Details for Dicranomyia (Dicranomyia) mitis
Name:Dicranomyia (Dicranomyia) mitis (Meigen, 1830)
Publication:Syst. Beschr. 6: 278 (as Limnobia)
Status:Recognized taxon, Synonym(s): excisa (Walker, 1848); unicolor (Strobl, 1898); maculosa (Strobl, 1900); lutea Lackschewitz, 1928; infuscata Lackschewitz, 1928.
Classification:Family Limoniidae
Subfamily Limoniinae
Revision:Stary and Stubbs, 2015.
Keys:Podenas and Gelhaus, 2007 (Limoniinae Mongolia and nearby); Stary and Freidberg, 2007 (Limoniidae Israel); Stary and Stubbs, 2015 (species Dicranomyia mitis complex); Boardman, 2016 (craneflies Shropshire); Stubbs and Kramer, 2016h; Stubbs, 2021 (British craneflies)
Dicranomyia (Dicranomyia) mitis : habitus - maleDicranomyia (Dicranomyia) mitis : body part(s) - clawDicranomyia (Dicranomyia) mitis : body part(s) - clawDicranomyia (Dicranomyia) mitis : body part(s) - clawDicranomyia (Dicranomyia) mitis : hypopygium
habitus
male
body part(s)
claw
body part(s)
claw
body part(s)
claw
hypopygium
plus 13 more images of hypopygium, hypopygium, hypopygium, ovipositor, ovipositor, wing, wing, wing, habitat, habitat, habitat, habitat and habitat.
Distribution:Albania, Andorra, Austria, Belarus, Belgium, Bosnia-Herzegovina, Bulgaria, Croatia, Czech Rep., Denmark (incl. Faroe Is), Finland, France (incl. Corsica), Germany, Great Britain, Greece, Hungary, Ireland, Italy, Lithuania, Luxembourg, Montenegro, Netherlands, North Macedonia, Norway, Poland, Romania, Serbia, Slovakia, Slovenia, Spain, Sweden, Switzerland, Turkey (European part: widespread), Ukraine; Russia: RUN, RUW, RUC, RUE, North Caucasus; Morocco, Algeria; Georgia, Azerbaijan, Turkey (Asiatic part: see citations for provinces), Israel;; Turkmenistan.
Note: see the manual for abbreviations: present-day Russia.
Region(s):Westpalaearctic;; Eastpalaearctic
General note:Added: Andorra (Geiger et al., 1994). Confirmed for Corsica by Podenas et al., 1997. For Italy check Stary and Oosterbroek, 1996, and citations below. For Finland check Salmela, 2012b, 2012c, and citations below. For Norway check Olsen et al., 2018, and citations below. For Great Britain and Ireland check Stubbs, 2021, and citations below. For Morocco check Kettani and Oosterbroek, 2022a, and citations below.
Map:Koc, 2004 (Mugla); Podenas et al., 2006 (Switzerland); Boardman, 2007 (Shropshire; separate maps for affinis Schummel, lutea Meigen, and mitis Meigen)
Figure
references:
Wing: Podenas et al., 2006; Stary and Stubbs, 2015
Hypopygium: Stary, 1969b; Alexander, 1975e; Podenas et al., 2006; Podenas and Gelhaus, 2007; Stary and Stubbs, 2015; Kramer, 2021; Kramer and Morris, 2022b
Ovipositor: Stary and Stubbs, 2015
Other figures: Stary, 1993a (male tarsal claw); Stary and Stubbs, 2015 (distal tarsomeres and claw)

Immature stages
Larva: Podeniene, 2003a

Miscellaneous
Quindroit, 2023 (habitat)
Citationson biology (mainly from 2000 onward):
Bulgaria
Habitat. Hubenov, 2025a: suppl: 4 (annotated list Diptera Stara Planina Mts, local distr, habitats and zoogeography).
Habitat. Hubenov, 2025b: suppl: 7 (annotated list Diptera high Bulgarian mts, local distr, habitats and zoogeography).
Czech Rep.
Habitat. Stary and Vonicka, 2018: 64 (on Limoniidae and Pediciidae of two localities in northern Bohemia, with list of localities, altitudes and habitats).
Denmark
Habitat. Nielsen and Nielsen, 2009: 121 (emerging from a forest floor in a Beech [Fagus] stand) (in Danish).
Habitat. Byriel and Rojas, 2017: 118 (among the 101 species in a study on cranefly hotspots in unmanaged and managed forests) (in Danish).
Finland
Habitat. Salmela, 2004: 9 (perhaps brooks and small rivers).
Habitat. Salmela, 2008: 11, 51 (49) (ecology or habitat: swamp).
Habitat. Autio and Salmela, 2010: 48 (habitats Aland Is) (as cf mitis).
France
Habitat. Kramer and Withers, 2007: 159 (marshland species with larvae in the more or less saturated organic mud of lake margins and marshes).
Biology. Quindroit, 2020a: 34, 45-48 (annotated list with information on abundancy and habitat of the Tipuloidea of the Pays de la Loire regoin, distr) (in French).
Habitat. Quindroit, 2020b: 134 (loc(s) Nord (59) and Pas-de-Calais (62) with notes on habitats and distr) (in French).
Habitat. Herbrecht et al., 2021: 57, 58 (loc(s) Loire-Atlantique (44) based on malaise-trapping with notes on habitats) (in French).
Habitat. Quindroit et al., 2021: 256-258 (collected during an entomological inventory in the canopy of Oak [Quercus] stands; sous-bois a tendance humide).
Habitat. Quindroit, 2021c: 424, 426 (loc(s) Mercantour National Park, Alpes-Maritimes (06), common species in France and Europe, with wide geographical distribution; ubiquitous, considered as common at all altitudes (Geiger, 1985); known from wet woodland in Northern France, to Mediterranean area (Var) and Corsica).
Habitat. Quindroit and Lemoine, 2022a: 22 (loc(s) Oise (60) with notes on habitats and distr) (in French).
Habitat. Quindroit, 2023: 62 (loc(s) Hautes-Alpes (05)) [locality and habitat are not given in the text but with the habitat figure].
Germany
Habitat. Reusch and Hohmann, 2009: 128, 129 (from spring brooks).
Great Britain
Habitat. Godfrey, 2000a: 20, 22.
Habitat. Godfrey, 2001a: 21 (as mitis s.l.).
Habitat. Rotheray, 2001: 15 (reared from Alder [Alnus] and Elm [Ulmus]).
Habitat. Hancock, 2002: 14 (recorded in this study emerging from water-logged wood).
Habitat. Stubbs, 2003: 50 (similar to Dicranomyia sericata and Limonia nigropunctata but more widespread).
Biology. Stubbs, 2006a: 134 (behaviour and feeding on Acer campestre leaf) (as Dicranomyia of the mitis group).
Habitat. Boardman, 2007: 77 (mainly a woodland cranefly, also to be found along woodland edge or hedgerows and amongst well established scrub).
Habitat. Phillips, 2007: 149 (flushes running down to beach).
Habitat. Cranston and Drake, 2010: 175 (larvae can be found in marginal situations along flowing waters).
Biology. Stary and Stubbs, 2015: 326 (the species is found mainly on well-drained clay or limestone soils; it is absent on acid soils, can be present under mesotrophic conditions, but shows greatest affinity to mild or strong calcareous situations, preferring scrub edges associated with calcareous grassland; it is found more patchily to the north and west; the species can occur within woodland, but rarely in numbers; it is a member of the vernal fauna (may–early june), but, on an exceptional occasion, it was found in the autumn (october)).
Habitat. Brighton, 2017c: 28, 38 (overall regional checklist Lancashire and Cheshire (VC58, 59 and 60) with notes on habitats and/or regional occurrence and/or status; see paper for details).
Greece
Habitat. Kramer, 2004: 20, 21 (found at a stream margin and grassy verge).
Lithuania
Habitat. Podeniene, 2003a: 1-295 (larva, descr, figs, habitat) (in Lithuanian).
Poland
Habitat. Wiedenska, 2014: 14 (on the occurence of Limoniidae in six selected plant communities in the Kamienica Valley, Gorce National Park, Western Carpathians, found only in Petasitetum kablikiani habitat) (in Polish, habitat tables also in English).
Phenology. Wiedenska, 2015a: 54 (on the flight activity of Limoniidae in the Kamienica Valley, Gorce National Park, Western Carpathians) (in Polish, tables also in English).
Habitat. Wiedenska, 2017a: 21-22 (faunistic and habitat data of species from the Gorce NP) (in Polish).
Romania
Habitat. Ujvarosi, 2005a: 244 (hygropetric zone, swamps, moist soil, citing various sources).
Habitat. Ujvarosi et al., 2011b: 108 (loc(s) Dupa Lanca marshy area near Voslobeni) (in Hungarian, species list with habitat information in English).
Habitat. Kolcsar et al., 2013: 70, 71 (loc(s) Cluj-Napoca and surroundings; list of habitats with altitudes; one of the most frequent species).
Russia
Habitat. Lantsov, 2009a: 75 (ecology) (in Russian).
Habitat. Lantsov, 2017b: 107 (on dominant species of the crane flies (Diptera, Tipuloidea) of the North Caucasus as biological indicators of habitats) (in Russian).
Habitat. Paramonov and Korobkov, 2019: 94-95 (loc(s) RUC: Tverskaya oblast, distr, habitat, citing various authors) (in Russian).
Turkey
Biology. Koc, 2004: 103, 111, 116-118, 153 (habitat; phenology) (in Turkish).
on characters and taxonomy (mainly from 2000 onward):
Country not relevant
Characters. Stary, 2006b: 201 (comparison).
Characters. Stary, 2009a: 56 (comparison).
Revision. Stary and Stubbs, 2015: 321-334 (revision, descr, comparison, key, figs).
East Palaearctic
Key. Podenas and Gelhaus, 2007: 47 (key, figs).
Great Britain
Taxonomy. Kramer, 2010c: 2-3 (member of mitis-complex, comparison).
Characters. Kramer, 2021: 1-3 (characters, identification, comparison, figs).
Ireland
Taxonomy. Ashe et al., 2008: 9 (comment on the mitis complex in Ireland).
Israel
Key. Stary and Freidberg, 2007: 305, 331-332, 335-336, 339-341 (key, comparison).
Lithuania
Immatures. Podeniene, 2003a: 1-295 (larva, descr, figs, habitat) (in Lithuanian).
on distribution (mainly from 2000 onward):
Algeria
First record. Vaillant, 1952: 248-249 (refera to didyma, confirmed by Stary in litt., june 2013).
Andorra
Distribution. Eiroa and Baez, 2002a: 55 (checklist).
Austria
Distribution. Reusch and Heiss, 2012: 314-322 (loc(s) Nat. Park Gesause, see PDF for months and altitudes).
Distribution. Aistleitner, 2015: 101 (loc(s) Vorarlberg).
Distribution. Stary and Stubbs, 2015: 324-326 (loc(s), distr).
Azerbaijan
Distribution. Snegovaya, 2021: 1136 (checklist with details on distr in Azerbaijan).
Belarus
First record. Kolcsar et al., 2021b: 53 (loc(s) Vitebsk).
Belgium
Distribution. Dek and Peeters, 2023a: 136 (loc(s) Botanic Garden Jean Massart, Brussels-Capital Region, common in Belgium).
Bosnia-Herzegowina
Distribution. Chvala, 2008: 251 (type-loc(s) of unicolor).
Bulgaria
Distribution. Hubenov, 2015: 221 (annotated list Diptera Pirin Mt, loc(s), faunistics and zoogeography).
Distribution. Stary and Stubbs, 2015: 324-326 (loc(s), distr).
Distribution. Hubenov, 2016: 49 (annotated checklist Diptera of the Rila Mountains based on data from the literature).
Distribution. Hubenov, 2017: 70 (on vertical distribution in the Pirin and Rila Mts; see paper for details on forests and subalpine-alpine zones, distr).
Distribution. Hubenov, 2018a: 16 (on the Diptera of the Vitosha Mt; see paper for details on vegetation belts and local distr).
Distribution. Hubenov, 2018b: 124 (review loc(s) Vrachanska Planina Mts).
Distribution. Hubenov, 2021a: 36 (checklist with summary of distr in Bulgaria, references).
Distribution. Hubenov, 2025a: suppl: 4 (annotated list Diptera Stara Planina Mts, local distr, habitats and zoogeography).
Distribution. Hubenov, 2025b: suppl: 7 (annotated list Diptera high Bulgarian mts, local distr, habitats and zoogeography).
Croatia
Distribution. Oosterbroek and Simova-Tosic, 2004: 447 (review literature).
Distribution. Kolcsar et al., 2015a: 33 (loc(s), distr) (identification confirmed in Ivkovic et al., 2020).
Distribution. Kolcsar et al., 2023b: 15 (loc(s)).
Czech Rep.
Distribution. Stary et al., 2005a: 25 (loc(s)).
Distribution. Stary, 2009i: webpage (checklist).
Distribution. Mocek, 2014: 156 (loc(s) NE Bohemia).
Distribution. Stary and Stubbs, 2015: 324-326, 333 (loc(s), distr, in the Czech Republic and Slovakia, this is the commonest and most widespread species [of the mitis-complex] occurring from high altitudes to lowland).
Distribution. Stary and Vonicka, 2018: 64 (on Limoniidae and Pediciidae of two localities in northern Bohemia, with list of localities, altitudes and habitats).
Denmark
Distribution. Petersen and Jong, 2001b: 149 (checklist).
Distribution. Nielsen and Nielsen, 2009: 121 (loc(s)).
Finland
Distribution. Salmela, 2004: 9 (loc(s)).
Distribution. Salmela, 2008: 11, 51 (49) (loc(s), known from all Finnish ecoregions).
Distribution. Autio and Salmela, 2010: 48 (loc(s) Aland Is) (as cf mitis).
Distribution. Salmela, 2012b: 236 (annotated list of Finnish crane flies).
Distribution. Salmela and Petrasiunas, 2014: 29 (checklist Finnish Tipulomorpha).
France
Distribution. Kramer and Withers, 2007: 159, 163 (loc(s) Ain (01)).
Distribution. Withers, 2008: 137 (loc(s) Ain (01)).
Distribution. Quindroit, 2020a: 34, 45-48 (annotated list with information on abundancy and habitat of the Tipuloidea of the Pays de la Loire regoin, distr) (in French).
Distribution. Quindroit, 2020b: 134 (loc(s) Nord (59) and Pas-de-Calais (62) with notes on habitats and distr) (in French).
Distribution. Quindroit, 2020d: 328 (collected at the Golf de Vidauban, Vidauban, Var (83)).
Distribution. Herbrecht et al., 2021: 57, 58 (loc(s) Loire-Atlantique (44) based on malaise-trapping with notes on habitats) (in French).
Distribution. Labat, 2021: 92 (on Diptera taxa recorded along the over-all length of the Dordogne River and some of its tributaries; see paper for the locality details) (in French).
Distribution. Quindroit et al., 2021: 257 (recorded for l Allier (03), Cher (18) and Loiret (45)).
Distribution. Quindroit, 2021c: 424, 426 (loc(s) Mercantour National Park, Alpes-Maritimes (06), common species in France and Europe, with wide geographical distribution; ubiquitous, considered as common at all altitudes (Geiger, 1985); known from wet woodland in Northern France, to Mediterranean area (Var) and Corsica).
Distribution. Tissot et al., 2021: 208 (the Diptera of the Reserves Naturelles Nationales du Doubs (25), results of Malaise trapping since 2009) (in French).
Distribution. Quindroit and Lemoine, 2022a: 22 (loc(s) Oise (60) with notes on habitats and distr) (in French).
Distribution. Quindroit and Lemoine, 2022a: 37 (table specifying the presence in the five departments of the Hauts-de-France region).
Distribution. Quindroit, 2022a: 25-28 (additions to the annotated list of the Tipuloidea of Pays de la Loire as given in Quindroit, 2020a).
Distribution. Lemoine and Quindroit, 2023: 8 (loc(s) Jura (39)).
Distribution. Quindroit et al., 2023: 502 (questions about the presence of D. mitis in Corsica; see paper for details).
Distribution. Quindroit, 2023: 62 (loc(s) Hautes-Alpes (05)) [locality and habitat are not given in the text but with the habitat figure].
Distribution. Quindroit and Racine, 2025: 359 (table specifying the presence in the five departments of the Pays de la Loire region).
Germany
Distribution. Schacht, 1999: 135 (checklist Bayern).
Distribution. Reusch and Bellstedt, 2000: 263 (loc(s) Thuringen).
Distribution. Reusch and Oosterbroek, 2000: 161 (checklist German Bundeslander).
Distribution. Reusch and Bellstedt, 2001b: 75 (checklist Thuringen).
Distribution. Reusch et al., 2004: 117 (loc(s) Brandenburg).
Distribution. Schacht, 2005: 4 (checklist Bayern).
Distribution. Reusch and Hohmann, 2009: 128, 129 (loc(s) Sachsen-Anhalt).
Distribution. Schacht, 2010: 16 (checklist Diptera Bayern).
Distribution. Stuke, 2019: 141 (checklist Niedersachsen and Bremen with reference to original sources).
Distribution. Kehlmaier et al., 2023: 209 (loc(s) Saarland).
Great Britain
Distribution. Godfrey, 2000a: 20, 22 (see paper for details of sites surveyed in Shropshire and into Wales).
Distribution. Godfrey, 2001a: 21 (recorded from sites in Shropshire and just over the country border in Montgmeryshire; see paper for details) (as mitis s.l.).
Distribution. Rotheray, 2001: 1, 14 (loc(s) Scotland).
Distribution. Hancock, 2002: 13 (loc(s) Scotland).
Distribution. Stubbs, 2006a: 134 (loc(s), behaviour and feeding on Acer campestre leaf) (as Dicranomyia of the mitis group).
Distribution. Boardman, 2007: 77 (loc(s), map).
Distribution. Phillips, 2007: 149 (loc(s) Isle of Rum)
Distribution. Kramer, 2009f: 1 (loc(s) Yorkshire).
Distribution. Skidmore, 2009: 129 (review of occurrence on the Western Isles of Scotland, distr).
Distribution. Kramer, 2011j: 21 (loc(s) Leicestershire and Rutland (VC55)).
Distribution. Hewitt, 2014: (provisional checklist Cumbrian Diptera).
Distribution. Stary and Stubbs, 2015: 325-326 (loc(s), the species is widespread in the southern half of England).
Distribution. Robinson et al., 2016: 80 (list of terrestrial invertebrates of Mingulay Is, Outer Hebrides).
Distribution. Brighton, 2017c: 28, 38 (overall regional checklist Lancashire and Cheshire (VC58, 59 and 60) with notes on habitats and/or regional occurrence and/or status; see paper for details).
Distribution. Kramer and Morris, 2022b: 18-19 (info on the species, on the distr and map of loc(s) Leicestershire and Rutland (VC55)).
Greece
First record. Kramer, 2004: 20, 21 (loc(s) Peloponnisos).
Hungary
Distribution. Toth, 2005: 169 (loc(s)).
Ireland
Distribution. Chandler et al., 2008: 13 (checklist).
Israel
First record. Stary and Freidberg, 2007: 305, 331-332, 335-336, 339-341 (loc(s), distr).
Italy
Distribution. Podenas and Podeniene, 2008: 347, 351 (loc(s), annotated checklist South Tyrol [Prov. Bolzano]).
Lithuania
Distribution. Pakalniskis et al., 2006: 18 (checklist).
Luxembourg
Distribution. Vogtenhuber, 2007: 354 (loc(s)).
Montenegro
Distribution. Oosterbroek and Simova-Tosic, 2004: 447 (review literature).
Morocco
Distribution. Parvu et al., 2006: 273 (loc(s)).
Distribution. Gavryushin in litt., 2012d: (loc(s) High Atlas, N. Vikhrev leg.).
Distribution. Driauach et al., 2013: 188 (review Limoniidae records of Morocco, loc(s)).
Distribution. Kettani and Oosterbroek, 2022a: 25-26 (annotated checklist, with review loc(s) and/or references).
Netherlands
Distribution. Jong and Oosterbroek, 2002b: 33 (checklist).
North Macedonia
Distribution. Oosterbroek and Simova-Tosic, 2004: 447 (review literature).
Norway
Distribution. Olsen et al., 2018: 160-161 (annotated checklist Nordic countries and for Norway according to the Strand regions).
Poland
Distribution. Wiedenska, 2007a: 74 (checklist).
Distribution. Wiedenska, 2008: 46 (loc(s) Parowy Janinowski nature reserve near Lodz) (in Polish).
Distribution. Wiedenska, 2010: 131 (loc(s) Parku Krajobrazowego Wzniesien Lodzkich, Lodz).
Distribution. Wiedenska, 2014: 14 (on the occurence of Limoniidae in six selected plant communities in the Kamienica Valley, Gorce National Park, Western Carpathians, found only in Petasitetum kablikiani habitat) (in Polish, habitat tables also in English).
Distribution. Wiedenska, 2017a: 21-22 (faunistic and habitat data of species from the Gorce NP) (in Polish).
Distribution. Wiedenska and Syratt, 2023: 145 (loc(s) Bieszczady Mountains, distr and remarks).
Romania
Distribution. Parvu, 2004: 185 (loc(s)).
Distribution. Ujvarosi, 2007: 228-231 (checklist Limoniidae).
Distribution. Parvu, 2009: 448 (on some Diptera from Bucuresti and its surroundings).
Distribution. Ujvarosi et al., 2011b: 108 (loc(s) Dupa Lanca marshy area near Voslobeni) (in Hungarian, species list with habitat information in English).
Distribution. Keresztes, 2012: 30 (loc(s) Rimetea area [Alba County]).
Distribution. Kolcsar et al., 2013: 70, 71 (loc(s) Cluj-Napoca and surroundings; list of habitats with altitudes; one of the most frequent species).
Russia
Distribution. Lantsov, 2003e: 83 (loc(s) Caucasus region, distr).
Distribution. Lantsov, 2009a: 75 (loc(s) Central Caucasus, distr).
Distribution. Lantsov, 2017b: 107 (on dominant species of the crane flies (Diptera, Tipuloidea) of the North Caucasus as biological indicators of habitats) (in Russian).
Distribution. Paramonov and Korobkov, 2019: 94-95 (loc(s) RUC: Tverskaya oblast, distr, habitat, citing various authors) (in Russian).
Distribution. Kolcsar et al., 2021b: 53-54 (loc(s) RUE: Bashkortostan Rep.).
Distribution. Kolcsar et al., 2021b: 54 (loc(s) RUN: Kareliya).
Distribution. Obona et al., 2024: 12 (loc(s) RUC: Mordoviya Rep.).
Serbia
Distribution. Oosterbroek and Simova-Tosic, 2004: 447 (review literature).
Distribution. Gavryushin in litt., 2015b: (loc(s) Serbia).
Slovakia
Distribution. Stary, 2009d: 34 (loc(s) Polana area).
Distribution. Stary, 2009i: webpage (checklist).
Distribution. Stary and Stubbs, 2015: 325-326 (loc(s), distr, in the Czech Republic and Slovakia, this is the commonest and most widespread species [of the mitis-complex] occurring from high altitudes to lowland).
Slovenia
Distribution. Oosterbroek and Simova-Tosic, 2004: 447 (review literature).
Spain
Distribution. Eiroa and Baez, 2002a: 55 (checklist).
Distribution. Chvala, 2008: 157 (type-loc(s) maculosa, loc(s)).
Distribution. Hancock et al., 2015a: 241 (loc(s) Lugo).
Distribution. Mederos and Eiroa, 2015: 266 (loc(s) Lerida).
Distribution. Keresztes et al., 2022: 195 (loc(s) Asturias and review distribution in Spain) (in Spanish).
Switzerland
Distribution. Podenas et al., 2006: 158, 224 (map, vertical distr per thermic level).
Distribution. Dufour and Merz, 2012: 354 (annotated checklist canton Geneva).
Turkey
First record. Koc, 2004: 103, 111, 116-118, 153 (loc(s), distr, map) (in Turkish).
Distribution. Ozgul et al., 2009: 63 (survey provinces SW Turkey).
Distribution. Bilgin et al., 2015: 6 (loc(s) Eskisehir, distr) (in Turkish).
Distribution. Koc et al., 2016: 11 (provinces Marmara region).
Distribution. Akpinar and Ozgul, 2023: 55 (loc(s) Aksaray).
Distribution. Provinces Asiatic part, 2025: (known from Aydin, Denizli, Eskisehir, Isparta, Mugla, Marmara region).
on flight period (mainly from 2000 onward):
Austria
Aistleitner, 2015: month(s): 5.
Stary and Stubbs, 2015: month(s): 5.
Belarus
Kolcsar et al., 2021b: month(s): 5.
Belgium
Dek and Peeters, 2023a: month(s): 5-6, 10.
Bosnia-Herzegowina
Chvala, 2008: month(s): 4.
Bulgaria
Hubenov, 2015: month(s): 6, 8.
Stary and Stubbs, 2015: month(s): 5.
Hubenov, 2016: month(s): 6, 8.
Croatia
Kolcsar et al., 2015a: month(s): 5-7.
Kolcsar et al., 2023b: month(s): 6.
Czech Rep.
Stary et al., 2005a: month(s): 5-6.
Stary and Stubbs, 2015: month(s): 5-6.
Stary and Vonicka, 2018: month(s): 5-6(7).
France
Kramer and Withers, 2007: month(s): 5.
Quindroit, 2020a: month(s): 4-5.
Quindroit, 2020b: month(s): 4-6(8).
Quindroit and Lemoine, 2022a: month(s): 6.
Quindroit, 2023: month(s): 6.
Germany
Reusch and Bellstedt, 2000: month(s): 5-7.
Reusch et al., 2004: month(s): 5-6.
Kehlmaier et al., 2023: month(s): 6.
Great Britain
Godfrey, 2000a: month(s): 6.
Godfrey, 2001a: month(s): 5.
Hancock, 2002: month(s): 5.
Stubbs, 2006a: month(s): 5.
Phillips, 2007: month(s): 8.
Kramer, 2009f: month(s): 5.
Kramer, 2011j: month(s): 5-6.
Stary and Stubbs, 2015: month(s): 5-6.
Greece
Kramer, 2004: month(s): 4.
Israel
Stary and Freidberg, 2007: month(s): 3.
Italy
Podenas and Podeniene, 2008: month(s): 6.
Morocco
Parvu et al., 2006: month(s): 3.
Gavryushin in litt., 2012d: month(s): 5.
Poland
Wiedenska, 2008: month(s): 6.
Wiedenska, 2014: month(s): 6-7.
Wiedenska, 2015a: month(s): 6-8.
Wiedenska, 2017a: month(s): 6-8, 10.
Wiedenska and Syratt, 2023: month(s): 6-8.
Romania
Parvu, 2004: month(s): 7.
Parvu, 2009: month(s): 4.
Ujvarosi et al., 2011b: month(s): 5.
Kolcsar et al., 2013: month(s): 4-6, 9.
Russia
Lantsov, 2003e: month(s): 7, 9.
Lantsov, 2009a: month(s): 9.
Paramonov and Korobkov, 2019: month(s): 5.
Kolcsar et al., 2021b: month(s): 8.
Kolcsar et al., 2021b: month(s): 6.
Obona et al., 2024: month(s): 6.
Serbia
Gavryushin in litt., 2015b: month(s): 5, 7.
Slovakia
Stary, 2009d: month(s): 5-6.
Stary and Stubbs, 2015: month(s): 5-6.
Spain
Chvala, 2008: month(s): 5.
Hancock et al., 2015a: month(s): 5.
Keresztes et al., 2022: month(s): 8.
Switzerland
Podenas et al., 2006: month(s): 4-11.
Turkey
Koc, 2004: month(s): 5.
on altitude (mainly from 2000 onward):
Austria
Aistleitner, 2015: altitude: 435 m.
Bulgaria
Hubenov, 2015: altitude: 1300-1900 m.
Hubenov, 2016: altitude: 1147-1850 m.
Hubenov, 2017: altitude: 1147-1900 m.
Hubenov, 2018a: altitude: 1400-1420 m.
Hubenov, 2018b: altitude: 380 m.
Hubenov, 2021a: altitude: 350-1900 m.
Hubenov, 2025a: altitude: 350-1900 m.
Hubenov, 2025b: altitude: 350-1900 m.
Croatia
Kolcsar et al., 2015a: altitude: 265-715 m.
Kolcsar et al., 2023b: altitude: 720 m.
Czech Rep.
Stary and Stubbs, 2015: altitude: 500-1330 m.
France
Labat, 2021: altitude: 1-119 m.
Quindroit, 2021c: altitude: 2000 m.
Lemoine and Quindroit, 2023: altitude: 5 m.
Quindroit, 2023: altitude: 1900 m.
Germany
Reusch et al., 2004: altitude: 20-90 m.
Reusch and Hohmann, 2009: altitude: 435-842 m.
Greece
Kramer, 2004: altitude: 875 m.
Italy
Podenas and Podeniene, 2008: altitude: 1930 m.
Morocco
Gavryushin in litt., 2012d: altitude: 1000-2600 m.
Driauach et al., 2013: altitude: 490-735 m.
Poland
Wiedenska, 2017a: altitude: 600-1180 m.
Wiedenska and Syratt, 2023: altitude: 650-800 m.
Russia
Lantsov, 2003e: altitude: 510-914 m.
Lantsov, 2009a: altitude: 1571-1600 m.
Kolcsar et al., 2021b: altitude: 607 m.
Kolcsar et al., 2021b: altitude: 5 m.
Serbia
Gavryushin in litt., 2015b: altitude: 800-1030 m.
Slovakia
Stary, 2009d: altitude: 630-780 m.
Stary and Stubbs, 2015: altitude: 700-850 m.
Spain
Mederos and Eiroa, 2015: altitude: 1300-1450 m.
Keresztes et al., 2022: altitude: 496 m.
Turkey
Koc, 2004: altitude: 641-700 m.

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