Details for Dicranomyia (Dicranomyia) modesta

Name: Dicranomyia (Dicranomyia) modesta (Meigen, 1818) Publication: Syst. Beschr. 1: 134 (as Limnobia) Status: Recognized taxon, Synonym(s): simplex (Meigen, 1830); nova (Meigen, 1830); takahashii Alexander, 1920; spinicauda Alexander, 1924. Classification: Family Limoniidae Subfamily Limoniinae Descriptive note: For authorship see Sabrosky, 1999. Synonymy: Alexander, 1948b, 1955c. Keys: Podenas and Gelhaus, 2007 (Limoniinae Mongolia and nearby); Oosterbroek et al., 2007 (Limoniidae Greenland); Reusch and Weber, 2013 (Luxembourg); Boardman, 2016 (craneflies Shropshire); Stubbs and Kramer, 2016h; Stubbs, 2021 (British craneflies); Podenas et al., 2019a (Dicranomyia Korea) habitus male habitus male habitus male habitus male habitus male plus 48 more images of habitus, habitus, habitus, habitus, habitus, body part(s), body part(s), body part(s), body part(s), body part(s), body part(s), body part(s), body part(s), body part(s), body part(s), body part(s), body part(s), hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, ovipositor, wing, wing, wing, wing, wing, wing, habitat, habitat, habitat, habitat, habitat, habitat, habitat, habitat, habitat, habitat, habitat, habitat, habitat, habitat, habitat, habitat, habitat and habitat. Distribution: Canada, USA (NWT, Utah); Greenland;; Albania, Austria, Belarus, Belgium, Bosnia-Herzegovina, Bulgaria, Croatia, Czech Rep., Denmark, Estonia, Finland, France (incl. Corsica), Germany, Great Britain, Greece, Hungary, Iceland, Ireland, Italy, Latvia, Lithuania, Luxembourg, Montenegro, Netherlands, North Macedonia, Norway, Poland, Portugal, Romania, Serbia, Slovakia, Slovenia, Spain (incl. Canary Is [Tenerife]), Sweden, Switzerland, Turkey (European part: Kirklareli), Ukraine; Russia: RUN, RUW, RUC, RUE, North Caucasus; Morocco; Georgia, Armenia, Azerbaijan, Turkey (Asiatic part: see citations for provinces), Iran;; Russia: WS, ES, FE; Kazakhstan, Turkmenistan, Tajikistan, Kyrgyzstan, Afghanistan; Mongolia, North Korea, South Korea, Japan (Hokkaido, Honshu, Shikoku, Kyushu), China (Inner Mongolia). Note: see the manual for abbreviations: Canada and USA; present-day Russia. Region(s): Nearctic;; Westpalaearctic;; Eastpalaearctic General note: For Italy check Stary and Oosterbroek, 1996, and citations below. For Finland check Salmela, 2012b, 2012c, and citations below. For Norway check Olsen et al., 2018, and citations below. For Spain check Eiroa and Carles-Tolra, 2019, and citations below. For Portugal check Oosterbroek et al., 2020, and citations below. For Great Britain and Ireland check Stubbs, 2021, and citations below. For Morocco check Kettani and Oosterbroek, 2022a, and citations below. Map: Koc, 2004 (Mugla); Podenas et al., 2006 (Switzerland); Oosterbroek et al., 2007 (Greenland); Boardman, 2007 (Shropshire); Podenas et al., 2019a (Korea) Biology: Schmidt, 1913 (habitat). Figure references: Habitus: Kato, 2023a (male); Quindroit et al., 2023 (male); Kato and Hirago, 2024 (male); Kato and Kawase, 2024 (female) Head: Podenas et al., 2006 (antenna); Podenas and Gelhaus, 2007 (antenna); Podenas et al., 2019a (antenna) Wing: Slipka and Stary, 1977b; Podenas et al., 2006; Oosterbroek et al., 2007; Podenas et al., 2019a Hypopygium: Alexander, 1924n (as spinicauda); Podenas et al., 2006; Oosterbroek et al., 2007; Podenas and Gelhaus, 2007; Podenas et al., 2019a; Kramer and Morris, 2022b Ovipositor: Podenas et al., 2019a Other figures: Frantsevich, 2004 (mid coxa); Podenas et al., 2019a (tarsomeres) Immature stages Larva: Podeniene, 2003a; Krivosheina and Krivosheina, 2011c Citations on biology (mainly from 2000 onward): Azerbaijan Habitat. Stary and Obona, 2020: 136 (loc(s); with list of localities, habitats and altitudes). Bulgaria Habitat. Hubenov, 2025b: suppl: 7 (annotated list Diptera high Bulgarian mts, local distr, habitats and zoogeography). Country not relevant Habitat. Krivosheina and Krivosheina, 2011c: 1-294 (larva: descr, figs, key, biology) (in Russian, key also in English) (based on Lindner, 1959). Czech Rep. Habitat. Stary et al., 2013b: 377 (on Limonidae from Vraz near Pisek, with general habitat description). Habitat. Stary and Vonicka, 2018: 64 (on Limoniidae and Pediciidae of two localities in northern Bohemia, with list of localities, altitudes and habitats). Denmark Habitat. Byriel and Rojas, 2017: 118 (among the 101 species in a study on cranefly hotspots in unmanaged and managed forests) (in Danish). Finland Phenology. Salmela, 2001a: 143, 144, 151 (obviously two generations per year). Habitat. Salmela, 2004: 9 (eurytopic, ubiquitous species inhabiting a wide spectrum of habitats). Habitat. Salmela and Ilmonen, 2005: 89 (the occurrence in relation to the trophic status of mires within an open boreal mire system is investigated). Habitat. Kahanpaa and Salmela, 2007: 43-45, 48 (found at a small stream with a width of three to five meters, a riverbed of soft muddy sediments, and surrounded by fully grown but not particularly old Spruce [Picea abies] forest, small stands of old Birch [Betula] and Aspen [Populus tremula] are present but there is little dead wood available in the forest) (this is a summary of the available habitat information in Finnish). Biology. Salmela, 2008: 11, 27, 51 (49) (ecology or habitat: eurytopic, wet; phenology). Habitat. Autio and Salmela, 2010: 48 (habitats Aland Is). Habitat. Rasimus, 2018: 17 (on saproxylic craneflies in the Tampere region) (in Finnish). France Habitat. Kramer and Withers, 2007: 159 (marshland species with larvae in the more or less saturated organic mud of lake margins and marshes). Habitat. Kramer, 2007a: 33 (adults were collected at an open marsh). Biology. Quindroit, 2020a: 34-35, 45-48 (annotated list with information on abundancy and habitat of the Tipuloidea of the Pays de la Loire region, distr); see also Quindroit and Racine, 2025: 337 (habitat) (in French). Habitat. Quindroit, 2020b: 134 (loc(s) Nord (59) and Pas-de-Calais (62) with notes on habitats and distr) (in French). Habitat. Quindroit and Lemoine, 2021a: 23, 24 (loc(s), resultats de deux campagnes de piegeage dans le Pas-de-Calais (62); see paper for details on habitat). Habitat. Quindroit, 2021a: 36 (loc(s), resultats d une campagne de piegeage dans le Pas-de-Calais (62); see paper for details on habitat). Habitat. Quindroit and Lemoine, 2022a: 22 (loc(s) Oise (60) and Somme (80) with notes on habitats and distr) (in French). Georgia Habitat. Stary and Obona, 2020: 136 (loc(s); with list of localities, habitats and altitudes). Germany Habitat. Reusch, 2006: 134 (Kalkquellmoor [carbonate fen complex]). Habitat. Dau, 2013: 42, Anhang 1: 32 (among species found in Salzquellen in Schleswig-Holstein). Great Britain Habitat. Godfrey, 2000a: 20, 22. Habitat. Godfrey, 2001a: 22, 24. Habitat. Boardman, 2007: 78 (occurring virtually anywhere that wet soils are present). Habitat. Irwin, 2010: 169 (a marine, estuarine or brackish species that can develop in mud). Habitat. Drake, 2011a: 24 (among the Diptera from a wet Alder [Alnus] wood in Devon). Habitat. Brighton, 2017c: 28, 38 (overall regional checklist Lancashire and Cheshire (VC58, 59 and 60) with notes on habitats and/or regional occurrence and/or status; see paper for details). Greece Habitat. Kramer, 2004: 20, 21 (found at a grassy verge). Greenland Biology. Oosterbroek et al., 2007: 27 (one of the most common eurytopic species of the genus (Savchenko, 1982; Salmela, 2004), occurring in a large variety of wet biotopes, such as different kinds of forest, shrub, bogs and meadows communities; in mountain regions it does not penetrate above forest level (Tjeder, 1958); in at least part of the range there are two or even more generations a year (Savchenko, 1982; Salmela, 2001a); females lay 130-160 eggs; the larvae are terrestrial to semi-aquatic, with habitats ranging from spring and brook environments to brackish (Lindner, 1959) or even halophilous conditions (Schmidt, 1913)). Kazakhstan Habitat. Devyatkov, 2013: 89 (collected at small freshwater pools and wet habitats near a salty steppe lake, surrounding vegetation was grazed grassland). Korea Habitat. Podenas et al., 2019a: 54 (open grassy areas near human settlements). Biology. Podenas et al., 2019a: 54 (period of flight; altitudes; attracted to light). Latvia Habitat. Spungis, 2008: 36 (raised bogs). Lithuania Habitat. Podeniene, 2003a: 1-295 (larva, descr, figs, habitat) (in Lithuanian). Habitat. Podeniene, 2003b: 11, 21, 22, 28 (larval habitat: sand and silt in riparian zone of small rivulets, mud in marshy meadows). Habitat. Gorban and Podeniene, 2021: 1-5 (aim of the study was to investigate the biodiversity of nematoceran flies associated with dead wood in a forest ecosystem in Lithuania; during this research, emergence traps were used on fallen tree trunks of Aspen [Populus tremula] and Ash [Fraxinus excelsior] at two localities in Lithania, findings (from dead Ash only) are accidental, larvae develop in mud in marshy areas; see paper for details). Biology. Gorban and Podeniene, 2022: 5 (loc(s); reared from Ash [Fraxinus excelsior]; see paper for details). Luxembourg Habitat. Reusch and Weber, 2013: 270-271 (found in a cave, summary of biology). Mongolia Habitat. Gelhaus and Podenas, 2006: 289, 295 (habitat preference: 100% lotic). Habitat. Yadamsuren et al., 2015: 474 (riparian zone). Habitat. Oliveira and Eck, 2025: Specieslist (collected in an open Pinus forest and along a river). Morocco Habitat. Driauach and Belqat, 2016: 140 (loc(s) Rif Mts with list of habitats). Netherlands Habitat. Hop and Moonen, 2021: 37 (on the occurrence (rarity) in Dutch inland waters) (in Dutch). Norway Habitat. Olsen and Andersen, 2021: 214 (loc(s) Innlandet, in former Hedmark, with habitats specified separately, for habitats see also Jonassen and Andersen, 2020). Poland Habitat. Wiedenska, 2014: 14 (on the occurence of Limoniidae in six selected plant communities in the Kamienica Valley, Gorce National Park, Western Carpathians, found only in Dentario glandulosae-Fagetum allietosum habitat) (in Polish, habitat tables also in English). Phenology. Wiedenska, 2015a: 54 (on the flight activity of Limoniidae in the Kamienica Valley, Gorce National Park, Western Carpathians) (in Polish, tables also in English). Habitat. Wiedenska, 2017a: 22 (faunistic and habitat data of species from the Gorce NP) (in Polish). Portugal Habitat. Stary, 2014b: 90 (collected at brook and marsh). Romania Habitat. Ujvarosi and Poti, 2006: 256 (habitat in Dupa Lunca marshland: predominantly at damp meadows). Habitat. Ujvarosi et al., 2011b: 108 (loc(s) Dupa Lanca marshy area near Voslobeni) (in Hungarian, species list with habitat information in English). Habitat. Kolcsar et al., 2013: 71 (loc(s) Cluj-Napoca and surroundings; list of habitats with altitudes). Russia Habitat. Gavryushin in litt., 2011f: (collected in a herbaceous vegetation with Urtica, Chamerion, umbellifers and grasses near a ditch). Habitat. Lantsov, 2017b: 107 (on dominant species of the crane flies (Diptera, Tipuloidea) of the North Caucasus as biological indicators of habitats) (in Russian). Habitat. Krivosheina et al., 2018: 6-7 (larvae are saprophages developing in silty accretions, moss hummocks, moist litter, humus-rich soils, and decomposing substrates; they typically occur in humid biotopes; it is an eurytopic species since it can also be found in saline habitats (Krivosheina and Krivosheina, 2011); it has not previously been recorded in a woody substrate, but according to the literature, some representatives of the genus are xylobionts; in particular, larvae of D. ochripes Alexander, 1955 develop in the outer layers of light and loose wood in fallen trunks of the Amur Cork tree [Phellodendron amurense] and the Castor Aralia [Kalopanax septemlobus] (Krivosheina and Krivosheina, 2011); larvae of this species were found under the bark of Abies nephrolepis trees partly abandoned by the four-eyed fir bark beetle in Primorskiy kray). Habitat. Hancock, 2019: 49 (loc(s) RUN: Murmanskaya oblast, collected from along banks of rivers). Habitat. Kolcsar et al., 2021b: 59 (Tatarstan Rep.: ravine, wetland). Slovakia Habitat. Beuk et al., 2023: 46, 54 (travertine rocks). Sweden Habitat. Lindstrom and Fritz, 2015: 44, 47 (distributin and ecology in five fens in Hallands County) (in Schwedish). Turkey Biology. Koc, 2004: 104, 111, 116-118, 153 (habitat; phenology) (in Turkish). on characters and taxonomy (mainly from 2000 onward): Country not relevant Characters. Frantsevich, 2004: 133-134 (structure of coxa, figs). Phylogeny. Boldgiv, 2006: 96-154 (one of the 84 morphological taxa from northern Mongolia in a consistency test of morphology based species identification of crane flies against DNA-barcoding identification using 536-base-pair sequence profiles of the COI gene, also known as the DNA barcode). Immatures. Krivosheina, N.P., 2009d: 140 (comparison larva). Immatures. Krivosheina and Krivosheina, 2011c: 1-294 (larva: descr, figs, key, biology) (in Russian, key also in English) (based on Lindner, 1959). Phylogeny. Kang et al., 2023: 735-741 (phylogeny Tipuloidea based on mitochondrial genome sequencing). Phylogeny. Song et al., 2023: 7-9 (phylogeny Tipuloidea based on mitochondrial genome sequencing). Phylogeny. Xu et al., 2023a: 5, 20-21 (phylogeny Tipuloidea based on mitochondrial genome sequencing). Phylogeny. Gao, Y. et al., 2024: 267-288 (phylogeny Tipuloidea based on mitochondrial genome sequencing). Phylogeny. Feng et al., 2025: 4-17 (phylogeny Tipuloidea based on mitochondrial genome sequencing). Great Britain Characters. Kramer and Morris, 2022b: 18 (diagnosis, figs). Greenland Key. Oosterbroek et al., 2007: 6-11 (key, figs). Korea Description. Podenas et al., 2019a: 12-15, 51-54 (type-mat takahashii, descr, key, comparison, figs). Lithuania Immatures. Podeniene, 2003a: 1-295 (larva, descr, figs, habitat) (in Lithuanian). Mongolia Key. Podenas and Gelhaus, 2007: 26, 37, 44-45 (key, figs). Portugal Characters. Ferreira et al., 2021: 8 (barcoding based on material mentioned in Oosterbroek et al. 2020). on distribution (mainly from 2000 onward): Albania First record. Kolcsar et al., 2021b: 54 (loc(s)). Armenia Distribution. Obona et al., 2016: 135, 140 (loc(s), checklist). Distribution. Hakobyan and Jenderedjian, 2023: 1514 (annotated checklist of the craneflies of Armenia with refs, distr and flight period in Caucasus Ecoregion with refs). Austria Distribution. Reusch and Heiss, 2012: 314-322 (loc(s) Nat. Park Gesause, see PDF for months and altitudes). Azerbaijan Distribution. Stary and Obona, 2020: 136 (loc(s); with list of localities, habitats and altitudes). Distribution. Snegovaya, 2021: 1136 (checklist with details on distr in Azerbaijan). Belarus First record. Kolcsar et al., 2021b: 54-55 (loc(s) various provinces). Belgium Distribution. Martens et al., 2014: 114, 123 (loc(s) Oost-Vlaanderen). Distribution. Dek and Peeters, 2023a: 136 (loc(s) Botanic Garden Jean Massart, Brussels-Capital Region, common in Belgium). Bosnia-Herzegowina Distribution. Oosterbroek and Simova-Tosic, 2004: 447 (review literature). Bulgaria Distribution. Hubenov, 2018b: 124 (review loc(s) Vrachanska Planina Mts). Distribution. Hubenov, 2021a: 36 (checklist with summary of distr in Bulgaria, references). Distribution. Hubenov, 2025b: suppl: 7 (annotated list Diptera high Bulgarian mts, local distr, habitats and zoogeography). Canary Is Distribution. Stary, 2014b: 90 (loc(s) Malaga and Tenerife, first record for Canary Is). Channel Is Distribution. Stubbs, 2022b: 5 (on a collection of craneflies from Jersey, collected between 1985 and 2002 by Tony Warne). China First record. Yu et al., 2016: 349 (loc(s) Inner Mongolia). Croatia Distribution. Oosterbroek and Simova-Tosic, 2004: 447 (review literature). Distribution. Kolcsar et al., 2015a: 33 (loc(s), distr). Czech Rep. Distribution. Stary et al., 2005a: 25, 30 (loc(s), among the most frequent limoniid species in the studied area). Distribution. Stary, 2009i: webpage (checklist). Distribution. Stary et al., 2013b: 377 (on Limonidae from Vraz near Pisek, with general habitat description). Distribution. Stary and Vonicka, 2018: 64 (on Limoniidae and Pediciidae of two localities in northern Bohemia, with list of localities, altitudes and habitats). Denmark Distribution. Petersen and Jong, 2001b: 149 (checklist). Estonia First record. Salmela in litt., 2003: (new for Estonia). Finland Distribution. Salmela, 2001a: 143, 144, 151 (loc(s)). Distribution. Salmela, 2002: 30-32 (loc(s) lake Iidesjarvi) (in Finnish). Distribution. Salmela, 2004: 9 (loc(s)). Distribution. Stary and Salmela, 2004: 47 (loc(s)). Distribution. Salmela and Ilmonen, 2005: 89 (loc(s)). Distribution. Kahanpaa and Salmela, 2007: 48 (loc(s)). Distribution. Salmela et al., 2007b: 37, 46 (abundance, loc(s)). Distribution. Salmela, 2008: 11, 51 (49) (loc(s), known from all Finnish ecoregions). Distribution. Autio and Salmela, 2010: 48 (loc(s) Aland Is). Distribution. Salmela, 2010a: 8, 12 (survey fauna Tahmela spring in city of Tampere) (in Finnish). Distribution. Salmela, 2012b: 236 (annotated list of Finnish crane flies). Distribution. Salmela and Petrasiunas, 2014: 29 (checklist Finnish Tipulomorpha). Distribution. Salmela et al., 2015: 100 (list of localities in extensive report on malaise-trapping of insects in conservation areas in Lapland in 2012–2014) (in Finnish). Distribution. Rasimus, 2018: 17 (on saproxylic craneflies in the Tampere region) (in Finnish). France Distribution. Kramer and Withers, 2007: 159, 163 (loc(s) Ain (01)). Distribution. Kramer, 2007a: 33 (loc(s) Ain (01)). Distribution. Withers, 2008: 137 (loc(s) Ain (01)). Distribution. Garrin and Herbrecht, 2016: Annexe 11 (loc(s) Notre-Dame-des-Landes, Loire-Atlantique (44)). Distribution. Quindroit, 2020a: 34-35, 45-48 (annotated list with information on abundancy and habitat of the Tipuloidea of the Pays de la Loire region, distr); see also Quindroit and Racine, 2025: 337 (habitat) (in French). Distribution. Quindroit, 2020b: 134 (loc(s) Nord (59) and Pas-de-Calais (62) with notes on habitats and distr) (in French). Distribution. Quindroit and Lemoine, 2021a: 23, 24 (loc(s), resultats de deux campagnes de piegeage dans le Pas-de-Calais (62); see paper for details on habitat). Distribution. Quindroit, 2021a: 36 (loc(s), resultats d une campagne de piegeage dans le Pas-de-Calais (62); see paper for details on habitat). Distribution. Quindroit and Lemoine, 2022a: 22 (loc(s) Oise (60) and Somme (80) with notes on habitats and distr) (in French). Distribution. Quindroit and Lemoine, 2022a: 37 (table specifying the presence in the five departments of the Hauts-de-France region). Distribution. Lemoine and Quindroit, 2023: 8 (loc(s) Seine-et-Marne (77)). Distribution. Quindroit and Racine, 2025: 359 (table specifying the presence in the five departments of the Pays de la Loire region). Georgia Distribution. Stary and Obona, 2020: 136 (loc(s); with list of localities, habitats and altitudes). Germany Distribution. Schacht, 1999: 135 (checklist Bayern). Distribution. Reusch and Oosterbroek, 2000: 161 (checklist German Bundeslander). Distribution. Reusch and Bellstedt, 2001b: 75 (checklist Thuringen). Distribution. Schacht, 2005: 4 (checklist Bayern). Distribution. Reusch, 2006: 134 (loc(s) Bayern). Distribution. Hable et al., 2010b: 90 (loc(s) Bayern). Distribution. Schacht, 2010: 16 (checklist Diptera Bayern). Distribution. Dau, 2013: 42, Anhang 1: 32 (among species found in Salzquellen in Schleswig-Holstein). Distribution. Dunk and Kraus, 2014: 169 (loc(s) Bayern). Distribution. Stuke, 2019: 141 (checklist Niedersachsen and Bremen with reference to original sources). Distribution. Stuke et al., 2020: 283 (loc(s) Niedersachsen). Distribution. Stuke et al., 2024: 161 (loc(s) Niedersachsen). Great Britain Distribution. Godfrey, 2000a: 20, 22 (see paper for details of sites surveyed in Shropshire and into Wales). Distribution. Godfrey, 2001a: 22, 24 (recorded from sites in Shropshire and just over the country border in Montgmeryshire; see paper for details). Distribution. Hancock, 2002: 13 (loc(s) Scotland). Distribution. Hewitt et al., 2005: 37 (loc(s) Cumbria) (as Limonia). Distribution. Boardman, 2007: 78 (loc(s), map). Distribution. Drake, 2008c: 224 (loc(s) Wiltshire, in study on invertebrates of two water meadows). Distribution. Skidmore, 2009: 129 (review of occurrence on the Western Isles of Scotland, distr). Distribution. Drake, 2011a: 24 (loc(s) Devon). Distribution. Hancock, 2011a: 2 (loc(s) Tiree and Coll, Southern Inner Hebrides). Distribution. Kramer, 2011j: 21 (loc(s) Leicestershire and Rutland (VC55)). Distribution. Kramer, 2013i: 8 (checklist of species from Leigh Woods NNR, Bristol district). Distribution. Hewitt, 2014: (provisional checklist Cumbrian Diptera). Distribution. Kramer, 2014c: 4 (in table of cranefly species recorded at Shapwick Heath NNR, as given in H. Audcent, 1949 and/or collected by J. Kramer in 2013). Distribution. Wolton et al., 2014: 26 (among Diptera associated with a hedge in Devon, caught in emergence trap). Distribution. Chandler, 2015: 90 (loc(s) Bushy Park, Middlesex). Distribution. Drake, 2016: 20 (study on the relative importance to Diptera of pasture and ditch margins on Catcott Reserve, Somerset). Distribution. Showers, 2016: 2 (loc(s) Northamptonshire). Distribution. Brighton, 2017c: 28, 38 (overall regional checklist Lancashire and Cheshire (VC58, 59 and 60) with notes on habitats and/or regional occurrence and/or status; see paper for details). Distribution. Showers, 2017a: 4 (loc(s) Northamptonshire). Distribution. Kramer and Morris, 2022b: 18 (info on the species, on the distr and map of loc(s) Leicestershire and Rutland (VC55)). Greece First record. Kramer, 2004: 20, 21 (loc(s) Peloponnisos). Greenland Distribution. Oosterbroek et al., 2007: 5, 8-9, 13, 27 (review record Greenland, distr, map). Iceland First record. Kolcsar et al., 2021b: 55 (loc(s)). Ireland Distribution. Ashe et al., 2008: 9 (additional Irish record(s), distr). Distribution. Chandler et al., 2008: 13 (checklist). Italy Distribution. Podenas and Podeniene, 2008: 347, 351-352 (loc(s), annotated checklist South Tyrol [Prov. Bolzano]). Japan Distribution. Kimura et al., 2011: 94 (on species composition and abundance in higland lakes on Honshu, loc(s)). Distribution. Podenas et al., 2019a: 51 (type-loc(s) takahashii Honshu). Distribution. Kato, 2023a: 25 (loc(s) Honshu) (in Japanese). Distribution. Kato and Hiraga, 2024: 33 (loc(s) Niigata, Honshu) (in Japanese). Distribution. Kato and Kawase, 2024: 9 (loc(s) Koka city, Shiga, Honshu) (in Japanese). Distribution. Kato and Yamauchi, 2024: 25 (loc(s) Toyama pref., Honshu). Distribution. Kato, 2024b: 9 (records from Tokamachi city, Honshu) (in Japanese). Kazakhstan Distribution. Devyatkov, 2013: 89 (loc(s) Pavlodarskaya oblast). Distribution. Devyatkov, 2019b: 246 (loc(s) Pavlodarskaya oblast, distr) (in Russian). Distribution. Devyatkov, 2020a: 50 (loc(s) East Kazakhstan, distr) (in Russian). Latvia Distribution. Stary, 2004f: (Fauna Europaea: added Latvia). Distribution. Spungis, 2008: 36 (loc(s)). Lithuania Distribution. Pakalniskis et al., 2006: 18 (checklist). Distribution. Gorban and Podeniene, 2021: 1-5 (aim of the study was to investigate the biodiversity of nematoceran flies associated with dead wood in a forest ecosystem in Lithuania; during this research, emergence traps were used on fallen tree trunks of Aspen [Populus tremula] and Ash [Fraxinus excelsior] at two localities in Lithania, findings (from dead Ash only) are accidental, larvae develop in mud in marshy areas; see paper for details). Distribution. Gorban and Podeniene, 2022: 5 (loc(s); reared from Ash [Fraxinus excelsior]; see paper for details). Luxembourg Distribution. Reusch and Weber, 2013: 270 (loc(s), map, distr). Mongolia Distribution. Boldgiv, 2006: 153 (loc(s)). First record. Gelhaus and Podenas, 2006: 289, 295 (loc(s)). Distribution. Yadamsuren et al., 2015: 474 (loc(s) Mongolia). Distribution. Oliveira and Eck, 2025: Specieslist (loc(s) Ulaanbaatar and Khentii aimags). Montenegro Distribution. Oosterbroek and Simova-Tosic, 2004: 447 (review literature). Morocco First record. Driauach and Belqat, 2016: 140 (loc(s) Rif Mts with list of habitats). Distribution. Kettani and Oosterbroek, 2022a: 26 (annotated checklist, with review loc(s) and/or references). Netherlands Distribution. Jong and Oosterbroek, 2002b: 33 (checklist). Distribution. Cuppen, 2009: 6 (loc(s) larval records). Distribution. Hoven, 2015: 10 (loc(s) Limburg). Distribution. Hoven, 2018: 16 (loc(s) Zeeland). Distribution. Stary, 2020a: 45 (Limoniidae Kaaistoep, Noord-Brabant). North Korea First record. Podenas et al., 2019a: 51, 104 (loc(s), map). North Macedonia Distribution. Oosterbroek and Simova-Tosic, 2004: 447 (review literature). Norway Distribution. Olsen et al., 2018: 160-161 (annotated checklist Nordic countries and for Norway according to the Strand regions). Distribution. Olsen and Andersen, 2021: 214 (loc(s) Innlandet, in former Hedmark, with habitats specified separately, for habitats see also Jonassen and Andersen, 2020). Poland Distribution. Wiedenska, 2007a: 74 (checklist). Distribution. Wiedenska, 2008: 46-47 (loc(s) Parowy Janinowski nature reserve near Lodz) (in Polish). Distribution. Wiedenska, 2010: 131 (loc(s) Parku Krajobrazowego Wzniesien Lodzkich, Lodz). Distribution. Wiedenska, 2014: 14 (on the occurence of Limoniidae in six selected plant communities in the Kamienica Valley, Gorce National Park, Western Carpathians, found only in Dentario glandulosae-Fagetum allietosum habitat) (in Polish, habitat tables also in English). Distribution. Wiedenska, 2015b: 56 (loc(s) near Lodz). Distribution. Wiedenska, 2017a: 22 (faunistic and habitat data of species from the Gorce NP) (in Polish). Distribution. Wiedenska and Syratt, 2023: 146 (loc(s) Bieszczady Mountains, distr and remarks). Portugal First record. Stary, 2014b: 90 (loc(s) Faro). Distribution. Oosterbroek et al., 2020: 330 (recorded from seven provinces, distr). Romania Distribution. Parvu, 2004: 185 (loc(s)). Distribution. Parvu, 2005: 149 (loc(s)). Distribution. Ujvarosi, 2005a: 244 (no details). Distribution. Parvu, 2006: 261 (loc(s)). Distribution. Ujvarosi and Poti, 2006: 256 (loc(s) eastern Carpathians). Distribution. Ujvarosi, 2007: 228-231 (checklist Limoniidae). Distribution. Ujvarosi et al., 2011b: 108 (loc(s) Dupa Lanca marshy area near Voslobeni) (in Hungarian, species list with habitat information in English). Distribution. Keresztes, 2012: 30 (loc(s) Rimetea area [Alba County]). Distribution. Kolcsar et al., 2013: 71 (loc(s) Cluj-Napoca and surroundings; list of habitats with altitudes). Russia Distribution. Lantsov, 2003e: 83 (loc(s) Caucasus region, distr). Distribution. Lantsov, 2004: 63, 66-67 (loc(s) Caucasus region, distr). Distribution. Polevoi and Humala, 2005: 181 (RUN: list of Diptera of Vepsskaya Volost, Kareliya) (in Russian). Distribution. Pilipenko and Sidorenko, 2006b: 147 (loc(s) Kedrovaya Pad, distr) (in Russian). Distribution. Humala and Polevoi, 2009: 66 (loc(s) RUN: Kareliya) (in Russian). Distribution. Pilipenko, 2009c: 334 (loc(s) Lazovsky Nature Reserve, Primorskiy kray). Distribution. Gavryushin in litt., 2011f: (loc(s) RUN: Arkhangelskaya oblast). Distribution. Paramonov, 2011: 41 (loc(s)) RUE: Mariy El Rep.). Distribution. Humala and Polevoi, 2012: 143 (loc(s) RUN: Kareliya) (in Russian). Distribution. Jakovlev et al., 2014: 309 (loc(s) RUN: Kareliya). Distribution. Paramonov and Klepikov, 2014: 36 (loc(s) Yaroslavskaya oblast). Distribution. Lantsov, 2016: 165-170 (on craneflies from Arkhyz, Karachay-Cherkessia Rep., North Caucasus) (in Russian). Distribution. Paramonov and Pilipenko, 2016: 99 (loc(s) RUC: Tverskaya oblast) (in Russian). Distribution. Lantsov, 2017b: 107 (on dominant species of the crane flies (Diptera, Tipuloidea) of the North Caucasus as biological indicators of habitats) (in Russian). Distribution. Paramonov, 2017a: 172 (loc(s) RUC: Chuvash Rep.) (in Russian). Distribution. Krivosheina et al., 2018: 7 (loc(s) Primorskiy kray). Distribution. Hancock, 2019: 49 (loc(s) RUN: Murmanskaya oblast, collected from along banks of rivers). Distribution. Pilipenko et al., 2020: 359 (loc(s) RUC: Mordoviya Rep., distr). Distribution. Kolcsar et al., 2021b: 55-61 (loc(s) RUE: Bashkortostan Rep. and Tatarstan Rep.). Distribution. Devyatkov and Dubatolov, 2025: 34 (loc(s) Khabarovskiy kray, Bolshekhekhtsirsky Nature Reserve, distr) (in Russian). Serbia Distribution. Oosterbroek and Simova-Tosic, 2004: 447 (review literature). Distribution. Gavryushin in litt., 2015b: (loc(s) Serbia). Slovakia Distribution. Stary, 2009d: 34 (loc(s) Polana area). Distribution. Stary, 2009i: webpage (checklist). Distribution. Beuk et al., 2023: 46, 54 (loc(s)). Slovenia Distribution. Oosterbroek and Simova-Tosic, 2004: 447 (review literature). South Korea First record. Podenas et al., 2019a: 51-53, 104 (loc(s), map). Spain Distribution. Eiroa and Baez, 2002a: 55 (checklist). Distribution. Hancock et al., 2015a: 241 (loc(s) Lugo). Distribution. Eiroa and Carles-Tolra, 2019: 131 (loc(s) Barcelona, Tarragona and Zaragoza, list of Spanish provinces, distr) (in Spanish). Distribution. Keresztes et al., 2022: 195 (loc(s) Asturias and review distribution in Spain) (in Spanish). Distribution. Mederos et al., 2025b: 161 (loc(s) Gerona, distr). Sweden Distribution. Salmela, 2010b: 133 (loc(s) Malmo). Distribution. Lindstrom and Fritz, 2015: 44, 47 (distributin and ecology in five fens in Hallands County) (in Schwedish). Switzerland Distribution. Podenas et al., 2006: 159, 224 (map, vertical distr per thermic level). Distribution. Dufour and Merz, 2012: 354 (annotated checklist canton Geneva). Tajikistan Distribution. Devyatkov, 2020c: 76 (loc(s), distr) (in Russian). Turkey First record. Koc, 2004: 104, 111, 116-118, 153 (loc(s), distr, map) (in Turkish). Distribution. Ozgul et al., 2009: 63 (survey provinces SW Turkey). Distribution. Bilgin et al., 2015: 6 (loc(s) Eskisehir, distr) (in Turkish). Distribution. Koc et al., 2016: 11 (provinces Marmara region). Distribution. Akpinar and Ozgul, 2023: 55 (loc(s) Aksaray). Distribution. Provinces Asiatic part, 2025: (known from Aydin, Denizli, Eskisehir, Mugla, Marmara region). on flight period (mainly from 2000 onward): Albania Kolcsar et al., 2021b: month(s): 6. Armenia Obona et al., 2016: month(s): 8-9. Azerbaijan Stary and Obona, 2020: month(s): 5, 9. Belarus Kolcsar et al., 2021b: month(s): 5-6. Belgium Martens et al., 2014: month(s): 5, 7, 9. Dek and Peeters, 2023a: month(s): 4-11. Canary Is Stary, 2014b: month(s): 1, 3-4. Channel Is Stubbs, 2022b: month(s): 6, 10. Croatia Kolcsar et al., 2015a: month(s): 5. Czech Rep. Stary et al., 2005a: month(s): 5-9. Stary et al., 2013b: month(s): 4 (8-10). Stary and Vonicka, 2018: month(s): 5-10. Finland Salmela, 2001a: month(s): 6, 8, 10. Salmela, 2008: month(s): 7-8. France Kramer and Withers, 2007: month(s): 5-7. Kramer, 2007a: month(s): 6. Quindroit, 2020a: month(s): 6-7, 9-10. Quindroit, 2020b: month(s): 5-11. Quindroit and Lemoine, 2022a: month(s): 5, 9. Lemoine and Quindroit, 2023: month(s): 5. Georgia Stary and Obona, 2020: month(s): 4-5. Germany Stuke et al., 2024: month(s): 6. Great Britain Godfrey, 2000a: month(s): 6. Godfrey, 2001a: month(s): 5. Hancock, 2002: month(s): 6-7. Hancock, 2011a: month(s): 6. Kramer, 2011j: month(s): 6-10. Kramer, 2014c: month(s): 6. Chandler, 2015: month(s): 10-11. Kramer and Morris, 2022b: month(s): 5-10. Greece Kramer, 2004: month(s): 4. Iceland Kolcsar et al., 2021b: month(s): 6. Ireland Ashe et al., 2008: month(s): 6, 8. Italy Podenas and Podeniene, 2008: month(s): 5-10. Japan Podenas et al., 2019a: month(s): 5. Kato, 2023a: month(s): 5-7, 10-11. Kato and Hiraga, 2024: month(s): 5, 9, 11. Kato and Kawase, 2024: month(s): 4-5, 11-12. Kato and Yamauchi, 2024: month(s): 7-9. Kato, 2024b: month(s): 11. Kazakhstan Devyatkov, 2013: month(s): 9. Devyatkov, 2019b: month(s): 6-9. Devyatkov, 2020a: month(s): 6-10. Korea Podenas et al., 2019a: month(s): 4-11. Luxembourg Reusch and Weber, 2013: month(s): 7. Mongolia Gelhaus and Podenas, 2006: month(s): 7. Oliveira and Eck, 2025: month(s): 7-8. Morocco Driauach and Belqat, 2016: month(s): 3-4. Netherlands Hoven, 2015: month(s): 5. Hoven, 2018: month(s): 5. Norway Olsen and Andersen, 2021: month(s): 6-9. Poland Wiedenska, 2008: month(s): 5-10. Wiedenska, 2014: month(s): 6. Wiedenska, 2015a: month(s): 6-7, 9. Wiedenska, 2017a: month(s): 6-8, 10. Wiedenska and Syratt, 2023: month(s): 6-9. Portugal Stary, 2014b: month(s): 3-5. Oosterbroek et al., 2020: month(s): 2, 5-6, 9. Romania Parvu, 2004: month(s): 6. Parvu, 2005: month(s): 5-6. Parvu, 2006: month(s): 6. Ujvarosi et al., 2011b: month(s): 5-6. Kolcsar et al., 2013: month(s): 6, 8-10. Russia Lantsov, 2003e: month(s): 10. Lantsov, 2004: month(s): 5, 10. Pilipenko and Sidorenko, 2006b: month(s): 7. Pilipenko, 2009c: month(s): 7. Gavryushin in litt., 2011f: month(s): 8. Paramonov, 2011: month(s): 6. Paramonov and Klepikov, 2014: month(s): 6-10. Paramonov and Pilipenko, 2016: month(s): 6-10. Paramonov, 2017a: month(s): 5, 7. Hancock, 2019: month(s): 8. Pilipenko et al., 2020: month(s): (7)8-9. Kolcsar et al., 2021b: month(s): 6-8. Devyatkov and Dubatolov, 2025: month(s): 8-9. Serbia Gavryushin in litt., 2015b: month(s): 7. Slovakia Stary, 2009d: month(s): 5-7, 9-10. Beuk et al., 2023: month(s): 6. Spain Hancock et al., 2015a: month(s): 5. Eiroa and Carles-Tolra, 2019: month(s): 4, 8, 11. Keresztes et al., 2022: month(s): 8. Mederos et al., 2025b: month(s): 5. Switzerland Podenas et al., 2006: month(s): 4-11. Tajikistan Devyatkov, 2020c: month(s): 7. Turkey Koc, 2004: month(s): 4-6. on altitude (mainly from 2000 onward): Albania Kolcsar et al., 2021b: altitude: 696 m. Armenia Obona et al., 2016: altitude: 940-1930 m. Belarus Kolcsar et al., 2021b: altitude: 156 m. Bulgaria Hubenov, 2021a: altitude: 10-720 m. Hubenov, 2025b: altitude: 10-720 m. Croatia Kolcsar et al., 2015a: altitude: 55 m. Germany Stuke et al., 2024: altitude: 200-270 m. Greece Kramer, 2004: altitude: 875 m. Iceland Kolcsar et al., 2021b: altitude: 15 m. Italy Podenas and Podeniene, 2008: altitude: 940-2030 m. Japan Kimura et al., 2011: altitude: 764-822 m. Kato and Yamauchi, 2024: altitude: 330-664 m. Kazakhstan Devyatkov, 2019b: altitude: 61-438 m. Devyatkov, 2020a: altitude: 158-1170 m. Korea Podenas et al., 2019a: altitude: 0-1700 m. Mongolia Gelhaus and Podenas, 2006: altitude: 1645 m. Oliveira and Eck, 2025: altitude: 950-1290 m. Morocco Driauach and Belqat, 2016: altitude: 36-394 m. Poland Wiedenska, 2017a: altitude: 720-1070 m. Wiedenska and Syratt, 2023: altitude: 470-1200 m. Portugal Stary, 2014b: altitude: 24-500 m. Romania Ujvarosi and Poti, 2006: altitude: 670 m. Russia Lantsov, 2003e: altitude: 914 m. Gavryushin in litt., 2011f: altitude: 20-25 m. Kolcsar et al., 2021b: altitude: 50-777 m. Devyatkov and Dubatolov, 2025: altitude: 70-100 m. Serbia Gavryushin in litt., 2015b: altitude: 800 m. Slovakia Stary, 2009d: altitude: 560-870 m. Beuk et al., 2023: altitude: 575 m. Spain Hancock et al., 2015a: altitude: 720 m. Keresztes et al., 2022: altitude: 250 m. Tajikistan Devyatkov, 2020c: altitude: 2250 m. Turkey Koc, 2004: altitude: 0-765 m.