Details for Phylidorea (Phylidorea) ferruginea

Name: Phylidorea (Phylidorea) ferruginea (Meigen, 1818) Publication: Syst. Beschr. 1: 128 (as Limnobia) Status: Recognized taxon, Synonym(s): fulvescens (Meigen, 1818); praeusta (Schummel, 1829); ?bifurcata (Zetterstedt, 1837); unicolor (Walker, 1856). Classification: Family Limoniidae Subfamily Limnophilinae Keys: Stary and Freidberg, 2007 (Limoniidae Israel); Krivosheina, M.G., 2010 (larvae of 6 Phylidorea species); Peeters en Oosterbroek, 2014c, 2016c (craneflies Netherlands, Belgium and Luxembourg) (in Dutch); Boardman, 2016 (craneflies Shropshire); Stubbs and Kramer, 2016g; Stubbs, 2021 (British craneflies) Variant spelling(s): ferrugina habitus male habitus male habitus female habitus female habitus copula plus 16 more images of body part(s), body part(s), hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, ovipositor, wing, wing, wing, habitat, habitat, habitat and habitat. Distribution: Austria, Belarus, Belgium, Bosnia-Herzegovina, Bulgaria, Croatia, Czech Rep., Denmark (incl. Faroe Is), Estonia, Finland, France (incl. Corsica), Germany, Great Britain, Greece, Hungary, Ireland, Italy (incl. Sardinia, Sicily), Latvia, Lithuania, Montenegro, Netherlands, North Macedonia, Norway, Poland, Portugal, Romania, Serbia, Slovakia, Slovenia, Spain (incl. Balearic Is [Mallorca]), Sweden, Switzerland, Turkey (European part: Istanbul, Kirklareli), Ukraine; Russia: RUN, RUW, RUC, RUE, RUS, North Caucasus; Georgia, Armenia, Azerbaijan, Turkey (Asiatic part: see citations for provinces), Israel;; Russia: WS (Tuva); Kazakhstan, Uzbekistan, Turkmenistan, Tajikistan, Kyrgyzstan; Mongolia. Note: see the manual for abbreviations: present-day Russia. Region(s): Westpalaearctic;; Eastpalaearctic General note: Added: Spain (Geiger et al., 1994). For Italy check Stary and Oosterbroek, 1996, and citations below. For Finland check Salmela, 2012b, 2012c, and citations below. For Norway check Olsen et al., 2018, and citations below. For Portugal check Oosterbroek et al., 2020, and citations below. For Great Britain and Ireland check Stubbs, 2021, and citations below. Map: Ashe et al., 2005a (Ireland); Koc et al., 2006 (SW Turkey); Boardman, 2007 (Shropshire) Figure references: Habitus: Quindroit, 2020a (male, female); Ferreira et al., 2021 (female); Kramer and Morris, 2022b (female) Wing: Alexander, 1972b; Podenas et al., 2006 Hypopygium: Alexander, 1972b; Podenas et al., 2006; Kramer and Morris, 2022b Immature stages Larva: Podeniene, 2003a Citations on biology (mainly from 2000 onward): Azerbaijan Habitat. Stary and Obona, 2020: 131 (loc(s); with list of localities, habitats and altitudes). Belarus Habitat. Paramonov and Sushko, 2010: 45 (loc(s), distr, about fauna of raised bogs, for details see paper) (in Russian). Bulgaria Habitat. Hubenov, 2025b: suppl: 3 (annotated list Diptera high Bulgarian mts, local distr, habitats and zoogeography). Czech Rep. Habitat. Stary et al., 2013b: 378 (on Limonidae from Vraz near Pisek, with general habitat description). Habitat. Stary and Vonicka, 2018: 52 (on Limoniidae and Pediciidae of two localities in northern Bohemia, with list of localities, altitudes and habitats). Denmark Habitat. Nielsen and Nielsen, 2009: 122 (emerging from a forest floor in a Beech [Fagus] stand) (in Danish). Habitat. Byriel and Rojas, 2017: 118 (among the 101 species in a study on cranefly hotspots in unmanaged and managed forests) (in Danish). Finland Habitat. Autio and Salmela, 2010: 48 (habitats Aland Is, a swamp-dwelling species). France Habitat. Kramer and Withers, 2007: 159 (marshland species with larvae in the more or less saturated organic mud of lake margins and marshes). Habitat. Kramer, 2007a: 33 (adults were collected along lake margins). Biology. Quindroit, 2020a: 32-33, 45-48 (annotated list with information on abundancy and habitat of the Tipuloidea of the Pays de la Loire regoin, distr) (in French). Habitat. Quindroit, 2020b: 141 (loc(s) Nord (59) and Pas-de-Calais (62) with notes on habitats and distr) (in French). Habitat. Herbrecht et al., 2021: 57 (loc(s) Loire-Atlantique (44) based on malaise-trapping with notes on habitats) (in French). Habitat. Quindroit and Lemoine, 2021a: 24, 25 (loc(s), resultats de deux campagnes de piegeage dans le Pas-de-Calais (62); see paper for details on habitat). Habitat. Quindroit and Lemoine, 2021b: 8 (loc(s) Nord (59); see paper for habitat description) (in French). Habitat. Quindroit et al., 2021: 256-258 (collected during an entomological inventory in the canopy of Oak [Quercus] stands; marais). Habitat. Quindroit, 2021a: 36 (loc(s), resultats d une campagne de piegeage dans le Pas-de-Calais (62); see paper for details on habitat). Habitat. Quindroit and Lemoine, 2022a: 30 (loc(s) Aisne (02), Oise (60) and Somme (80) with notes on habitats and distr) (in French). Georgia Habitat. Stary and Obona, 2020: 131 (loc(s); with list of localities, habitats and altitudes). Germany Habitat. Reusch, 2006: 134 (Kalkquellmoor [carbonate fen complex]). Habitat. Lehmann and Reusch, 2009: 122 (from around lowland springs). Habitat. Reiff et al., 2015: 234 (loc(s) Baden-Wurttemberg with list of habitats). Great Britain Habitat. Godfrey, 2003a: 44 (associated with woody debris). Habitat. Boardman, 2007: 71 (lush vegetation on wet ground such as lake and pond edges, fen, mire, upland moorland and wet Juncus grassland). Habitat. Phillips, 2007: 150 (flushes running to beach) (as Limnophila). Habitat. Cranston and Drake, 2010: 175 (Phylidorea larvae can be found in marginal situations along standing waters [no species mentioned]). Habitat. Kramer, 2011j: 20 (a common marshland species). Habitat. Chandler, 2015: 85 (collected at a wooded area surrounded by ditches). Habitat. Showers, 2015: 2 (on craneflies in moth traps at Pitsford Water Nature Reserve in 2013 and 2014, including information for the two traps and photos of their habitat). Greece Habitat. Kramer, 2004: 20-22 (found at a lake margin). Ireland Habitat. Ashe et al., 2005a: 317, 332-333, 335 (bred from mossy woodland litter, citing Blackith et al., 1991). Italy Habitat. Podenas, 2011: 863 (larvae develop in damp soil near water reservoirs). Latvia Habitat. Spungis, 2008: 36 (raised bogs). Lithuania Habitat. Podeniene, 2001a: 83, 85 (larvae were found in the intertidal zone of small river in silt and leaf fallings, and in leaf fallings in low-laying marshland in Alnus forest (Carici elongatae-Alnetum)). Habitat. Podeniene, 2002: 297, 299 (larvae develop in mud and leaf litter in low-laying marsh (Carici elongatae-Alnetum) in forest and in riparian zones of river). Habitat. Podeniene, 2003a: 1-295 (larva, descr, figs, habitat) (in Lithuanian). Habitat. Podeniene, 2003b: 10, 21-23, 29 (larval habitat: sand and silt in riparian zone of small rivulets, swamps at the bottom of karst pits in old deciduous forest, mud (in bogs of low lying woody swamps, on swampy riversides), fallings and uppermost clay soil layer in Salix groves, sandy loam under grassy vegetation or under fallings not completely covered by forest). Netherlands Habitat. Dek and Oosterbroek, 2013: 87 (collected in a damp deciduous forest with undergrowth largely of Aegopodium podagraria and forest floor with much dead wood). Norway Habitat. Olsen and Andersen, 2021: 211 (loc(s) Innlandet, in former Hedmark, with habitats specified separately, for habitats see also Jonassen and Andersen, 2020). Romania Habitat. Parvu, 2003: 233. Habitat. Ujvarosi, 2005a: 247 (standing water, peat bog, swamps, citing various sources). Habitat. Ujvarosi and Poti, 2006: 256 (habitat in Dupa Lunca marshland: damp meadows). Habitat. Ujvarosi et al., 2011b: 108 (loc(s) Dupa Lanca marshy area near Voslobeni) (in Hungarian, species list with habitat information in English). Habitat. Kolcsar et al., 2013: 73 (loc(s) Cluj-Napoca and surroundings; list of habitats with altitudes). Russia Habitat. Przhiboro, 2003: 364 (larvae common in the water margin zone of an oligotrophic lake; rearing). Habitat. Przhiboro, 2004: 103, 111 (in Russian). Biology. Przhiboro, 2012: 260 (on the abundance and biomass values of Tipuloidea immatures at five sites of the lake water margin zone, Rakovye lakes, Leningradskaya oblast) (in Russian). Habitat. Lantsov, 2014b: 153 (loc(s) North Caucasus: Karachay-Cherkess Rep.; found in same habitats as Prionocera turcica). Spain Habitat. Stary and Oosterbroek, 2008: 10 (marshland; wet Quercus forest). Habitat. Stary, 2014b: 88 (collected at saltmarsh and canal). Sweden Habitat. Lindstrom and Fritz, 2015: 44, 48 (distributin and ecology in five fens in Hallands County) (in Schwedish). on characters and taxonomy (mainly from 2000 onward): Country not relevant Characters. Frantsevich, 2004: 133 (structure of coxa). Phylogeny. Boldgiv, 2006: 96-154 (one of the 84 morphological taxa from northern Mongolia in a consistency test of morphology based species identification of crane flies against DNA-barcoding identification using 536-base-pair sequence profiles of the COI gene, also known as the DNA barcode). Immatures. Krivosheina, M.G., 2010a: 19 (key to larvae of 6 Phylidorea species) (for key in English see Krivosheina, M.G., 2010b). Immatures. Podeniene and Gelhaus, 2010: 200-202 (larvae of the genus Phylidorea can be divided into the abdominalis, ferruginea and squalens groups based on the sclerotization of the spiracular field, the ferruginea group consists of 5 species, ferruginea, nervosa, fulvonervosa, nigronotata and temelskin). Great Britain Characters. Kramer and Morris, 2022b: 12 (diagnosis, figs). Israel Key. Stary and Freidberg, 2007: 305, 307, 313 (key). Lithuania Immatures. Podeniene, 2003a: 1-295 (larva, descr, figs, habitat) (in Lithuanian). Portugal Characters. Ferreira et al., 2021: 7 (barcoding based on material mentioned in Oosterbroek et al. 2020). on distribution (mainly from 2000 onward): Armenia First record. Obona et al., 2016: 128 (loc(s)). Distribution. Hakobyan and Jenderedjian, 2023: 1512 (annotated checklist of the craneflies of Armenia with refs, distr and flight period in Caucasus Ecoregion with refs). Azerbaijan Distribution. Stary and Obona, 2020: 131 (loc(s); with list of localities, habitats and altitudes). Distribution. Snegovaya, 2021: 1148 (checklist with details on distr in Azerbaijan). Belarus First record. Paramonov and Sushko, 2010: 45 (loc(s), distr, about fauna of raised bogs, for details see paper) (in Russian). Distribution. Gavryushin in litt., 2013d: (loc(s) Minsk region). Belgium Distribution. Martens et al., 2013: 125 (loc(s) Oost-Vlaanderen). Distribution. Martens et al., 2014: 114, 124 (loc(s) Oost-Vlaanderen). Distribution. Dek and Peeters, 2023a: 139 (loc(s) Botanic Garden Jean Massart, Brussels-Capital Region, common in Belgium). Bosnia-Herzegowina Distribution. Oosterbroek and Simova-Tosic, 2004: 446 (review literature). Bulgaria Distribution. Hubenov, 2016: 47 (annotated checklist Diptera of the Rila Mountains based on data from the literature). Distribution. Hubenov, 2017: 68 (on vertical distribution in the Pirin and Rila Mts; see paper for details on forests and subalpine-alpine zones, distr). Distribution. Hubenov, 2018a: 16 (on the Diptera of the Vitosha Mt; see paper for details on vegetation belts and local distr). Distribution. Hubenov, 2021a: 30 (checklist with summary of distr in Bulgaria, references). Distribution. Hubenov, 2025b: suppl: 3 (annotated list Diptera high Bulgarian mts, local distr, habitats and zoogeography). Channel Is Distribution. Stubbs, 2022b: 5 (on a collection of craneflies from Jersey, collected between 1985 and 2002 by Tony Warne). Croatia Distribution. Oosterbroek and Simova-Tosic, 2004: 446 (review literature). Czech Rep. Distribution. Stary et al., 2005a: 29 (loc(s)). Distribution. Stary, 2009i: webpage (checklist). Distribution. Stary et al., 2013b: 378 (on Limonidae from Vraz near Pisek, with general habitat description). Distribution. Stary and Vonicka, 2018: 52 (on Limoniidae and Pediciidae of two localities in northern Bohemia, with list of localities, altitudes and habitats). Denmark Distribution. Petersen and Jong, 2001b: 151 (checklist). Distribution. Nielsen and Nielsen, 2009: 122 (loc(s)). Estonia Distribution. Stary, 2004f: (Fauna Europaea: added Estonia). Finland Distribution. Salmela, 2002: 30-32 (loc(s) lake Iidesjarvi) (in Finnish). Distribution. Salmela et al., 2007b: 45 (loc(s)). Distribution. Autio and Salmela, 2010: 48 (loc(s) Aland Is). Distribution. Salmela, 2010a: 12 (survey fauna Tahmela spring in city of Tampere) (in Finnish). Distribution. Salmela, 2012b: 239 (annotated list of Finnish crane flies). Distribution. Salmela and Petrasiunas, 2014: 28 (checklist Finnish Tipulomorpha). Distribution. Salmela et al., 2015: 106 (list of localities in extensive report on malaise-trapping of insects in conservation areas in Lapland in 2012–2014) (in Finnish). France Distribution. Kramer and Withers, 2007: 159, 163 (loc(s) Ain (01)). Distribution. Kramer, 2007a: 33 (loc(s) Ain (01)). Distribution. Withers, 2008: 137 (loc(s) Ain (01)). Distribution. Garrin and Herbrecht, 2016: Annexe 11 (loc(s) Notre-Dame-des-Landes, Loire-Atlantique (44)). Distribution. Quindroit, 2020a: 32-33, 45-48 (annotated list with information on abundancy and habitat of the Tipuloidea of the Pays de la Loire regoin, distr) (in French). Distribution. Quindroit, 2020b: 141 (loc(s) Nord (59) and Pas-de-Calais (62) with notes on habitats and distr) (in French). Distribution. Quindroit, 2020d: 328 (collected at the Golf de Vidauban, Vidauban, Var (83)). Distribution. Herbrecht et al., 2021: 57 (loc(s) Loire-Atlantique (44) based on malaise-trapping with notes on habitats) (in French). Distribution. Quindroit and Lemoine, 2021a: 24, 25 (loc(s), resultats de deux campagnes de piegeage dans le Pas-de-Calais (62); see paper for details on habitat). Distribution. Quindroit and Lemoine, 2021b: 8 (loc(s) Nord (59); see paper for habitat description) (in French). Distribution. Quindroit et al., 2021: 257 (recorded for Loiret (45)). Distribution. Quindroit, 2021a: 36 (loc(s), resultats d une campagne de piegeage dans le Pas-de-Calais (62); see paper for details on habitat). Distribution. Tissot et al., 2021: 209 (the Diptera of the Reserves Naturelles Nationales du Doubs (25), results of Malaise trapping since 2009) (in French). Distribution. Quindroit and Lemoine, 2022a: 30 (loc(s) Aisne (02), Oise (60) and Somme (80) with notes on habitats and distr) (in French). Distribution. Quindroit and Lemoine, 2022a: 38 (table specifying the presence in the five departments of the Hauts-de-France region). Distribution. Lemoine and Quindroit, 2023: 9 (loc(s) Cote-d'Or (21), Moselle (57), Nievre (58)). Distribution. Quindroit and Racine, 2025: 359 (table specifying the presence in the five departments of the Pays de la Loire region). Georgia First record. Stary and Obona, 2020: 131 (loc(s); with list of localities, habitats and altitudes). Germany Distribution. Schacht, 1999: 134 (checklist Bayern). Distribution. Reusch and Oosterbroek, 2000: 156 (checklist German Bundeslander). Distribution. Reusch and Bellstedt, 2001b: 72 (checklist Thuringen). Distribution. Schacht, 2005: 6 (checklist Bayern). Distribution. Reusch, 2006: 134 (loc(s) Bayern). Distribution. Lehmann and Reusch, 2009: 122 (loc(s) Niedersachsen). Distribution. Hable et al., 2010b: 92 (loc(s) Bayern). Distribution. Schacht, 2010: 18 (checklist Diptera Bayern). Distribution. Reiff et al., 2015: 234 (loc(s) Baden-Wurttemberg with list of habitats). Distribution. Stuke, 2019: 145 (checklist Niedersachsen and Bremen with reference to original sources). Great Britain Distribution. Godfrey, 2003a: 44 (distr). Distribution. Hewitt et al., 2005: 38 (loc(s) Cumbria) (as Limnophila). Distribution. Boardman, 2007: 71 (distr, map). Distribution. Phillips, 2007: 150 (loc(s) Isle of Rum) (as Limnophila). Distribution. Drake, 2008c: 223 (loc(s) Wiltshire, in study on invertebrates of two water meadows). Distribution. Skidmore, 2008a: 59 (loc(s) Tiree, Western Isles, Scotland). Distribution. Skidmore, 2008b: 68 (loc(s) Kerrera, Western Isles, Scotland). Distribution. Skidmore, 2009: 128 (review of occurrence on the Western Isles of Scotland, distr). Distribution. Hancock, 2011a: 2 (loc(s) Tiree and Coll, Southern Inner Hebrides). Distribution. Kramer, 2011j: 20 (loc(s) Leicestershire and Rutland (VC55)). Distribution. Hewitt, 2014: (provisional checklist Cumbrian Diptera). Distribution. Kramer, 2014c: 4 (in table of cranefly species recorded at Shapwick Heath NNR, as given in H. Audcent, 1949 and/or collected by J. Kramer in 2013). Distribution. Wolton et al., 2014: 27 (among Diptera associated with a hedge in Devon). Distribution. Chandler, 2015: 90 (loc(s) Bushy Park, Middlesex). Distribution. Showers, 2015: 2 (on craneflies in moth traps at Pitsford Water Nature Reserve in 2013 and 2014, including information for the two traps and photos of their habitat). Distribution. Robinson et al., 2016: 79 (list of terrestrial invertebrates of Mingulay Is, Outer Hebrides). Distribution. Showers, 2016: 2 (loc(s) Northamptonshire). Distribution. Brighton, 2017c: 28 (overall regional checklist Lancashire and Cheshire (VC58, 59 and 60) with notes on habitats and/or regional occurrence and/or status; see paper for details). Distribution. Showers, 2017a: 4 (loc(s) Northamptonshire). Distribution. Lovegrove et al., 2018: 6-7 (among the cranefly species found in a survey of the historical site for Idiocera sexguttat at Stony Moors in the New Forest). Distribution. MacDonald, 2018: 216 (loc(s) Scottish Highland). Distribution. Kramer and Morris, 2022b: 12 (info on the species, on the distr and map of loc(s) Leicestershire and Rutland (VC55)). Greece First record. Kramer, 2004: 20-22 (loc(s) Peloponnisos). Distribution. Stary and Oosterbroek, 2008: 10 (loc(s), distr). Hungary Distribution. Toth, 2005: 169 (loc(s)). Ireland Distribution. Ashe et al., 2005a: 317, 332-333, 335 (review Irish records). Distribution. Ashe et al., 2008: 14 (additional Irish record(s), distr). Distribution. Chandler et al., 2008: 12 (checklist). Israel First record. Stary and Freidberg, 2007: 305, 307, 313 (loc(s), distr). Italy Distribution. Podenas, 2011: 863 (loc(s) Sardinia, distr). Kazakhstan Distribution. Devyatkov, 2019b: 246 (loc(s) Pavlodarskaya oblast, distr) (in Russian). Distribution. Devyatkov, 2020d: 271 (loc(s) East Kazakhstan, distr) (in Russian). Latvia First record. Spungis, 2008: 36 (loc(s)). Lithuania Distribution. Podeniene, 2001a: 83, 85 (loc(s)). Distribution. Podeniene, 2002: 297, 299 (loc(s)). Distribution. Pakalniskis et al., 2006: 23 (checklist). Mongolia Distribution. Boldgiv, 2006: 154 (loc(s)). Montenegro Distribution. Oosterbroek and Simova-Tosic, 2004: 446 (review literature). Netherlands Distribution. Jong and Oosterbroek, 2002b: 32 (checklist). Distribution. Dek and Oosterbroek, 2013: 87 (loc(s) Zeeland). Distribution. Hoven, 2019: 20 (loc(s) Gaasterland). Distribution. Steenis, 2019: 15 (loc(s) Noord-Brabant, Biesbosch). Distribution. Oosterbroek and Dek, 2020: 17 (Limoniidae and Tipulidae Nationaal Park Hollandse Duinen in 2018, Zuid-Holland). Distribution. Stary, 2020a: 45 (Limoniidae Kaaistoep, Noord-Brabant). North Macedonia Distribution. Oosterbroek and Simova-Tosic, 2004: 446 (review literature). Norway Distribution. Olsen et al., 2018: 160-161 (annotated checklist Nordic countries and for Norway according to the Strand regions). Distribution. Olsen and Andersen, 2021: 211 (loc(s) Innlandet, in former Hedmark, with habitats specified separately, for habitats see also Jonassen and Andersen, 2020). Poland Distribution. Palaczyk, 2001: 269 (loc(s)). Distribution. Wiedenska, 2007a: 76 (checklist). Distribution. Wiedenska, 2010: 135 (loc(s) Parku Krajobrazowego Wzniesien Lodzkich, Lodz). Portugal First record. Stary, 2014b: 88 (loc(s) Faro). Distribution. Oosterbroek et al., 2020: 328 (recorded for six provinces, distr). Romania Distribution. Parvu, 2003: 233 (loc(s)). Distribution. Ujvarosi and Poti, 2006: 256 (loc(s) eastern Carpathians). Distribution. Ujvarosi, 2007: 228-231 (checklist Limoniidae). Distribution. Ujvarosi et al., 2011b: 108 (loc(s) Dupa Lanca marshy area near Voslobeni) (in Hungarian, species list with habitat information in English). Distribution. Kolcsar et al., 2013: 73 (loc(s) Cluj-Napoca and surroundings; list of habitats with altitudes). Russia Distribution. Przhiboro, 2003: 364 (loc(s) NW Russia). Distribution. Przhiboro, 2004: 103, 111 (loc(s) RUW: Leningradskaya oblast) (in Russian). Distribution. Polevoi, 2006: 96 (loc(s) RUN: Kareliya) (in Russian). Distribution. Gavryushin in litt., 2011f: (loc(s) RUN: Arkhangelskaya oblast). Distribution. Przhiboro, 2012: 260 (on the abundance and biomass values of Tipuloidea immatures at five sites of the lake water margin zone, Rakovye lakes, Leningradskaya oblast) (in Russian). Distribution. Jakovlev et al., 2014: 309 (loc(s) RUN: Kareliya). Distribution. Lantsov, 2014b: 153 (loc(s) North Caucasus: Karachay-Cherkess Rep.; found in same habitats as Prionocera turcica). Distribution. Paramonov and Klepikov, 2014: 36 (loc(s) Yaroslavskaya oblast). Distribution. Lantsov, 2016: 165-170 (on craneflies from Arkhyz, Karachay-Cherkessia Rep., North Caucasus) (in Russian). Distribution. Paramonov and Pilipenko, 2016: 98 (loc(s) RUC: Tverskaya oblast) (in Russian). Distribution. Paramonov, 2018a: 183 (loc(s) RUC: Chuvash Rep., distr) (in Russian). Distribution. Paramonov and Korobkov, 2019: 98 (loc(s) RUC: Tverskaya oblast, distr, citing various authors) (in Russian). Distribution. Pilipenko et al., 2020: 358 (loc(s) RUC: Mordoviya Rep., distr). Distribution. Kolcsar et al., 2021b: 203-204 (loc(s) RUE: Bashkortostan Rep.). Serbia First record. Kolcsar et al., 2021b: 204 (loc(s)). Slovakia Distribution. Stary, 2009d: 26 (loc(s) Polana area). Distribution. Stary, 2009i: webpage (checklist). Slovenia Distribution. Oosterbroek and Simova-Tosic, 2004: 446 (review literature). Spain Distribution. Eiroa and Baez, 2002a: 55 (checklist). Distribution. Stary and Oosterbroek, 2008: 10 (loc(s), Spanish provinces, distr). Distribution. Stary, 2014b: 88 (loc(s) Mallorca, first record for Balearic Is). Distribution. Keresztes et al., 2022: 194 (loc(s) Asturias and review distribution in Spain) (in Spanish). Sweden Distribution. Salmela, 2010b: 133 (loc(s) Malmo). Distribution. Lindstrom and Fritz, 2015: 44, 48 (distributin and ecology in five fens in Hallands County) (in Schwedish). Switzerland Distribution. Podenas et al., 2006: 147, 321 (map, vertical distr per thermic level). Distribution. Dufour and Merz, 2012: 354 (annotated checklist canton Geneva). Tajikistan Distribution. Devyatkov, 2020c: 78 (loc(s), distr) (in Russian). Turkey First record. Koc et al., 2006: 19-21 (loc(s), distr, map). Distribution. Stary and Oosterbroek, 2008: 10 (loc(s), Turkish provinces, distr). Distribution. Ozgul et al., 2009: 64 (survey provinces SW Turkey). Distribution. Bilgin et al., 2015: 6 (loc(s) Eskisehir, distr) (in Turkish). Distribution. Koc et al., 2016: 11 (provinces Marmara region). Distribution. Akpinar and Ozgul, 2023: 55 (loc(s) Aksaray). Distribution. Provinces Asiatic part, 2025: (known from Aksaray, Balikesir, Burdur, Bursa, Canakkale, Erzurum, Eskisehir). on flight period (mainly from 2000 onward): Armenia Obona et al., 2016: month(s): 9. Azerbaijan Stary and Obona, 2020: month(s): 5. Belarus Paramonov and Sushko, 2010: month(s): 8. Gavryushin in litt., 2013d: month(s): 7. Belgium Martens et al., 2013: month(s): 8. Martens et al., 2014: month(s): 4-9. Dek and Peeters, 2023a: month(s): 5-10. Bulgaria Hubenov, 2016: month(s): 5-7. Channel Is Stubbs, 2022b: month(s): 6, 10. Czech Rep. Stary et al., 2005a: month(s): 5-8. Stary et al., 2013b: month(s): 9-10. Stary and Vonicka, 2018: month(s): 5-6. France Kramer and Withers, 2007: month(s): 4-6. Kramer, 2007a: month(s): 6. Quindroit, 2020a: month(s): 3-10. Quindroit, 2020b: month(s): 4-10. Quindroit and Lemoine, 2021b: month(s): 5. Quindroit and Lemoine, 2022a: month(s): 4-9, 11. Lemoine and Quindroit, 2023: month(s): 4, 7. Georgia Stary and Obona, 2020: month(s): 7, 9. Germany Reiff et al., 2015: month(s): 6. Great Britain Phillips, 2007: month(s): 8. Skidmore, 2008a: month(s): 6. Skidmore, 2008b: month(s): 6. Hancock, 2011a: month(s): 6. Kramer, 2011j: month(s): 7. Kramer, 2014c: month(s): 6. Chandler, 2015: month(s): 7-8. Kramer and Morris, 2022b: month(s): 4-10. Greece Kramer, 2004: month(s): 4. Stary and Oosterbroek, 2008: month(s): 7, 10. Ireland Ashe et al., 2005a: month(s): 5-10. Ashe et al., 2008: month(s): 6, 8. Israel Stary and Freidberg, 2007: month(s): 12. Italy Podenas, 2011: month(s): 11. Kazakhstan Devyatkov, 2019b: month(s): 6. Devyatkov, 2020d: month(s): 5-8. Netherlands Dek and Oosterbroek, 2013: month(s): 10. Hoven, 2019: month(s): 5. Oosterbroek and Dek, 2020: month(s): 4-7, 9. Norway Olsen and Andersen, 2021: month(s): 7. Portugal Stary, 2014b: month(s): 4-5. Oosterbroek et al., 2020: month(s): 5-6, 9. Romania Parvu, 2003: month(s): 7. Ujvarosi et al., 2011b: month(s): 5. Kolcsar et al., 2013: month(s): 7. Russia Przhiboro, 2003: month(s): 6-7. Gavryushin in litt., 2011f: month(s): 8. Paramonov and Klepikov, 2014: month(s): 9. Paramonov and Pilipenko, 2016: month(s): 6, 9. Paramonov, 2018a: month(s): 5, 8. Paramonov and Korobkov, 2019: month(s): 5-6, 8. Pilipenko et al., 2020: month(s): 5, 8-9. Kolcsar et al., 2021b: month(s): 7. Serbia Kolcsar et al., 2021b: month(s): 9. Slovakia Stary, 2009d: month(s): 5-6. Spain Stary and Oosterbroek, 2008: month(s): 4-7. Stary, 2014b: month(s): 5. Keresztes et al., 2022: month(s): 6. Switzerland Podenas et al., 2006: month(s): 4-10. Tajikistan Devyatkov, 2020c: month(s): 6-7. Turkey Koc et al., 2006: month(s): 5. Stary and Oosterbroek, 2008: month(s): 5-6. on altitude (mainly from 2000 onward): Armenia Obona et al., 2016: altitude: 1705 m. Azerbaijan Stary and Obona, 2020: altitude: 200 m. Bulgaria Hubenov, 2016: altitude: 1700 m. Hubenov, 2017: altitude: 1700 m. Hubenov, 2021a: altitude: 200-1700 m. Hubenov, 2025b: altitude: 200-1700 m. Czech Rep. Stary and Vonicka, 2018: altitude: 330-350 m. Georgia Stary and Obona, 2020: altitude: 1115-1970 m. Germany Reiff et al., 2015: altitude: 650 m. Greece Kramer, 2004: altitude: 875 m. Stary and Oosterbroek, 2008: altitude: 1600 m. Kazakhstan Devyatkov, 2019b: altitude: 384-456 m. Devyatkov, 2020d: altitude: 280-727 m. Norway Olsen and Andersen, 2021: altitude: 170 m. Portugal Stary, 2014b: altitude: 126 m. Romania Parvu, 2003: altitude: 800 m. Ujvarosi and Poti, 2006: altitude: 670 m. Russia Gavryushin in litt., 2011f: altitude: 19 m. Kolcsar et al., 2021b: altitude: 550 m. Serbia Kolcsar et al., 2021b: altitude: 1500 m. Spain Stary and Oosterbroek, 2008: altitude: 600-1600 m. Stary, 2014b: altitude: 0-1 m. Keresztes et al., 2022: altitude: 1485-1670 m. Tajikistan Devyatkov, 2020c: altitude: 2250-2440 m. Turkey Koc et al., 2006: altitude: 930 m. Stary and Oosterbroek, 2008: altitude: 1000-2300 m.