: month(s): 7. Great Britain Green, 2001: month(s): 6-8. Stubbs, 2003: month(s): 6-8. Godfrey, 2008: month(s): 6-7. Whiteley, 2010: month(s): 7. Dickson, 2011: month(s): 7. Dickson, 2011: month(s): 7. Kramer, 2012i: month(s): 7. Kramer, 2013h: month(s): 6-7. Waring, 2013: month(s): 7. Sims, 2014a: month(s): 7. Finch, 2017: month(s): 6. Miles, 2017: month(s): 7. Leftwich, 2018: month(s): 7. Boardman and Owens, 2021: month(s): 7. Ireland Ashe et al., 2007a: month(s): 5. Liechtenstein Aistleitner, 2015: month(s): 6. Netherlands Oosterbroek and Jong, 2001: month(s): 6-8. Oosterbroek, 2012: month(s): 4-7. Poland Hebda, 2010: month(s): 7. Bakowski et al., 2011: month(s): 5-7. Dominiak, 2011: month(s): 4-5. Malkiewicz and Nalewalska, 2011: month(s): 6-8. Kowalczyk and Senn, 2017: month(s): 8. Kania et al., 2023: month(s): 6-7. Portugal Grosso-Silva and Soares-Vieira, 2011: month(s): 7. Oosterbroek et al., 2020: month(s): 7. Romania Kolcsar et al., 2013: month(s): 6-8. Russia Pilipenko, 2016: month(s): 7. Spain Carles-Tolra, 2010: month(s): 5-8. Camano Portela et al., 2011: month(s): 7. Pino Perez and Pino Perez, 2014: month(s): 8. Pino Perez and Pino Perez, 2016: month(s): 8. Eiroa and Carles-Tolra, 2019: month(s): 7. Switzerland Weibel and Kessler, 2021: month(s): 6. on altitude (mainly from 2000 onward): Austria Vogtenhuber and Kofler, 2017: altitude: 640 m. Croatia Kolcsar et al., 2023b: altitude: 170 m. Cyprus Kolcsar et al., 2023a: altitude: 530 m. France Oosterbroek in litt., 2024: altitude: 200 m. Liechtenstein Aistleitner, 2015: altitude: 600 m. Portugal Oosterbroek et al., 2020: altitude: 850 m. Spain Camano Portela et al., 2011: altitude: 980 m. Pino Perez and Pino Perez, 2014: altitude: 618 m. Pino Perez and Pino Perez, 2016: altitude: 55 m. Switzerland Weibel and Kessler, 2021: altitude: 890 m.