Details for Phylidorea (Paraphylidorea) fulvonervosa

Name: Phylidorea (Paraphylidorea) fulvonervosa (Schummel, 1829) Publication: Beitr. Ent., Breslau 1: 164 (as Limnobia) Status: Recognized taxon, Synonym(s): lineolella (Verrall, 1887); sapporensis (Alexander, 1921). Classification: Family Limoniidae Subfamily Limnophilinae Keys: Krivosheina, M.G., 2010 (larvae 6 Phylidorea species); Boardman, 2016 (craneflies Shropshire); Peeters en Oosterbroek, 2016c (craneflies Netherlands, Belgium and Luxembourg) (in Dutch); Stubbs and Kramer, 2016g; Stubbs, 2021 (British craneflies) habitus male habitus male habitus male habitus male body part(s) antenna plus 32 more images of body part(s), body part(s), body part(s), hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, ovipositor, wing, wing, wing, habitat, habitat, habitat, habitat, habitat, habitat, habitat, habitat, habitat, habitat, habitat, habitat, habitat, habitat, habitat and habitat. Distribution: Austria, Belarus, Belgium, Bulgaria, Czech Rep., Denmark, Estonia, Finland, France, Germany, Great Britain, Hungary, Ireland, Italy, Lithuania, Montenegro, Netherlands, Norway, Poland, Portugal, Romania, Slovakia, Spain, Sweden, Switzerland, Ukraine; Russia: RUN, RUW, RUC, RUE;; Russia: FE (Khabarovskiy kray, Primorskiy kray, Sakhalin (incl. Kuril Is)); Kazakhstan (east); Japan (Hokkaido). Note: see the manual for abbreviations: present-day Russia. Region(s): Westpalaearctic;; Eastpalaearctic General note: Added: Montenegro (Simova-Tosic and Vukovic, 1996). For Italy check Stary and Oosterbroek, 1996, and citations below. For Finland check Salmela, 2012b, 2012c, and citations below. For Norway check Olsen et al., 2018, and citations below. For Portugal check Oosterbroek et al., 2020, and citations below. For Great Britain and Ireland check Stubbs, 2021, and citations below. Map: Ashe et al., 2005a (Ireland); Podenas et al., 2006 (Switzerland); Boardman, 2007 (Shropshire) Figure references: Wing: Slipka and Stary, 1977b; Podenas et al., 2006 Hypopygium: Kramer, 2006; Podenas et al., 2006; Kramer and Morris, 2022b Immature stages Larva: Podeniene, 2003a Citations on biology (mainly from 2000 onward): Bulgaria Habitat. Hubenov, 2025b: suppl: 3 (annotated list Diptera high Bulgarian mts, local distr, habitats and zoogeography). Czech Rep. Habitat. Stary et al., 2013b: 378 (on Limonidae from Vraz near Pisek, with general habitat description). Habitat. Stary and Vonicka, 2018: 51-52 (on Limoniidae and Pediciidae of two localities in northern Bohemia, with list of localities, altitudes and habitats). Denmark Habitat. Byriel and Rojas, 2017: 118 (among the 101 species in a study on cranefly hotspots in unmanaged and managed forests) (in Danish). Finland Habitat. Salmela, 2001a: 144, 146, 150 (inhabits marshes and swamps, citing Brinkmann, 1991). Habitat. Salmela, 2004: 9 (ubiquitous in moist and aquatic environments). Habitat. Salmela and Ilmonen, 2005: 89 (the occurrence in relation to the trophic status of mires within an open boreal mire system is investigated; found at a mesotrophic site only). Habitat. Salmela and Autio, 2007b: 49 (eurytopic dweller of wetlands and moist biotopes). Biology. Salmela, 2008: 10, 24, 27, 32, 50 (49) (ecology or habitat: eurytopic, wet; phenology). Habitat. Autio and Salmela, 2010: 48, 50 (habitats Aland Is). Habitat. Autio et al., 2013: 1128 (a wetland generalist, one of the species found in a study on species richness and rarity in a boreal mire). Habitat. Rasimus, 2018: 17 (on saproxylic craneflies in the Tampere region) (in Finnish). France Biology. Quindroit, 2020a: 33, 45-48 (annotated list with information on abundancy and habitat of the Tipuloidea of the Pays de la Loire regoin, distr) (in French). Habitat. Quindroit, 2020b: 141 (loc(s) Pas-de-Calais (62) with notes on habitats and distr) (in French). Habitat. Quindroit and Lemoine, 2021a: 25 (loc(s), resultats de deux campagnes de piegeage dans le Pas-de-Calais (62); see paper for details on habitat). Habitat. Quindroit, 2021a: 36 (loc(s), resultats d une campagne de piegeage dans le Pas-de-Calais (62); see paper for details on habitat). Habitat. Quindroit and Lemoine, 2022a: 30 (loc(s) Aisne (02) with notes on habitats and distr) (in French). Germany Habitat. Reusch and Hohmann, 2009: 129 (from spring brooks). Habitat. Reiff et al., 2015: 234 (loc(s) Baden-Wurttemberg with list of habitats). Great Britain Habitat. Godfrey, 2000a: 22. Habitat. Boardman, 2007: 71 (boggy sites, wet woodland, acid carr and seepages). Habitat. Kramer, 2008b: 7 (at boggy groud with Bog Asphodel). Habitat. Cranston and Drake, 2010: 175 (Phylidorea larvae can be found in marginal situations along standing waters [no species mentioned]). Habitat. Drake, 2011a: 25 (among the Diptera from a wet Alder [Alnus] wood in Devon). Habitat. Kramer, 2011j: 20 (a common marshland species). Habitat. Alexander, 2015: 3 (result of a study on sampling techniques in Cornwall, showing productive pitfall-trapping for moorland crane flies). Habitat. Stubbs, 2018b: 2 (prefers wet scrub carr). Biology. Wolton, 2022: 6-8 (on observations on the phenology and sex ratios of craneflies (Limoniidae) found in emergencetraps, showing phenology charts for six species of Limoniidae from four emergence traps running between the beginning of may and the end of october 2020 in a wet woodland at Locks Park Farm, Devon; see paper for details on this apparently univoltine species). Ireland Habitat. Ashe et al., 2005a: 317, 331-332, 335 (bred from Phragmites stem bases, citing Blackith et al., 1991). Italy Habitat. Stary, 2003a: 125. Lithuania Habitat. Podeniene, 2003a: 1-295 (larva, descr, figs, habitat) (in Lithuanian). Habitat. Podeniene, 2003b: 10, 22, 28 (larval habitat: mud in littoral zone of swampy lakes). Norway Habitat. Olsen and Andersen, 2021: 211-212 (loc(s) Innlandet, in former Hedmark, with habitats specified separately, for habitats see also Jonassen and Andersen, 2020). Poland Habitat. Wiedenska, 2014: 12 (on the occurence of Limoniidae in six selected plant communities in the Kamienica Valley, Gorce National Park, Western Carpathians, found only in bog spring with Caltha-Chaerophyllum) (in Polish, habitat tables also in English). Phenology. Wiedenska, 2015a: 52 (on the flight activity of Limoniidae in the Kamienica Valley, Gorce National Park, Western Carpathians) (in Polish, tables also in English). Habitat. Wiedenska, 2017a: 14 (faunistic and habitat data of species from the Gorce NP) (in Polish). Romania Habitat. Ujvarosi and Poti, 2006: 256 (habitat in Dupa Lunca marshland: predominantly swamp with bushes and trees). Habitat. Ujvarosi et al., 2011b: 108 (loc(s) Dupa Lanca marshy area near Voslobeni) (in Hungarian, species list with habitat information in English). Russia Habitat. Przhiboro, 2003: 364 (larvae in the water margin zone of an oligotrophic lake; rearing). Habitat. Gavryushin in litt., 2011f: (collected in a herbaceous vegetation with Urtica, Chamerion, umbellifers and grasses near a ditch). Sweden Habitat. Lindstrom and Fritz, 2015: 44, 48 (distributin and ecology in five fens in Hallands County) (in Schwedish). on characters and taxonomy (mainly from 2000 onward): Country not relevant Characters. Kramer, 2006b: 3 (figs, short cut to identification). Immatures. Krivosheina, M.G., 2010a: 19 (key to larvae of 6 Phylidorea species) (for key in English see Krivosheina, M.G., 2010b). Immatures. Podeniene and Gelhaus, 2010: 200-202 (larvae of the genus Phylidorea can be divided into the abdominalis, ferruginea and squalens groups based on the sclerotization of the spiracular field, the ferruginea group consists of 5 species, ferruginea, nervosa, fulvonervosa, nigronotata and temelskin). Phylogeny. Santos and Ribeiro, 2024: 4-14 (phylogentic analysis of the Idioptera-Eloeophila complex: outgroup taxon). Great Britain Characters. Kramer and Morris, 2022b: 12 (diagnosis, figs). Lithuania Immatures. Podeniene, 2003a: 1-295 (larva, descr, figs, habitat) (in Lithuanian). on distribution (mainly from 2000 onward): Austria Distribution. Reusch and Heiss, 2012: 333-342 (loc(s) Nat. Park Gesause, see PDF for months and altitudes). Belarus First record. Kolcsar et al., 2021b: 204 (loc(s) Minsk region). Bulgaria Distribution. Hubenov, 2016: 47 (annotated checklist Diptera of the Rila Mountains based on data from the literature). Distribution. Hubenov, 2017: 68 (on vertical distribution in the Pirin and Rila Mts; see paper for details on forests and subalpine-alpine zones, distr). Distribution. Hubenov, 2021a: 30 (checklist with summary of distr in Bulgaria, references). Distribution. Hubenov, 2025b: suppl: 3 (annotated list Diptera high Bulgarian mts, local distr, habitats and zoogeography). Channel Is Distribution. Stubbs, 2022b: 5 (on a collection of craneflies from Jersey, collected between 1985 and 2002 by Tony Warne). Czech Rep. Distribution. Stary et al., 2005a: 29 (loc(s)). Distribution. Stary, 2009i: webpage (checklist). Distribution. Stary et al., 2013b: 378 (on Limonidae from Vraz near Pisek, with general habitat description). Distribution. Stary and Vonicka, 2018: 51-52 (on Limoniidae and Pediciidae of two localities in northern Bohemia, with list of localities, altitudes and habitats). Denmark Distribution. Petersen and Jong, 2001b: 151 (checklist). Estonia Distribution. Stary, 2004f: (Fauna Europaea: added Estonia). Finland Distribution. Salmela, 2001a: 144, 146, 150 (loc(s)). Distribution. Salmela, 2002: 30-32 (loc(s) lake Iidesjarvi) (in Finnish). Distribution. Salmela, 2004: 9 (loc(s)). Distribution. Stary and Salmela, 2004: 47 (loc(s)). Distribution. Salmela and Ilmonen, 2005: 89 (loc(s)). Distribution. Salmela and Autio, 2007b: 54 (loc(s)). Distribution. Salmela et al., 2007b: 37, 45 (abundance, loc(s)). Distribution. Salmela, 2008: 10, 50 (49) (loc(s), known from all Finnish ecoregions). Distribution. Autio and Salmela, 2010: 48 (loc(s) Aland Is). Distribution. Salmela, 2010a: 8, 12 (survey fauna Tahmela spring in city of Tampere) (in Finnish). Distribution. Salmela, 2012b: 229, 239 (annotated list of Finnish crane flies). Distribution. Salmela, 2013: 18 (belongs to the most abundant and frequent cranefly species of Finland, see table above under Habitat). Distribution. Salmela and Petrasiunas, 2014: 28 (checklist Finnish Tipulomorpha). Distribution. Salmela et al., 2015: 106 (list of localities in extensive report on malaise-trapping of insects in conservation areas in Lapland in 2012–2014) (in Finnish). Distribution. Rasimus, 2018: 17 (on saproxylic craneflies in the Tampere region) (in Finnish). France Distribution. Quindroit, 2020a: 33, 45-48 (annotated list with information on abundancy and habitat of the Tipuloidea of the Pays de la Loire regoin, distr) (in French). Distribution. Quindroit, 2020b: 141 (loc(s) Pas-de-Calais (62) with notes on habitats and distr) (in French). Distribution. Quindroit and Lemoine, 2021a: 25 (loc(s), resultats de deux campagnes de piegeage dans le Pas-de-Calais (62); see paper for details on habitat). Distribution. Quindroit, 2021a: 36 (loc(s), resultats d une campagne de piegeage dans le Pas-de-Calais (62); see paper for details on habitat). Distribution. Quindroit and Lemoine, 2022a: 30 (loc(s) Aisne (02) with notes on habitats and distr) (in French). Distribution. Quindroit and Lemoine, 2022a: 38 (table specifying the presence in the five departments of the Hauts-de-France region). Distribution. Quindroit, 2022a: 25-28 (additions to the annotated list of the Tipuloidea of Pays de la Loire as given in Quindroit, 2020a). Distribution. Lemoine and Quindroit, 2023: 9 (loc(s) Vosges (88)). Distribution. Quindroit and Racine, 2025: 359 (table specifying the presence in the five departments of the Pays de la Loire region). Germany Distribution. Schacht, 1999: 134 (checklist Bayern). Distribution. Reusch and Oosterbroek, 2000: 156 (checklist German Bundeslander). Distribution. Reusch and Bellstedt, 2001b: 72 (checklist Thuringen). Distribution. Reusch et al., 2004: 112 (loc(s) Baden-Wurttemberg). Distribution. Schacht, 2005: 6 (checklist Bayern). Distribution. Reusch and Hohmann, 2009: 129 (loc(s) Sachsen-Anhalt). Distribution. Hable et al., 2010b: 92 (loc(s) Bayern). Distribution. Schacht, 2010: 17 (checklist Diptera Bayern). Distribution. Reiff et al., 2015: 234 (loc(s) Baden-Wurttemberg with list of habitats). Distribution. Stuke, 2019: 145 (checklist Niedersachsen and Bremen with reference to original sources). Distribution. Kehlmaier et al., 2023: 210 (loc(s) Saarland). Distribution. Stuke et al., 2024: 161 (loc(s) Niedersachsen). Great Britain Distribution. Godfrey, 2000a: 20, 22 (see paper for details of sites surveyed in Shropshire and into Wales). Distribution. Hewitt et al., 2005: 38 (loc(s) Cumbria) (as Limnophila). Distribution. Boardman, 2007: 71 (loc(s), map). Distribution. Kramer, 2008b: 7 (loc(s) Wales). Distribution. Skidmore, 2009: 128 (review of occurrence on the Western Isles of Scotland, distr). Distribution. Drake, 2011a: 25 (loc(s) Devon). Distribution. Kramer, 2011g: 2 (loc(s) Devon). Distribution. Kramer, 2011j: 20 (loc(s) Leicestershire and Rutland (VC55)). Distribution. Hewitt, 2014: (provisional checklist Cumbrian Diptera). Distribution. Kramer, 2014c: 4 (in table of cranefly species recorded at Shapwick Heath NNR, as given in H. Audcent, 1949 and/or collected by J. Kramer in 2013). Distribution. Wolton et al., 2014: 27 (among Diptera associated with a hedge in Devon). Distribution. Alexander, 2015: 3 (result of a study on sampling techniques in Cornwall, showing productive pitfall-trapping for moorland crane flies). Distribution. Drake, 2016: 20 (study on the relative importance to Diptera of pasture and ditch margins on Catcott Reserve, Somerset). Distribution. Brighton, 2017c: 28 (overall regional checklist Lancashire and Cheshire (VC58, 59 and 60) with notes on habitats and/or regional occurrence and/or status; see paper for details). Distribution. Stubbs, 2018b: 2 (loc(s) Staffordshire). Distribution. Kramer and Morris, 2022b: 12 (info on the species, on the distr and map of loc(s) Leicestershire and Rutland (VC55)). Distribution. Wolton, 2022: 6-8 (on observations on the phenology and sex ratios of craneflies (Limoniidae) found in emergencetraps, showing phenology charts for six species of Limoniidae from four emergence traps running between the beginning of may and the end of october 2020 in a wet woodland at Locks Park Farm, Devon; see paper for details on this apparently univoltine species). Hungary First record. Thalhammer: 1899: XXX (loc(s)). Distribution. Stary, 2001b: 200 (loc(s), first documented record for Hungary, distr). Ireland Distribution. Ashe et al., 2005a: 317, 331-332, 335 (review Irish records). Distribution. Ashe et al., 2008: 14 (additional Irish record(s), distr). Distribution. Chandler et al., 2008: 12 (checklist). Italy Distribution. Stary, 2003a: 125 (loc(s)). Kazakhstan First record. Devyatkov, 2020d: 271 (loc(s) East Kazakhstan, distr) (in Russian). Lithuania Distribution. Pakalniskis et al., 2006: 23 (checklist). Montenegro Distribution. Oosterbroek and Simova-Tosic, 2004: 446 (review literature). Netherlands Distribution. Jong and Oosterbroek, 2002b: 32 (checklist). Norway Distribution. Olsen et al., 2018: 158-159 (annotated checklist Nordic countries and for Norway according to the Strand regions). Distribution. Olsen and Andersen, 2021: 211-212 (loc(s) Innlandet, in former Hedmark, with habitats specified separately, for habitats see also Jonassen and Andersen, 2020). Poland Distribution. Wiedenska, 2007a: 76 (checklist). Distribution. Wiedenska, 2010: 135 (loc(s) Parku Krajobrazowego Wzniesien Lodzkich, Lodz). Distribution. Wiedenska, 2014: 12 (on the occurence of Limoniidae in six selected plant communities in the Kamienica Valley, Gorce National Park, Western Carpathians, found only in bog spring with Caltha-Chaerophyllum) (in Polish, habitat tables also in English). Distribution. Wiedenska, 2017a: 14 (faunistic and habitat data of species from the Gorce NP) (in Polish). Distribution. Wiedenska and Syratt, 2023: 132 (loc(s) Bieszczady Mountains, distr and remarks). Portugal First record. Eiroa and Baez, 2002a: 55-56 (checklist, loc(s) Guarda). Distribution. Oosterbroek et al., 2020: 327 (known from Guarda only, distr). Romania Distribution. Ujvarosi, 2005a: 247 (no details). Distribution. Ujvarosi and Poti, 2006: 256 (loc(s) eastern Carpathians). Distribution. Ujvarosi, 2007: 228-231 (checklist Limoniidae). Distribution. Ujvarosi et al., 2011b: 108 (loc(s) Dupa Lanca marshy area near Voslobeni) (in Hungarian, species list with habitat information in English). Russia Distribution. Polevoi and Humala, 2003: 70 (loc(s) RUN: Kareliya). Distribution. Przhiboro, 2003: 364 (loc(s) NW Russia). Distribution. Polevoi and Humala, 2005: 181 (RUN: list of Diptera of Vepsskaya Volost, Kareliya) (in Russian). Distribution. Pilipenko and Sidorenko, 2006b: 145 (checklist Kedrovaya Pad, distr) (in Russian). Distribution. Polevoi, 2006: 96 (loc(s) RUN: Kareliya) (in Russian). Distribution. Polevoi and Humala, 2007: 135 (RUN: list of Diptera of Chukozero area, Kareliya) (in Russian). Distribution. Humala and Polevoi, 2009: 66 (loc(s) RUN: Kareliya) (in Russian). Distribution. Pilipenko, 2009c: 332 (loc(s) Lazovsky Nature Reserve, Primorskiy kray). Distribution. Gavryushin in litt., 2011e: (loc(s) RUN: Murmanskaya oblast). Distribution. Gavryushin in litt., 2011f: (loc(s) RUN: Arkhangelskaya oblast). Distribution. Gavryushin in litt., 2013b: (loc(s) Shikotan, Kuril Is). Distribution. Jakovlev et al., 2014: 309 (loc(s) RUN: Kareliya). Distribution. Paramonov and Pilipenko, 2016: 98 (loc(s) RUC: Tverskaya oblast) (in Russian). Distribution. Paramonov, 2018a: 183 (loc(s) RUC: Chuvash Rep., distr) (in Russian). Distribution. Paramonov and Korobkov, 2019: 97 (loc(s) RUC: Tverskaya oblast, distr, citing various authors) (in Russian). Distribution. Pilipenko et al., 2020: 358 (loc(s) RUC: Mordoviya Rep., distr). Distribution. Kolcsar et al., 2021b: 204-205 (loc(s) RUE: Bashkortostan Rep.). Distribution. Humala and Polevoi, 2022: 33 (loc(s) RUN: Arkhangelskaya oblast). Distribution. Devyatkov and Dubatolov, 2025: 33 (loc(s) Khabarovskiy kray, Bolshekhekhtsirsky Nature Reserve, distr) (in Russian). Slovakia Distribution. Stary, 2009d: 26 (loc(s) Polana area). Distribution. Stary, 2009i: webpage (checklist). Spain First record. Stary and Oosterbroek, 2008: 10 (loc(s) Teruel, distr). Distribution. Oosterbroek, 2009c: 181 (loc(s) Pontevedra). Sweden Distribution. Lindstrom and Fritz, 2015: 44, 48 (distributin and ecology in five fens in Hallands County) (in Schwedish). Switzerland Distribution. Podenas et al., 2006: 146, 322 (map, vertical distr per thermic level). Distribution. Dufour and Merz, 2012: 354 (annotated checklist canton Geneva). on flight period (mainly from 2000 onward): Belarus Kolcsar et al., 2021b: month(s): 7. Bulgaria Hubenov, 2016: month(s): 8. Channel Is Stubbs, 2022b: month(s): 6. Czech Rep. Stary et al., 2005a: month(s): 5-6. Stary et al., 2013b: month(s): 5-6. Stary and Vonicka, 2018: month(s): 5-8(9). Finland Salmela, 2001a: month(s): 6-8. Salmela, 2008: month(s): 7. France Quindroit, 2020a: month(s): 5-7. Quindroit, 2020b: month(s): 7. Quindroit and Lemoine, 2022a: month(s): 6. Lemoine and Quindroit, 2023: month(s): 6. Germany Reusch et al., 2004: month(s): 6. Reiff et al., 2015: month(s): 5. Kehlmaier et al., 2023: month(s): 6. Stuke et al., 2024: month(s): 6. Great Britain Godfrey, 2000a: month(s): 6. Kramer, 2008b: month(s): 7. Kramer, 2011j: month(s): 6. Kramer, 2014c: month(s): 6. Stubbs, 2018b: month(s): 6. Kramer and Morris, 2022b: month(s): 5-8. Hungary Stary, 2001b: month(s): 6. Ireland Ashe et al., 2005a: month(s): 5-10. Ashe et al., 2008: month(s): 5-6. Kazakhstan Devyatkov, 2020d: month(s): 6-8. Norway Olsen and Andersen, 2021: month(s): 6-8. Poland Wiedenska, 2014: month(s): 8. Wiedenska, 2015a: month(s): 7-8. Wiedenska, 2017a: month(s): 7-8. Wiedenska and Syratt, 2023: month(s): 6-7. Portugal Eiroa and Baez, 2002a: month(s): 7. Romania Ujvarosi et al., 2011b: month(s): 5. Russia Przhiboro, 2003: month(s): 7. Pilipenko, 2009c: month(s): 7. Gavryushin in litt., 2011e: month(s): 7. Gavryushin in litt., 2011f: month(s): 8. Gavryushin in litt., 2013b: month(s): 7. Paramonov and Pilipenko, 2016: month(s): 6, 8. Paramonov, 2018a: month(s): 6-7. Paramonov and Korobkov, 2019: month(s): 6. Pilipenko et al., 2020: month(s): 8-9. Kolcsar et al., 2021b: month(s): 7. Devyatkov and Dubatolov, 2025: month(s): 7. Slovakia Stary, 2009d: month(s): 5-7, 9. Spain Stary and Oosterbroek, 2008: month(s): 7. Switzerland Podenas et al., 2006: month(s): 5-10. on altitude (mainly from 2000 onward): Belarus Kolcsar et al., 2021b: altitude: 161 m. Bulgaria Hubenov, 2016: altitude: 1876 m. Hubenov, 2017: altitude: 1876 m. Hubenov, 2021a: altitude: 1876 m. Hubenov, 2025b: altitude: 1876 m. Germany Reusch et al., 2004: altitude: 355 m. Reusch and Hohmann, 2009: altitude: 375-842 m. Reiff et al., 2015: altitude: 666 m. Stuke et al., 2024: altitude: 200-270 m. Italy Stary, 2003a: altitude: 1500-1600 m. Kazakhstan Devyatkov, 2020d: altitude: 852-1468 m. Poland Wiedenska, 2017a: altitude: 760-1100 m. Wiedenska and Syratt, 2023: altitude: 620-800 m. Romania Ujvarosi and Poti, 2006: altitude: 670 m. Russia Gavryushin in litt., 2011f: altitude: 25 m. Kolcsar et al., 2021b: altitude: 499-765 m. Devyatkov and Dubatolov, 2025: altitude: 100 m. Slovakia Stary, 2009d: altitude: 630-870 m. Spain Stary and Oosterbroek, 2008: altitude: 1500 m.