Details for Achyrolimonia neonebulosa

Name: Achyrolimonia neonebulosa (Alexander, 1924) Publication: Philipp. J. Sci. 24: 555 (as Limonia) Status: Recognized taxon, Synonym(s): nebulosa (Alexander, 1913); delicatula (Lackschewitz, 1964). Classification: Family Limoniidae Subfamily Limoniinae Revision: Podenas and Byun, 2014a. Descriptive note: As new name for Dicranomyia nebulosa Alexander, 1913. Keys: Podenas and Gelhaus, 2007 (Limoniinae Mongolia and nearby); Peeters en Oosterbroek, 2014c (craneflies Netherlands, Belgium and Luxembourg) (in Dutch); Podenas et al., 2015c, Podenas, 2016c (Achyrolimonia Korea) habitus male habitus male habitus female body part(s) abdomen hypopygium plus 21 more images of hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, ovipositor, ovipositor, ovipositor, wing, wing, wing, wing, wing, wing and wing. Distribution: USA (Mass, NY);; Austria, Bulgaria, Croatia, Czech Rep., Denmark, France, Germany, Great Britain, Hungary, Italy, Lithuania, Netherlands, Norway, Poland, Romania, Serbia, Slovakia, Switzerland, Ukraine; Russia: RUC, North Caucasus;; Iran; Russia: WS (Altay), FE (Khabarovskiy kray, Primorskiy kray, Sakhalin); North Korea, South Korea, Japan (Hokkaido, Honshu, Shikoku), China (see citations for provinces). Note: see the manual for abbreviations: Canada and USA; present-day Russia. Region(s): Nearctic;; Westpalaearctic;; Eastpalaearctic General note: Added: Serbia (Simova-Tosic and Vukovic, 1996). For Norway check Olsen et al., 2018, and citations below. For Great Britain check Stubbs, 2021, and citations below. Map: Podenas et al., 2006 (Switzerland); Podenas and Byun, 2014a (Korea) Biology: Stary, 1998b (habitat). Figure references: Habitus: James and Kramer, 2019 (male); Wiedenska and Syratt, 2021 (male) Wing: Alexander, 1913e (as nebulosa); Podenas et al., 2006; Podenas and Gelhaus, 2007; Podenas and Byun, 2014a; Podenas, 2016c; James and Kramer, 2019 Hypopygium: Alexander, 1913e (as nebulosa); Podenas et al., 2006; Kramer and Withers, 2007; Podenas and Gelhaus, 2007; Podenas and Byun, 2014a; Podenas, 2016c; James and Kramer, 2019; Sugaro and Kato, 2023; Kolcsar et al., 2025 Ovipositor: Podenas, 2016c; Kolcsar et al., 2025 Other figures: James and Kramer, 2019 (abdomen) Immature stages Larva: Krivosheina and Krivosheina, 2011c Pupa: James and Kramer, 2019 Citations on biology (mainly from 2000 onward): Country not relevant Habitat. Krivosheina and Krivosheina, 2011c: 1-294 (larva: loc(s), descr, figs, biology, key) (in Russian, key also in English). Denmark Habitat. Kolcsar et al., 2025: 19 (loc(s) Copenhagen, Vallensbæk Strand, Plantorama, plant and garden center, the same plant shop as Atypophthalmus umbratus in 2021). Europe Biology. Kolcsar et al., 2025: 19-22 (review habitats and biology; recent reports show that the species has also been found in greenhouses, or emerged from indoor plant pots in the Netherlands (Oosterbroek 2009), Great Britain (James & Kramer 2019), Denmark (this paper), represented the first records for these countries, and Romania (this paper), where it has also been reported from a natural habitat (Erhan 1984, Ujvarosi et al. 2011)). France Habitat. Kramer and Withers, 2007: 158, 160 (species with larvae in dry or more or less wet decaying wood, citing Brindle, 1960c; also citing Stary, 2002a; habitat details of locality). Habitat. Withers, 2008: 133 (dead wood species). Great Britain Biology. James and Kramer, 2019: 195-196 (an adult male emerged from the pupa on 24 september 2019 from the compost of a plant pot containing an imported plant of Sarracenia (a carnivorous pitcher plant) which was imported from the Netherlands; see paper for details). Italy Habitat. Stary, 2002a: 106 (semi-dry Quercus forest). Japan Biology. Hayashi and Naba, 1991: 31 (in Japanese with short English summary: record of Limonia (Limonia) neonebulosa Alexander injures the sphagnum and root of the orchid). Poland Biology. Wiedenska, 2015b: 57-58 (review biology, citing various authors) (in Polish). Habitat. Wiedenska and Syratt, 2021: 428 (short descr, figs, loc(s), distr, habitat) (in Polish). Romania Habitat. Ujvarosi et al., 2011b: 108 (loc(s) Dupa Lanca marshy area near Voslobeni) (in Hungarian, species list with habitat information in English). Biology. Kolcsar et al., 2025: 19 (at Cluj-Napoca, Flower Time Garden center, trapped on the same Drosera sp. plant as a male of Rhipidia domestica; based on the label of the plant pot, it originated from the Netherlands). Russia Habitat. Krivosheina, N.P., 2009a: 125-133 (review Russian xylophilous Limoniidae, mainly inhabiting bark and wood of decomposing trees, with new original data on larval biology [found in wood of deciduous trees]) (in Russian). Habitat. Dvorak et al., 2020: 5 (on Diptera collected in beer traps in the RUC: Mordoviya Rep.; collected in a mixed forest). Slovakia Biology. Obona et al., 2025b: 100, 102 (sporadic non-target catch in CO₂-baited BG-Sentinel 2 traps used for mosquito surveillance). South Korea Habitat. Podenas and Byun, 2014a: 170 (collected at light in a small hotel, which was surrounded by agricultural fields with irrigation channel. Forest was at the far end of the field; according to Savchenko (1985), adults are found in wet broad leaved and mixed forests and in Eriophorum - Carex community in marsh surrounded by pine forest in Ukraine). on characters and taxonomy (mainly from 2000 onward): China Description. Zhang, X. et al., 2023: 25 (descr, figs wing, hypopygium after Alexander; loc(s) Zhejiang, list of Chinese provinces, mentioning Sichuan only) (in Chinese). Country not relevant Immatures. Krivosheina and Krivosheina, 2011c: 1-294 (larva: loc(s), descr, figs, biology, key) (in Russian, key also in English). East Palaearctic Key. Podenas and Gelhaus, 2007: 29 (key, figs). Great Britain Characters. James and Kramer, 2019: 195-196 (char, comparison, figs). Immatures. James and Kramer, 2019: 195-196 (char, fig pupa). Poland Characters. Wiedenska and Syratt, 2021: 428 (short descr, figs, loc(s), distr, habitat) (in Polish). South Korea Revision. Podenas and Byun, 2014a: 169-170 (revision, figs). Characters. Podenas, 2016c: 48 (key, figs). on distribution (mainly from 2000 onward): Bulgaria Distribution. Hubenov, 2021a: 36 (checklist with summary of distr in Bulgaria, references). China Distribution. Zhang, X. et al., 2023: 25 (descr, figs wing, hypopygium after Alexander; loc(s) Zhejiang, list of Chinese provinces, mentioning Sichuan only) (in Chinese). Distribution. Provinces Eastpalaearctic part, 2026: (as far south as Zhejiang and Sichuan, provinces to be worked out). Country not relevant Distribution. Krivosheina and Krivosheina, 2011c: 1-294 (larva: loc(s), descr, figs, biology, key) (in Russian, key also in English). Croatia First record. Oosterbroek and Simova-Tosic, 2004: 446 (loc(s)). Czech Rep. Distribution. Stary, 2009i: webpage (checklist). Denmark First record. Kolcsar et al., 2025: 19 (loc(s) Copenhagen, Vallensbæk Strand, Plantorama, plant and garden center, the same plant shop as Atypophthalmus umbratus in 2021). Europe Distribution. Kolcsar et al., 2025: 19-22 (review distributional history; it is suggested to treat the species as an invasive non-native species, in the countries where it was introduced through plant trade; see paper for details). France Distribution. Kramer and Withers, 2007: 158, 160, 163 (loc(s) Ain (01)). First record. Kramer, 2007a: 33 (loc(s) Ain (01)). Distribution. Kramer, 2007d: 1 (loc(s) Verot Nat. Res.). Distribution. Withers, 2008: 133, 137 (loc(s) Ain (01)). Germany Distribution. Schacht, 1999: 135 (checklist Bayern). Distribution. Reusch and Oosterbroek, 2000: 160 (checklist German Bundeslander). Distribution. Schacht, 2005: 4 (checklist Bayern). Distribution. Hable et al., 2010b: 90 (loc(s) Bayern). Distribution. Schacht, 2010: 16 (checklist Diptera Bayern). Distribution. Dunk and Kraus, 2014: 169 (loc(s) Bayern). Distribution. Stuke, 2019: 141 (checklist Niedersachsen and Bremen with reference to original sources). Great Britain First record. James and Kramer, 2019: 195-196 (loc(s) East Suffolk, Norfolk). Hungary First record. Kolcsar and Soltesz, 2018: 165 (loc(s) Pest county). Italy First record. Stary, 2002a: 106 (loc(s) Mantova). Japan Distribution. Kato and Suzuki, 2017: 13 (revised list of the crane flies of Kanagawa pref., Honshu, distr, refs). Distribution. Kato and Yamauchi, 2024: 22-23 (loc(s) Toyama pref., Honshu). Distribution. Suzuki et al., 2024: 6 (records from Hadano city, Honshu) (in Japanese). Distribution. Kato and Endo, 2025: 17 (loc(s) Honshu) (in Japanese). Distribution. Kato and Kawase 2025: 16 (annotated checklist of the Gifu prefecture, Honshu). Lithuania First record. Pakalniskis et al., 2000: 7 (checklist) (details in Podenas and Pakalniskis, 2000). Distribution. Podenas and Pakalniskis, 2000: 22 (loc(s)). Distribution. Pakalniskis et al., 2006: 17 (checklist). Netherlands Distribution. Oosterbroek, 2009c: 180 (found in an Orchid greenhouse). First record. Pijnakker et al., 2006: 5, 10, 12 (found in Zuid-Holland at Wateringen in Cambria orchid) (in Dutch). Distribution. Stary, 2020a: 45 (Limoniidae Kaaistoep, Noord-Brabant). North Korea First record. Podenas et al., 2017: 261 (loc(s)). Norway Distribution. Olsen et al., 2018: 160-161 (annotated checklist Nordic countries and for Norway according to the Strand regions). Poland First record. Wiedenska, 2015b: 56 (loc(s) near Lodz). Distribution. Wiedenska and Syratt, 2021: 428 (short descr, figs, loc(s), distr, habitat) (in Polish). Romania Distribution. Ujvarosi, 2007: 228-231 (checklist Limoniidae). Distribution. Ujvarosi et al., 2011b: 108 (loc(s) Dupa Lanca marshy area near Voslobeni) (in Hungarian, species list with habitat information in English). Distribution. Kolcsar et al., 2025: 19 (at Cluj-Napoca, Flower Time Garden center, trapped on the same Drosera sp. plant as a male of Rhipidia domestica; based on the label of the plant pot, it originated from the Netherlands). Russia Distribution. Pilipenko and Sidorenko, 2006b: 146 (checklist Kedrovaya Pad, distr) (in Russian). Distribution. Pilipenko, 2009c: 333 (loc(s) Lazovsky Nature Reserve, Primorskiy kray). Distribution. Paramonov and Pilipenko, 2016: 99 (loc(s) RUC: Tverskaya oblast, distr) (in Russian). Distribution. Dvorak et al., 2020: 5 (on Diptera collected in beer traps in the RUC: Mordoviya Rep.; collected in a mixed forest). Distribution. Pilipenko et al., 2020: 359 (loc(s) RUC: Mordoviya Rep., distr). Distribution. Devyatkov and Dubatolov, 2025: 33 (loc(s) Khabarovskiy kray, Bolshekhekhtsirsky Nature Reserve, distr) (in Russian). Serbia Distribution. Oosterbroek and Simova-Tosic, 2004: 446 (review literature). Slovakia Distribution. Stary, 2009i: webpage (checklist). South Korea First record. Podenas and Byun, 2014a: 170, 181 (loc(s), map). Switzerland Distribution. Podenas et al., 2006: 152, 197 (map, vertical distr per thermic level). USA Distribution. Brodo et al., 2022a: 53 (list of Tipuloidea collected in the NY, Putnam County, town of Kent). on flight period (mainly from 2000 onward): Denmark Kolcsar et al., 2025: month(s): 12. France Kramer and Withers, 2007: month(s): 6. Kramer, 2007a: month(s): 6. Kramer, 2007d: month(s): 5. Great Britain James and Kramer, 2019: month(s): 9. Hungary Kolcsar and Soltesz, 2018: month(s): 6, 8. Japan Kato and Yamauchi, 2024: month(s): 7-9. Suzuki et al., 2024: month(s): 11. Kato and Endo, 2025: month(s): 5, 8-9. Lithuania Podenas and Pakalniskis, 2000: month(s): 6, 8. Netherlands Oosterbroek, 2009c: month(s): 1. Pijnakker et al., 2006: month(s): 1. North Korea Podenas et al., 2017: month(s): 7-8. Poland Wiedenska, 2015b: month(s): 6-8. Wiedenska and Syratt, 2021: month(s): 5. Romania Ujvarosi et al., 2011b: month(s): 6. Kolcsar et al., 2025: month(s): 3. Russia Paramonov and Pilipenko, 2016: month(s): 6. Dvorak et al., 2020: month(s): 8-9. Pilipenko et al., 2020: month(s): 8. Devyatkov and Dubatolov, 2025: month(s): 9. South Korea Podenas and Byun, 2014a: month(s): 6. Switzerland Podenas et al., 2006: month(s): 7, 9. on altitude (mainly from 2000 onward): Bulgaria Hubenov, 2021a: altitude: 500 m. Hungary Kolcsar and Soltesz, 2018: altitude: 284-385 m. Japan Kato and Yamauchi, 2024: altitude: 330-1120 m. North Korea Podenas et al., 2017: altitude: 1372-2134 m. Romania Kolcsar et al., 2025: altitude: 320 m. Russia Devyatkov and Dubatolov, 2025: altitude: 80 m. South Korea Podenas and Byun, 2014a: altitude: 633 m.