Details for Dicranomyia (Dicranomyia) didyma
Name:Dicranomyia (Dicranomyia) didyma (Meigen, 1804)
Publication:Klass. Beschr. 1: 55 (as Limonia)
Status:Recognized taxon, Synonym(s): trinotata (Meigen and Wiedemann, 1818); oscillans Haliday, 1833.
Classification:Family Limoniidae
Subfamily Limoniinae
Keys:Podenas and Gelhaus, 2007 (Limoniinae Mongolia and nearby); Peeters en Oosterbroek, 2014c (craneflies Netherlands, Belgium and Luxembourg) (in Dutch); Boardman, 2016 (craneflies Shropshire); Stubbs and Kramer, 2016h; Stubbs, 2021 (British craneflies)
Variant spelling(s):dydima
Dicranomyia (Dicranomyia) didyma : habitus - maleDicranomyia (Dicranomyia) didyma : habitus - maleDicranomyia (Dicranomyia) didyma : habitus - maleDicranomyia (Dicranomyia) didyma : habitus - maleDicranomyia (Dicranomyia) didyma : habitus - female
habitus
male
habitus
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habitus
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habitus
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habitus
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plus 21 more images of body part(s), hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, wing, wing, wing, wing, habitat, habitat, habitat, habitat, habitat, habitat, habitat, habitat, habitat and habitat.
Distribution:Albania, Austria, Belgium, Bosnia-Herzegovina, Bulgaria, Croatia, Czech Rep., Denmark (incl. Faroe Is), Estonia, Finland, France (incl. Corsica), Germany, Great Britain, Hungary, Iceland, Ireland, Italy, Latvia, Lithuania, Montenegro, Netherlands, North Macedonia, Norway, Poland, Portugal, Romania, Serbia, Slovakia, Slovenia, Spain, Sweden, Switzerland, Ukraine; Russia: RUN, RUW, RUC, RUE, North Caucasus; Morocco, Algeria; Georgia, Armenia, Azerbaijan, Turkey (Asiatic part: see citations for provinces), Iran;; Afghanistan; Mongolia.
Note: see the manual for abbreviations: present-day Russia.
Region(s):Westpalaearctic;; Eastpalaearctic
General note:Added: Algeria (Vaillant, 1952: 248-249, as mitis, confirmed by Stary in litt., june 2013). For Italy check Stary and Oosterbroek, 1996, and citations below. For Finland check Salmela, 2012b, 2012c, and citations below. For Norway check Olsen et al., 2018, and citations below. For Portugal check Oosterbroek et al., 2020, and citations below. For Great Britain and Ireland check Stubbs, 2021, and citations below. For Morocco check Kettani and Oosterbroek, 2022a, and citations below. To be confirmed for Russia: RUE (Middle Volga region) (in Chuzhekova, 2015, based on larvae only) and China: Shanxi.
Map:Podenas et al., 2006 (Switzerland); Boardman, 2007 (Shropshire); Przhiboro, 2018c (St Petersburg region)
Biology:Alexander, 1920r (as trinotata Meigen) (mating behaviour, habitat, short description immature stages).
Figure
references:
Habitus: Przhiboro, 2018c (male)
Wing: Podenas and Gelhaus, 2007
Hypopygium: Alexander, 1975e; Podenas et al., 2006; Podenas and Gelhaus, 2007; Kramer and Morris, 2022b

Immature stages
Larva: Podeniene, 2003a; Werner et al., 2007 (larvae in aggregations of larvae and pupae Simulium); Przhiboro, 2018c

Miscellaneous
Werner et al., 2007 (adult prayed upon by adult Limnophora (Muscidae); habitat)
Citationson biology (mainly from 2000 onward):
Azerbaijan
Habitat. Stary and Obona, 2020: 135 (loc(s); with list of localities, habitats and altitudes).
Czech Rep.
Habitat. Stary and Vonicka, 2018: 63 (on Limoniidae and Pediciidae of two localities in northern Bohemia, with list of localities, altitudes and habitats).
Finland
Habitat. Salmela, 2004: 9 (brooks and small river, very rarely recorded from springs).
Biology. Salmela, 2008: 11, 28, 51 (49) (ecology or habitat: lotic; phenology).
Habitat. Salmela, 2011a: 1-28 (study on three wetland habitats: aapa mires, springs and headwater streams; found in headwater streams only).
Georgia
Habitat. Stary and Obona, 2020: 135 (loc(s); with list of localities, habitats and altitudes).
Germany
Habitat. Reusch, 2006: 134 (Kalkquellmoor [carbonate fen complex]).
Biology. Werner et al., 2007: 101-111 (early larval stages are strictly detritivorous, older larvae are also predaceous, preying on larvae and pupae of Chironomidae and Simuliidae; larval feeding behaviour; adult mating behaviour; oviposition; adults are prayed upon by adult Limnophora (Muscidae), incl. photo; habitat, incl. photo; univoltine but under optimal conditions possibly bivoltine).
Habitat. Reusch and Hohmann, 2009: 129 (from spring brooks).
Habitat. Dunk, 2018: 79, 96 (loc(s) Bayern with list of habtats).
Great Britain
Habitat. Godfrey, 1999: 79 (can be found on exposed riverine sediments).
Habitat. Godfrey, 2000a: 13, 22 (waterfalls and vertical seepages).
Habitat. Godfrey, 2001a: 21.
Habitat. Boardman, 2007: 76 (wooded mossy streams, dingle woodlands).
Habitat. Kramer, 2008b: 7 (near the source of a stream).
Habitat. Kramer, 2009c: 6 (founf at mossy fringes of waterfalls).
Habitat. Cranston and Drake, 2010: 175 (larvae can be found in marginal situations along flowing waters and among aquatic or sub-aquatic mosses).
Habitat. Kramer, 2011j: 21 (by waterfalls, lays it eggs in wet moss).
Habitat. Boardman, 2013a: 2 (found on the splashzone at the outflow of pools).
Habitat. Kramer, 2013i: 7 (occurs near waterfalls and the larvae feed in surrounding damp mosses, also known from wet woodlands).
Biology. Kramer, 2016c: 3 (a common but local species of waterfalls, laying its eggs in adjacent wet moss).
Habitat. Kramer and Morris, 2022b: 17 (info on the species, on the distr and map of loc(s) Leicestershire and Rutland (VC55); seems to be found near running water, especially waterfalls, where the eggs are laid in damp moss).
Ireland
Habitat. Ashe et al., 2008: 9 (associated with running water).
Lithuania
Habitat. Podeniene, 2003a: 1-295 (larva, descr, figs, habitat) (in Lithuanian).
Mongolia
Habitat. Yadamsuren et al., 2015: 474 (terrestrial).
Poland
Habitat. Wiedenska, 2017a: 21 (faunistic and habitat data of species from the Gorce NP) (in Polish).
Romania
Habitat. Ujvarosi, 2005a: 244 (springs, swamps, moist soil, citing various sources).
Russia
Habitat. Lantsov, 2009a: 74 (ecology) (in Russian).
Habitat. Paramonov and Pilipenko, 2016: 99 (habitat, citing Savchenko, 1985) (in Russian).
Biology. Przhiboro, 2018c: 32, 387 (Red data book St Petersburg; figs larva, adult male, short descr, distr, local records with map, habitats incl larval habitats, rearing, data on bionomics, proposals for protection of local populations) (in Russian).
Spain
Habitat. Hancock and Hewitt, 2020: 45-49 (in the Cazorla Parque Natural, Jaen Province, Spain, in june 2019, an area of cliff face was seen covered with a butterwort, Pinguicula vallisneriifolia (Webb), the leaves of which had trapped large numbers of insects, including 19 species of craneflies; the effectiveness of the leaf glandular secretions in relation to immobilising different sizes of flies is discussed; see paper for details, habitat figs).
Habitat. Hancock, 2020: 95 (common near river bank and surrounding areas, also found attached to leaves of butterwort (Pinguicula vallisneriifolia Webb)) (see also Hancock and Hewitt, 2020).
on characters and taxonomy (mainly from 2000 onward):
Country not relevant
Phylogeny. Boldgiv, 2006: 96-154 (one of the 84 morphological taxa from northern Mongolia in a consistency test of morphology based species identification of crane flies against DNA-barcoding identification using 536-base-pair sequence profiles of the COI gene, also known as the DNA barcode).
Characters. Stary, 2006b: 201 (comparison).
Characters. Stary and Freidberg, 2007: 336, 341 (comparison).
Characters. Stary, 2009a: 56 (comparison).
Phylogeny. Salmela et al., 2014: 4, 15 (DNA-barcoding, phylogeny).
Great Britain
Characters. Kramer, 2016c: 3 (species look a-likes: D. didyma and D. consimilis; these two species have a similar appearnce, the wings having a rom of 4 dark spots along the costal edge and could easily be confused).
Characters. Kramer and Morris, 2022b: 17 (diagnosis, figs).
Lithuania
Immatures. Podeniene, 2003a: 1-295 (larva, descr, figs, habitat) (in Lithuanian).
Mongolia
Key. Podenas and Gelhaus, 2007: 42 (key, figs).
Portugal
Characters. Ferreira et al., 2021: 8 (barcoding based on material mentioned in Oosterbroek et al. 2020).
Russia
Immatures. Krivosheina, N.P., 2009d: 140 (comparison larva).
Description. Przhiboro, 2018c: 32, 387 (Red data book St Petersburg; figs larva, adult male, short descr, distr, local records with map, habitats incl larval habitats, rearing, data on bionomics, proposals for protection of local populations) (in Russian).
on distribution (mainly from 2000 onward):
Algeria
First record. Vaillant, 1952: 248-249 (as mitis, confirmed by Stary in litt., june 2013).
Armenia
Distribution. Obona et al., 2016: 134, 140 (loc(s), checklist).
Distribution. Hakobyan and Jenderedjian, 2023: 1513 (annotated checklist of the craneflies of Armenia with refs, distr and flight period in Caucasus Ecoregion with refs).
Austria
Distribution. Reusch and Heiss, 2012: 314-322 (loc(s) Nat. Park Gesause, see PDF for months and altitudes).
Azerbaijan
Distribution. Stary and Obona, 2020: 135 (loc(s); with list of localities, habitats and altitudes).
Distribution. Snegovaya, 2021: 1135-1136 (checklist with details on distr in Azerbaijan).
Bosnia-Herzegowina
Distribution. Oosterbroek and Simova-Tosic, 2004: 447 (review literature).
Distribution. Kolcsar et al., 2023b: 12 (loc(s)).
Bulgaria
Distribution. Hubenov, 2018b: 124 (review loc(s) Vrachanska Planina Mts).
Distribution. Hubenov, 2021a: 36 (checklist with summary of distr in Bulgaria, references).
Croatia
Distribution. Oosterbroek and Simova-Tosic, 2004: 447 (review literature).
Distribution. Kolcsar et al., 2015a: 31-32 (loc(s), distr).
Distribution. Kolcsar et al., 2023b: 12-14 (loc(s)).
Czech Rep.
Distribution. Stary et al., 2005a: 25 (loc(s)).
Distribution. Stary, 2009i: webpage (checklist).
Distribution. Stary and Vonicka, 2018: 63 (on Limoniidae and Pediciidae of two localities in northern Bohemia, with list of localities, altitudes and habitats).
Denmark
Distribution. Petersen and de Jong, 2001b: 148 (checklist).
Estonia
Distribution. Stary, 2004f: (Fauna Europaea: added Estonia).
Finland
Distribution. Salmela, 2001a: 144, 151 (loc(s)).
Distribution. Salmela, 2004: 9 (loc(s)).
Distribution. Stary and Salmela, 2004: 47 (loc(s)).
Distribution. Salmela et al., 2007b: 45 (loc(s)).
Distribution. Salmela, 2008: 11, 51 (49) (loc(s), known from all Finnish ecoregions).
Distribution. Salmela, 2011a: 24 (loc(s) subalpine northern boreal Finland).
Distribution. Salmela, 2012b: 236 (annotated list of Finnish crane flies).
Distribution. Salmela and Petrasiunas, 2014: 29 (checklist Finnish Tipulomorpha).
Distribution. Salmela et al., 2014: 4 (loc(s)).
Distribution. Salmela et al., 2015: 100 (list of localities in extensive report on malaise-trapping of insects in conservation areas in Lapland in 2012–2014) (in Finnish).
France
Distribution. Kramer and Langlois, 2019b: 228 (list of craneflies captured 25-27 june 2019 in Ravin de Valbois, Doubs (25)).
Distribution. Tissot et al., 2021: 208 (the Diptera of the Reserves Naturelles Nationales du Doubs (25), results of Malaise trapping since 2009) (in French).
Georgia
Distribution. Stary and Obona, 2020: 135 (loc(s); with list of localities, habitats and altitudes).
Germany
Distribution. Schacht, 1999: 135 (checklist Bayern).
Distribution. Reusch and Oosterbroek, 2000: 161 (checklist German Bundeslander).
Distribution. Reusch and Bellstedt, 2001b: 74 (checklist Thuringen).
Distribution. Schacht, 2005: 4 (checklist Bayern).
Distribution. Reusch, 2006: 134 (loc(s)).
Distribution. Werner et al., 2007: 102 (loc(s)).
Distribution. Reusch and Hohmann, 2009: 129 (loc(s) Sachsen-Anhalt).
Distribution. Schacht, 2010: 16 (checklist Diptera Bayern).
Distribution. Dunk, 2018: 79, 96 (loc(s) Bayern with list of habtats).
Distribution. Stuke, 2019: 141 (checklist Niedersachsen and Bremen with reference to original sources).
Great Britain
Distribution. Godfrey, 2000a: 13, 22 (loc(s)).
Distribution. Godfrey, 2001a: 21 (loc(s)).
Distribution. Hewitt et al., 2005: 37 (loc(s) Cumbria) (as Limonia).
Distribution. Boardman, 2007: 76 (loc(s), map).
Distribution. Kramer, 2008b: 7 (loc(s) Wales).
Distribution. Skidmore, 2009: 128 (review of occurrence on the Western Isles of Scotland, distr).
Distribution. Kramer, 2011j: 21 (loc(s) Leicestershire and Rutland (VC55)).
Distribution. Boardman, 2013a: 2 (loc(s) Shropshire).
Distribution. Kramer, 2013g: 2 (loc(s) Lancaster).
Distribution. Kramer, 2013i: 8 (checklist of species from Leigh Woods NNR, Bristol district).
Distribution. Hewitt, 2014: (provisional checklist Cumbrian Diptera).
Distribution. Halstead, 2016: 11 (loc(s) Surrey).
Distribution. Kramer, 2016c: 3 (a common but local species of waterfalls, laying its eggs in adjacent wet moss).
Distribution. Brighton, 2017c: 28 (overall regional checklist Lancashire and Cheshire (VC58, 59 and 60) with notes on habitats and/or regional occurrence and/or status; see paper for details).
Distribution. Showers, 2017a: 4 (loc(s) Northamptonshire).
Distribution. Kramer, 2018f: 1 (loc(s) Staffordshire).
Distribution. Kramer and Morris, 2022b: 17 (info on the species, on the distr and map of loc(s) Leicestershire and Rutland (VC55); seems to be found near running water, especially waterfalls, where the eggs are laid in damp moss).
Iceland
Distribution. Olsen in litt., 2018: (collected in Iceland in july 2018).
Ireland
Distribution. Ashe et al., 2008: 9 (additional Irish record(s), distr).
Distribution. Chandler et al., 2008: 12 (checklist).
Lithuania
Distribution. Pakalniskis et al., 2006: 18 (checklist).
Mongolia
Distribution. Boldgiv, 2006: 153 (loc(s)).
First record. Podenas and Gelhaus, 2007: 40 (new for Mongolia, without further details).
Distribution. Yadamsuren et al., 2015: 474 (loc(s) Mongolia).
Montenegro
Distribution. Oosterbroek and Simova-Tosic, 2004: 447 (loc(s), review literature).
Morocco
Distribution. Gavryushin in litt., 2012d: (loc(s) High Atlas, N. Vikhrev leg.).
Distribution. Driauach et al., 2013: 188 (review Limoniidae records of Morocco, loc(s)).
Distribution. Kettani and Oosterbroek, 2022a: 25 (annotated checklist, with review loc(s) and/or references).
Netherlands
Distribution. de Jong and Oosterbroek, 2002b: 33 (checklist).
North Macedonia
Distribution. Oosterbroek and Simova-Tosic, 2004: 447 (review literature).
Norway
Distribution. Olsen et al., 2018: 160-161 (annotated checklist Nordic countries and for Norway according to the Strand regions).
Poland
Distribution. Wiedenska, 2007a: 74 (checklist).
Distribution. Wiedenska, 2010: 131 (loc(s) Parku Krajobrazowego Wzniesien Lodzkich, Lodz).
Distribution. Wiedenska, 2017a: 21 (faunistic and habitat data of species from the Gorce NP) (in Polish).
Distribution. Wiedenska and Syratt, 2023: 145 (loc(s) Bieszczady Mountains, distr and remarks).
Portugal
Distribution. Eiroa and Baez, 2002a: 55 (checklist).
Distribution. Oosterbroek et al., 2020: 329 (loc(s) Braganca, Aveiro and Guarda, distr).
Romania
Distribution. Ujvarosi, 2007: 228-231 (checklist Limoniidae).
Distribution. Keresztes, 2012: 30 (loc(s) Rimetea area [Alba County]).
Russia
Distribution. Lantsov, 2009a: 74 (loc(s) Central Caucasus, distr).
Distribution. Gavryushin in litt., 2011e: (loc(s) RUN: Murmanskaya oblast).
Distribution not confirmed. Chuzhekova, 2015: 211 (on macrozoobenthos communities in spring brooks of Middle Volga) (PhD thesis, in Russian, 242 pages; for English summary see Chuzhekova, 2016, 37 pages).
Distribution. Paramonov and Pilipenko, 2016: 99 (loc(s) RUC: Tverskaya oblast, distr) (in Russian).
Distribution. Przhiboro, 2018c: 32, 387 (Red data book St Petersburg; figs larva, adult male, short descr, distr, local records with map, habitats incl larval habitats, rearing, data on bionomics, proposals for protection of local populations) (in Russian).
Distribution. Kolcsar et al., 2021b: 36 (loc(s) RUE: Samarskaya oblast).
Serbia
Distribution. Oosterbroek and Simova-Tosic, 2004: 447 (loc(s), review literature).
Distribution. Gavryushin in litt., 2015b: (loc(s) Serbia).
Slovakia
Distribution. Stary, 2009d: 34 (loc(s) Polana area).
Distribution. Stary, 2009i: webpage (checklist).
Slovenia
Distribution. Oosterbroek and Simova-Tosic, 2004: 447 (review literature).
Spain
Distribution. Eiroa and Baez, 2002a: 55 (checklist).
Distribution. Mederos and Eiroa, 2015: 266 (loc(s) Lerida).
Distribution. Hancock and Hewitt, 2020: 45-49 (in the Cazorla Parque Natural, Jaen Province, Spain, in june 2019, an area of cliff face was seen covered with a butterwort, Pinguicula vallisneriifolia (Webb), the leaves of which had trapped large numbers of insects, including 19 species of craneflies; the effectiveness of the leaf glandular secretions in relation to immobilising different sizes of flies is discussed; see paper for details, habitat figs).
Distribution. Hancock, 2020: 95 (loc(s) Jaen, Cazorla National Park).
Distribution. Keresztes et al., 2022: 194 (loc(s) Asturias and review distribution in Spain) (in Spanish).
Sweden
Distribution. Salmela, 2010b: 133 (loc(s) Malmo).
Switzerland
Distribution. Podenas et al., 2006: 155, 215 (map, vertical distr per thermic level).
Distribution. Stary, 2009a: 68 (loc(s)).
Distribution. Dufour and Merz, 2012: 354 (annotated checklist canton Geneva).
Turkey
First record. Ozgul and Koc, 2012: 749 (first record for Turkey, prov. Afyonkarahisar and Usak).
Distribution. Bilgin et al., 2015: 6 (loc(s) Eskisehir, distr) (in Turkish).
Distribution. Akpinar and Ozgul, 2023: 55 (loc(s) Aksaray).
Distribution. Provinces Asiatic part, 2024: (known from Afyonkarahisar, Eskisehir, Usak).
on flight period (mainly from 2000 onward):
Armenia
Obona et al., 2016: month(s): 8-9.
Azerbaijan
Stary and Obona, 2020: month(s): 5, 10.
Bosnia-Herzegowina
Kolcsar et al., 2023b: month(s): 4.
Croatia
Kolcsar et al., 2015a: month(s): 2-12.
Kolcsar et al., 2023b: month(s): 5-10.
Czech Rep.
Stary and Vonicka, 2018: month(s): 5-7, 9.
Finland
Salmela, 2001a: month(s): 8.
Salmela, 2008: month(s): 8.
France
Kramer and Langlois, 2019b: month(s): 6.
Georgia
Stary and Obona, 2020: month(s): 7.
Germany
Werner et al., 2007: month(s): 5-7.
Dunk, 2018: month(s): 6.
Great Britain
Godfrey, 2000a: month(s): 6.
Godfrey, 2001a: month(s): 5.
Kramer, 2008b: month(s): 7.
Kramer, 2011j: month(s): 5-6.
Boardman, 2013a: month(s): 8.
Kramer, 2013g: month(s): 7.
Halstead, 2016: month(s): 6.
Kramer, 2018f: month(s): 6.
Kramer and Morris, 2022b: month(s): 6-9.
Ireland
Ashe et al., 2008: month(s): 6.
Morocco
Gavryushin in litt., 2012d: month(s): 5.
Poland
Wiedenska, 2017a: month(s): 7-9.
Wiedenska and Syratt, 2023: month(s): 6-8.
Portugal
Oosterbroek et al., 2020: month(s): 3, 5.
Russia
Lantsov, 2009a: month(s): 9.
Gavryushin in litt., 2011e: month(s): 7.
Paramonov and Pilipenko, 2016: month(s): 7.
Kolcsar et al., 2021b: month(s): 7.
Serbia
Gavryushin in litt., 2015b: month(s): 7.
Slovakia
Stary, 2009d: month(s): 6-7, 9.
Spain
Hancock and Hewitt, 2020: month(s): 6.
Hancock, 2020: month(s): 5-6.
Keresztes et al., 2022: month(s): 8.
Switzerland
Podenas et al., 2006: month(s): 5-11.
Stary, 2009a: month(s): 7.
Turkey
Ozgul and Koc, 2012: month(s): 4-5.
on altitude (mainly from 2000 onward):
Armenia
Obona et al., 2016: altitude: 976-2195 m.
Azerbaijan
Stary and Obona, 2020: altitude: 825-1510 m.
Bosnia-Herzegowina
Kolcsar et al., 2023b: altitude: 308 m.
Bulgaria
Hubenov, 2018b: altitude: 380 m.
Hubenov, 2021a: altitude: 200-400 m.
Croatia
Kolcsar et al., 2015a: altitude: 45-720 m.
Kolcsar et al., 2023b: altitude: 545-720 m.
Georgia
Stary and Obona, 2020: altitude: 1565-2250 m.
Germany
Reusch and Hohmann, 2009: altitude: 375-435 m.
Great Britain
Kramer, 2008b: altitude: 500 m.
Morocco
Gavryushin in litt., 2012d: altitude: 1750-2600 m.
Kettani and Oosterbroek, 2022a: altitude: 2000 m.
Poland
Wiedenska, 2017a: altitude: 450-1000 m.
Wiedenska and Syratt, 2023: altitude: 470-800 m.
Russia
Lantsov, 2009a: altitude: 1571-1659 m.
Kolcsar et al., 2021b: altitude: 30 m.
Serbia
Gavryushin in litt., 2015b: altitude: 800 m.
Slovakia
Stary, 2009d: altitude: 630-780 m.
Spain
Mederos and Eiroa, 2015: altitude: 1450 m.
Keresztes et al., 2022: altitude: 1236 m.
Switzerland
Stary, 2009a: altitude: 340 m.
Turkey
Ozgul and Koc, 2012: altitude: 680-1022 m.

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