| Name: | Pilaria fuscipennis (Meigen, 1818) | | Publication: | Syst. Beschr. 1: 125 (as Limnobia) | | Status: | Recognized taxon. | | Classification: | Family Limoniidae Subfamily Limnophilinae
| | Keys: | Boardman, 2016 (craneflies Shropshire); Peeters en Oosterbroek, 2016c (craneflies Netherlands, Belgium and Luxembourg) (in Dutch); Stubbs and Kramer, 2016g; Stubbs, 2021 (British craneflies) |  |  |  |  |  |
habitus male | habitus male | habitus male | habitus female | body part(s) antenna |
plus 17 more images of body part(s), body part(s), body part(s), body part(s), body part(s), hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, ovipositor, wing, wing, wing, wing, habitat and habitat. | | | | Distribution: | Austria, Belarus, Belgium, Bulgaria, Czech Rep., Denmark, Estonia, France (incl. Corsica), Germany, Great Britain, Greece, Hungary, Ireland, Italy, Latvia, Lithuania, Netherlands, North Macedonia, Norway, Poland, Romania, Slovakia, Sweden, Switzerland, Turkey (European part: Kirklareli), Ukraine; Russia: RUW, RUC, North Caucasus; Turkey (Asiatic part: see citations for provinces);; Russia: WS (Altay). Note: see the manual for abbreviations: present-day Russia. | | Region(s): | Westpalaearctic;; Eastpalaearctic | | General note: | For Italy check Stary and Oosterbroek, 1996, and citations below. For Norway check Olsen et al., 2018, and citations below. For Great Britain and Ireland check Stubbs, 2021, and citations below. Removed: Finland (Salmela, 2012b). To be confirmed for Far East Russia (see Alexander, 1925n). | | Map: | Ashe et al., 2005a (Ireland); Podenas et al., 2006 (Switzerland); Boardman, 2007 (Shropshire) | Figure references: | Head: Drake and Stubbs, 2014 (antenna) Hypopygium: Podenas et al., 2006; Drake and Stubbs, 2014 Other figures: Drake and Stubbs, 2014 (head, antennae, thorax); Kramer and Morris, 2022b (head, antennae, thorax)
Immature stages Larva: Podeniene, 2003a
| | | | Citations | | | Bulgaria Habitat. Hubenov, 2025a: suppl: 3 (annotated list Diptera Stara Planina Mts, local distr, habitats and zoogeography). Habitat. Hubenov, 2025b: suppl: 3 (annotated list Diptera high Bulgarian mts, local distr, habitats and zoogeography). | | Country not relevant Biology. Olsen et al., 2018: 145 (adults are found at seepages in marsh, carr and wet woodland, along flowing and standing waters, avoiding open sunny situations (Boardman 2007, 2016, Drake & Stubbs 2014, Godfrey 2000); larvae live in mud of bogs, woody swamps and along margins of flowing or standing water (Cranston & Drake 2010, Mendl 1978, Podenas & Podeniene 2008, Podeniene 2001, 2003); in Norway collected at a swamp forest near small tarn, with ferns and horsetails; the period of flight is june-august). | | Czech Rep. Habitat. Stary and Vonicka, 2018: 52 (on Limoniidae and Pediciidae of two localities in northern Bohemia, with list of localities, altitudes and habitats). | | Denmark Habitat. Byriel and Rojas, 2017: 118 (among the 101 species in a study on cranefly hotspots in unmanaged and managed forests) (in Danish). | | France Biology. Quindroit, 2020a: 33, 45-48 (annotated list with information on abundancy and habitat of the Tipuloidea of the Pays de la Loire regoin, distr) (in French). Habitat. Herbrecht et al., 2021: 57 (loc(s) Loire-Atlantique (44) based on malaise-trapping with notes on habitats) (in French). Habitat. Quindroit and Lemoine, 2021a: 25 (loc(s), resultats de deux campagnes de piegeage dans le Pas-de-Calais (62); see paper for details on habitat). Habitat. Quindroit, 2022d: 417 (loc(s) Ariege (09); zone de suintement boueux en bordure de sous-bois). | | Germany Habitat. Ruckert, 2005: 47, 53 (found in a study on the association with the type of source: limnokrene, rheokrene, or helokrene in Schlesig-Holstein; see paper for details, as well as Anhang A2-A10). Habitat. Wittrock, 2005: 85 (summer emergence; helokrene and rheokrene). Habitat. Ruckert et al., 2007: 56 (Limnophil). | | Great Britain Habitat. Godfrey, 2000a: 13, 22 (woodland seepages, wet woodland and carr). Habitat. Howe et al., 2001: 139, 143-146. Habitat. Stubbs, 2003: 52 (carr). Habitat. Boardman, 2007: 72 (seepages in carr and wet woodland). Habitat. Drake, 2008b: 168 (collected in a small wet woodland with base rich seepages in Dorset). Habitat. Cranston and Drake, 2010: 175 (larvae can be found in marginal situations along flowing and standing waters). Habitat. Drake, 2011a: 12, 14, 25 (among the Diptera from a wet Alder [Alnus] wood in Devon). Habitat. Drake and Stubbs, 2014: 184 (species of wet woodland, especially where groundwater saepages are present; it avoids open sunny situations but exceptionally has been found where tall vegetation provides sufficient shade). Habitat. Showers, 2015: 2 (on craneflies in moth traps at Pitsford Water Nature Reserve in 2013 and 2014, including information for the two traps and photos of their habitat). Habitat. Brighton, 2017c: 28, 37 (overall regional checklist Lancashire and Cheshire (VC58, 59 and 60) with notes on habitats and/or regional occurrence and/or status; see paper for details). | | Greece Habitat. Kolcsar et al., 2021b: 215 (brook). | | Italy Habitat. Podenas and Podeniene, 2008: 352 (larvae developing in rich organic mud near water reservoirs). | | Lithuania Habitat. Podeniene, 2001a: 84 (larvae were found in the intertidal zone of a small river in silt and leaf fallings). Habitat. Podeniene, 2003a: 1-295 (larva, descr, figs, habitat) (in Lithuanian). Habitat. Podeniene, 2003b: 10, 22, 28 (larval habitat: mud in bogs of low lying woody swamps). | | Poland Habitat. Wiedenska, 2014: 12 (on the occurence of Limoniidae in six selected plant communities in the Kamienica Valley, Gorce National Park, Western Carpathians) (in Polish, habitat tables also in English). Biology. Wiedenska, 2015a: 52 (on the flight activity of Limoniidae in the Kamienica Valley, Gorce National Park, Western Carpathians; observed emergence of adults in july) (in Polish, tables also in English). Habitat. Wiedenska, 2017a: 14 (faunistic and habitat data of species from the Gorce NP) (in Polish). | | Romania Habitat. Kolcsar et al., 2013: 73 (loc(s) Cluj-Napoca and surroundings; list of habitats with altitudes). | | Sweden Habitat. Lindstrom and Fritz, 2015: 44, 48 (distributin and ecology in five fens in Hallands County) (in Schwedish). | | Turkey Habitat. Koc, 2008: 4 (collected under marshy conditions at a grassy area by a lake and by a small creek). | | | | Country not relevant Characters. Drake and Stubbs, 2014: 184-186 (comparison, figs). Characters. Olsen et al., 2018: 144-145 (differs from other Pilaria species by its entirely orange thorax, entirely yellow femora and male apodeme without a bulbous outgrowth; fig). | | Great Britain Characters. Kramer and Morris, 2022b: 13 (diagnosis, figs). | | Lithuania Immatures. Podeniene, 2003a: 1-295 (larva, descr, figs, habitat) (in Lithuanian). | | | | Belarus First record. Kolcsar et al., 2021b: 214-215 (loc(s) Minsk region). | | Belgium Distribution. Dek and Peeters, 2023a: 139 (loc(s) Botanic Garden Jean Massart, Brussels-Capital Region, very rare in Belgium, the Jardin being the only recent locality). | | Bulgaria Distribution. Hubenov, 2015: 220 (annotated list Diptera Pirin Mt, loc(s), faunistics and zoogeography). Distribution. Hubenov, 2017: 68 (on vertical distribution in the Pirin and Rila Mts; see paper for details on forests and subalpine-alpine zones, distr). Distribution. Hubenov, 2018b: 123 (review loc(s) Vrachanska Planina Mts). Distribution. Hubenov, 2021a: 31 (checklist with summary of distr in Bulgaria, references). Distribution. Hubenov, 2025a: suppl: 3 (annotated list Diptera Stara Planina Mts, local distr, habitats and zoogeography). Distribution. Hubenov, 2025b: suppl: 3 (annotated list Diptera high Bulgarian mts, local distr, habitats and zoogeography). | | Channel Is Distribution. Stubbs, 2022b: 5 (on a collection of craneflies from Jersey, collected between 1985 and 2002 by Tony Warne). | | Czech Rep. Distribution. Stary et al., 2005a: 29 (loc(s)). Distribution. Stary, 2009i: webpage (checklist). Distribution. Stary and Vonicka, 2018: 52 (on Limoniidae and Pediciidae of two localities in northern Bohemia, with list of localities, altitudes and habitats). | | Denmark Distribution. Petersen and Jong, 2001b: 151 (checklist). | | Estonia Distribution. Stary, 2004f: (Fauna Europaea: added Estonia). | | Finland Distribution. Salmela, 2012b: 227, 229 (removed from list of Finnish crane flies; Finnish record(s) refer to nigropunctata). | | France Distribution. Quindroit, 2020a: 33, 45-48 (annotated list with information on abundancy and habitat of the Tipuloidea of the Pays de la Loire regoin, distr) (in French). Distribution. Herbrecht et al., 2021: 57 (loc(s) Loire-Atlantique (44) based on malaise-trapping with notes on habitats) (in French). Distribution. Quindroit and Lemoine, 2021a: 25 (loc(s), resultats de deux campagnes de piegeage dans le Pas-de-Calais (62); see paper for details on habitat). Distribution. Quindroit and Lemoine, 2022a: 38 (table specifying the presence in the five departments of the Hauts-de-France region). Distribution. Quindroit, 2022a: 25-28 (additions to the annotated list of the Tipuloidea of Pays de la Loire as given in Quindroit, 2020a). Distribution. Quindroit, 2022d: 417 (loc(s) Ariege (09); zone de suintement boueux en bordure de sous-bois). Distribution. Quindroit and Racine, 2025: 359 (table specifying the presence in the five departments of the Pays de la Loire region). | | Germany Distribution. Schacht, 1999: 134 (checklist Bayern). Distribution. Reusch and Oosterbroek, 2000: 156 (checklist German Bundeslander). Distribution. Reusch and Bellstedt, 2001b: 72 (checklist Thuringen). Distribution. Ruckert, 2005: 47, 53 (found in a study on the association with the type of source: limnokrene, rheokrene, or helokrene in Schlesig-Holstein; see paper for details, as well as Anhang A2-A10). Distribution. Schacht, 2005: 6 (checklist Bayern). Distribution. Wittrock, 2005: 85 (loc(s) Schleswig-Holstein). Distribution. Ruckert et al., 2007: 56 (loc(s) Schleswig-Holstein). Distribution. Hable et al., 2010b: 92 (loc(s) Bayern). Distribution. Schacht, 2010: 18 (checklist Diptera Bayern). Distribution. Stuke, 2019: 146 (checklist Niedersachsen and Bremen with reference to original sources). Distribution. Stuke et al., 2024: 161 (loc(s) Niedersachsen). | | Great Britain Distribution. Godfrey, 2000a: 13, 22 (see paper for details of sites surveyed in Shropshire and into Wales). Distribution. Howe et al., 2001: 139, 143-146 (loc(s) Dorset). Distribution. Gibbs, 2002a: 1 (loc(s) Bristol Region). Distribution. Crossley, 2003b: 62 (loc(s) Sheffield). Distribution. Boardman, 2007: 72-73 (loc(s), map). Distribution. Skidmore, 2009: 128 (review of occurrence on the Western Isles of Scotland, distr). Distribution. Drake, 2011a: 25 (loc(s) Devon). Distribution. Kramer, 2011j: 20 (loc(s) Leicestershire and Rutland (VC55)). Distribution. Hewitt, 2014: (provisional checklist Cumbrian Diptera). Distribution. Showers, 2015: 2 (on craneflies in moth traps at Pitsford Water Nature Reserve in 2013 and 2014, including information for the two traps and photos of their habitat). Distribution. Brighton, 2017c: 28, 37 (overall regional checklist Lancashire and Cheshire (VC58, 59 and 60) with notes on habitats and/or regional occurrence and/or status; see paper for details). Distribution. Showers, 2018c: 4 (loc(s) Northampshire). Distribution. Cunningham, 2019: 16-17 (kocs Devon). Distribution. Kramer and Morris, 2022b: 13 (info on the species, on the distr and map of loc(s) Leicestershire and Rutland (VC55)). | | Greece First record. Kolcsar et al., 2021b: 215 (loc(s) Peloponnisos). | | Ireland Distribution. Ashe et al., 2005a: 317, 334-336 (review single Irish record). Distribution. Chandler et al., 2008: 12 (checklist). | | Italy Distribution. Podenas and Podeniene, 2008: 352 (annotated checklist South Tyrol [Prov. Bolzano]). | | Latvia Distribution. Stary, 2004f: (Fauna Europaea: added Latvia). | | Lithuania Distribution. Podeniene, 2001a: 84 (loc(s)). Distribution. Pakalniskis et al., 2006: 24 (checklist). | | Netherlands Distribution. Jong and Oosterbroek, 2002b: 32 (checklist). Distribution. Stary, 2020a: 45 (Limoniidae Kaaistoep, Noord-Brabant). | | North Macedonia Distribution. Oosterbroek and Simova-Tosic, 2004: 446 (review literature). | | Norway First record. Olsen et al., 2018: 144, 160-161 (loc(s) O, Moss; distr; annotated checklist Nordic countries and for Norway according to the Strand regions). | | Poland Distribution. Wiedenska, 2007a: 76 (checklist). Distribution. Wiedenska, 2010: 135 (loc(s) Parku Krajobrazowego Wzniesien Lodzkich, Lodz). Distribution. Wiedenska, 2014: 12 (on the occurence of Limoniidae in six selected plant communities in the Kamienica Valley, Gorce National Park, Western Carpathians) (in Polish, habitat tables also in English). Distribution. Wiedenska, 2017a: 14 (faunistic and habitat data of species from the Gorce NP) (in Polish). Distribution. Wiedenska and Syratt, 2023: 133 (loc(s) Bieszczady Mountains, distr and remarks). | | Romania Distribution. Ujvarosi, 2007: 228-231 (checklist Limoniidae). Distribution. Kolcsar et al., 2013: 73 (loc(s) Cluj-Napoca and surroundings; list of habitats with altitudes). | | Russia Distribution. Pilipenko, 2017a: 389 (investigation of Tipuloidea of the Altay, first record for Altay and first confirmation for Eastpalaearctic) (in Russian). Distribution. Pilipenko et al., 2020: 359 (loc(s) RUC: Mordoviya Rep., distr). | | Slovakia Distribution. Stary, 2009d: 26 (loc(s) Polana area). Distribution. Stary, 2009i: webpage (checklist). | | Sweden Distribution. Lindstrom and Fritz, 2015: 44, 48 (distributin and ecology in five fens in Hallands County) (in Schwedish). | | Switzerland Distribution. Podenas et al., 2006: 149, 327 (map). | | Turkey First record. Koc, 2008: 4 (loc(s) Afyonkarahisar, Isparta). Distribution. Ozgul et al., 2009: 64 (survey provinces SW Turkey). Distribution. Bilgin et al., 2015: 6 (loc(s) Eskisehir, distr) (in Turkish). Distribution. Koc et al., 2016: 11 (provinces Marmara region). Distribution. Provinces Asiatic part, 2025: (known from Afyonkarahisar, Bursa, Eskisehir, Isparta). | | | | Belarus Kolcsar et al., 2021b: month(s): 7. | | Belgium Dek and Peeters, 2023a: month(s): 9. | | Bulgaria Hubenov, 2015: month(s): 8. | | Channel Is Stubbs, 2022b: month(s): 6-7. | | Czech Rep. Stary and Vonicka, 2018: month(s): 6-7. | | France Quindroit, 2020a: month(s): 5-7. Quindroit, 2022d: month(s): 7. | | Germany Stuke et al., 2024: month(s): 6. | | Great Britain Godfrey, 2000a: month(s): 6. Gibbs, 2002a: month(s): 7. Crossley, 2003b: month(s): 6, 8. Kramer, 2011j: month(s): 6, 8. Cunningham, 2019: month(s): 7, 9. Kramer and Morris, 2022b: month(s): 6-7. | | Greece Kolcsar et al., 2021b: month(s): 5. | | Norway Olsen et al., 2018: month(s): 6. | | Poland Wiedenska, 2014: month(s): 7. Wiedenska, 2015a: month(s): 7-8. Wiedenska and Syratt, 2023: month(s): 7. | | Romania Kolcsar et al., 2013: month(s): 6. | | Russia Pilipenko et al., 2020: month(s): 8. | | Slovakia Stary, 2009d: month(s): 6-7. | | Switzerland Podenas et al., 2006: month(s): 6-7. | | Turkey Koc, 2008: month(s): 6. | | | | Belarus Kolcsar et al., 2021b: altitude: 156-161 m. | | Bulgaria Hubenov, 2015: altitude: 1960-2914 m. Hubenov, 2017: altitude: 1960-2914 m. Hubenov, 2018b: altitude: 380 m. Hubenov, 2021a: altitude: 0-1914 m. Hubenov, 2025a: altitude: 0-2914 m. Hubenov, 2025b: altitude: 0-2914 m. | | Czech Rep. Stary and Vonicka, 2018: altitude: 450-630 m. | | France Quindroit, 2022d: altitude: 800 m. | | Germany Stuke et al., 2024: altitude: 150-170 m. | | Greece Kolcsar et al., 2021b: altitude: 200-250 m. | | Poland Wiedenska and Syratt, 2023: altitude: 500 m. | | Slovakia Stary, 2009d: altitude: 560-780 m. | | Turkey Koc, 2008: altitude: 834-1350 m. |
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