Details for Helius (Helius) calviensis

Name: Helius (Helius) calviensis Edwards, 1928 Publication: Encycl. Ent. (B II) Dipt. 4: 183 (as Helius) Status: Recognized taxon. Classification: Family Limoniidae Subfamily Limoniinae Keys: Quindroit, 2022c (European Helius, in French and English); Solodkii and Pilipenko, 2025 (Helius Russia) habitus female body part(s) head and antenna hypopygium hypopygium hypopygium plus 10 more images of hypopygium, hypopygium, hypopygium, ovipositor, wing, wing, wing, wing, habitat and habitat. Distribution: Albania, Bulgaria, France (incl. Corsica), Greece, Italy (incl. Sardinia, Sicily), Montenegro, Portugal, Romania, Spain (incl. Balearic Is [Ibiza, Mallorca]); Russia: North Caucasus; Algeria, Tunisia; Turkey (Asiatic part: Afyonkarahisar, Balikesir, Canakkale). Note: see the manual for abbreviations: present-day Russia. Region(s): Westpalaearctic General note: For Italy check Stary and Oosterbroek, 1996, and citations below. For Spain check Eiroa and Carles-Tolra, 2019, and citations below. For Portugal check Oosterbroek et al., 2020, and citations below. For France check Quindroit, 2022c, and citations below. Map: Quindroit et al., 2021; Quindroit, 2022c (France) Figure references: Habitus: Quindroit et al., 2021 (female) Head: Quindroit, 2022c Wing: Quindroit, 2022c Hypopygium: Mendl, 1979a; Quindroit et al., 2021; Quindroit, 2022c Ovipositor: Quindroit, 2022c Citations on biology (mainly from 2000 onward): Algeria Habitat. Stary and Oosterbroek, 2008: 14 (marsh and Pistaccia carr). France Habitat. Quindroit et al., 2021: 256-258 (collected during an entomological inventory in the canopy of Oak [Quercus] stands, see paper for habitat and forest details; marais). Habitat. Quindroit, 2022a: 22, 28 (annotated list with information on abundancy and habitat of the Tipuloidea of the Pays de la Loire regoin, distr) (in French). Habitat. Quindroit, 2022c: 343-350 (review of the five French species of Helius, loc(s) 7 departements, map) (in French). Habitat. Quindroit et al., 2023: 503 (on Corsica collected at sea level, in a riparian undergrowth of Alnus). Russia Biology. Lantsov, 2015a: 366-367 (loc(s) North Caucasus: Kabardino-Balkar. Rep., Dagestan, with data on habitats, ecology and chorology) (in Russian) (as flavus, see Latsov, 2020). Habitat. Lantsov, 2020: 293 (North Caucasus: found in Dagestan, in the Samur liana forest; in Lantsov, 2015a [and 2019c] recorded as Helius flavus). Spain Habitat. Stary and Oosterbroek, 2008: 14 (meadows along river). Habitat. Stary, 2014b: 92 (collected at saltmarsh and canal). Habitat. Mederos et al., 2025b: 161-162 (loc(s) Barcelona, distr; see habitat photo). on characters and taxonomy (mainly from 2000 onward): Country not relevant Characters. Quindroit et al., 2021: 258 (comparison genital characters, figs). Characters. Solodkii and Pilipenko, 2025: 400, 406 (notes on the morphology of Helius, comparison). France Key. Quindroit, 2022c: 343-350 (review of the five French species of Helius, key, figs) (in French, key also in English). Portugal Characters. Ferreira et al., 2021: 10 (barcoding based on material mentioned in Oosterbroek et al. 2020). on distribution (mainly from 2000 onward): Albania First record. Kolcsar et al., 2021b: 123 (loc(s)). Algeria First record. Stary and Oosterbroek, 2008: 14 (loc(s), distr). Bulgaria Distribution. Hubenov, 2021a: 36 (checklist with summary of distr in Bulgaria, references). France Distribution. Quindroit et al., 2021: 257 (recorded for Loiret (45); first record for mainland France, distr, map). Distribution. Quindroit, 2022a: 22, 28 (annotated list with information on abundancy and habitat of the Tipuloidea of the Pays de la Loire regoin, distr) (in French). Distribution. Quindroit, 2022c: 343-350 (review of the five French species of Helius, loc(s) 7 departements, map) (in French). Distribution. Quindroit and Racine, 2025: 360 (table specifying the presence in the five departments of the Pays de la Loire region). Greece First record. Kolcsar et al., 2021b: 123 (loc(s) Eastern Macedonia and Thrace). Italy Distribution. Podenas, 2011: 865 (first record for Sardinia, distr). Montenegro First record. Kolcsar et al., 2021b: 123 (loc(s)). Portugal First record. Stary, 2014b: 92 (loc(s) Faro). Distribution. Oosterbroek et al., 2020: 333 (loc(s) Setubal, also known from Faro, distr). Romania First record. Kolcsar et al., 2021b: 123 (loc(s) Tulcea). Russia First record. Lantsov, 2015a: 366-367 (loc(s) North Caucasus: Kabardino-Balkar. Rep., Dagestan, with data on habitats, ecology and chorology) (in Russian) (as flavus, see Latsov, 2020). Distribution. Lantsov, 2019c: 134 (North Caucasus: on tipuloid dipterans of the protected areas of Dagestan) (in Russian) (as flavus, see Latsov, 2020). Distribution. Lantsov, 2020: 293 (North Caucasus: found in Dagestan, in the Samur liana forest; in Lantsov, 2015a [and 2019c] recorded as Helius flavus). Spain First record. Stary and Oosterbroek, 2008: 14 (loc(s), distr). Distribution. Oosterbroek, 2009c: 181 (new for La Coruna). Distribution. Stary, 2014b: 92 (loc(s) Malaga, Murcia, Mallorca). Distribution. Eiroa and Carles-Tolra, 2019: 132 (loc(s) Zaragoza, list of Spanish provinces, distr) (in Spanish). Distribution. Mederos et al., 2025b: 161-162 (loc(s) Barcelona, distr; see habitat photo). Tunisia First record. Stary and Oosterbroek, 2008: 14 (loc(s), distr). Turkey First record. Ozgul and Koc, 2012: 750 (first record for Turkey, prov. Afyonkarahisar). First record. Koc et al., 2016: 194 (loc(s) Asiatic part, Marmara region) (as flavus, teste J. Stary). on flight period (mainly from 2000 onward): Albania Kolcsar et al., 2021b: month(s): 5. Algeria Stary and Oosterbroek, 2008: month(s): 4. France Quindroit, 2022a: month(s): 5-6, 9. Quindroit, 2022c: month(s): 4-6, 8-9. Greece Kolcsar et al., 2021b: month(s): 5. Italy Podenas, 2011: month(s): 5. Montenegro Kolcsar et al., 2021b: month(s): 5. Portugal Stary, 2014b: month(s): 4. Oosterbroek et al., 2020: month(s): 5. Romania Kolcsar et al., 2021b: month(s): 6 or 7. Russia Lantsov, 2015a: month(s): 5, 7. Spain Stary and Oosterbroek, 2008: month(s): 5. Stary, 2014b: month(s): 3-5, 8. Eiroa and Carles-Tolra, 2019: month(s): 9. Mederos et al., 2025b: month(s): 6. Tunisia Stary and Oosterbroek, 2008: month(s): 4. Turkey Ozgul and Koc, 2012: month(s): 5. Koc et al., 2016: month(s): 4. on altitude (mainly from 2000 onward): Albania Kolcsar et al., 2021b: altitude: 12 m. Algeria Stary and Oosterbroek, 2008: altitude: 0 m. Bulgaria Hubenov, 2021a: altitude: 0-20 m. Greece Kolcsar et al., 2021b: altitude: 10 m. Montenegro Kolcsar et al., 2021b: altitude: 20 m. Portugal Stary, 2014b: altitude: 24-500 m. Romania Kolcsar et al., 2021b: altitude: 3 m. Russia Lantsov, 2015a: altitude: 20-870 m. Spain Stary and Oosterbroek, 2008: altitude: 150 m. Stary, 2014b: altitude: 0-820 m. Mederos et al., 2025b: altitude: 170 m. Tunisia Stary and Oosterbroek, 2008: altitude: 0 m. Turkey Ozgul and Koc, 2012: altitude: 848-85 m. Koc et al., 2016: altitude: 23-190 m.