| Name: | Paradelphomyia fuscula (Loew, 1873) | | Publication: | Beschr. europ. Dipt. 3: 64 (as Cladura) | | Status: | Recognized taxon, Synonym(s): furcata (Kuntze, 1919). | | Classification: | Family Limoniidae Subfamily Limnophilinae
| | Descriptive note: | Also described as nov spec in Loew, 1873b: Berl. ent. Z. 17: 35. | | Keys: | Kramer, 2015g (European Paradelphomyia except the Caucasus); Boardman, 2016 (craneflies Shropshire); Stubbs and Kramer, 2016g; Stubbs, 2021 (British craneflies) |  |  |  |  |  |
habitus male | habitus male | habitus male | body part(s) head and thorax | body part(s) head and thorax |
plus 22 more images of hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, wing, wing, wing, habitat, habitat, habitat, habitat, habitat, habitat, habitat and habitat. | | | | Distribution: | Austria, Bulgaria, Croatia, Czech Rep., Denmark, Estonia, Finland, France, Germany, Great Britain, Hungary, Ireland, Lithuania, Norway, Poland, Romania, Slovakia, Sweden, Switzerland, Ukraine; Russia: RUW, RUC, RUE; Armenia, Iran. Note: see the manual for abbreviations: present-day Russia. | | Region(s): | Westpalaearctic | | General note: | For Finland check Salmela, 2012b, 2012c, and citations below. For Norway check Olsen et al., 2018, and citations below. For Great Britain and Ireland check Stubbs, 2021, and citations below. For possible records from Belgium see Dek and Peeters, 2023a. | | Map: | Ashe et al., 2005a (Ireland); Podenas et al., 2006 (Switzerland); Boardman, 2007 (Shropshire) | Figure references: | Hypopygium: Podenas et al., 2006; Kramer, 2015g (after Edwards, 1938); Kramer and Morris, 2022b Other figures: Kramer, 2015g (head and thorax)
Immature stages Larva: Podeniene, 2002; Podeniene, 2003a; Krivosheina and Krivosheina, 2011c
Miscellaneous Lundstrom and Fritz, 2015 (habitat)
| | | | Citations | | | Bulgaria Habitat. Hubenov, 2025b: suppl: 2 (annotated list Diptera high Bulgarian mts, local distr, habitats and zoogeography). | | Country not relevant Biology. Olsen et al., 2018: 143, 153 (to be found around springs (Reusch & Hohmann 2009, Salmela 2001, 2004), acid seepages in woodland (Boardman 2007, 2016), shaded streams (Godfrey 2000); larvae live in the riparian zone of small rivulets, between sand, silt and leaf litter, and littoral zone of lakes, mud, springy slopes of lakes and rivers, in bogs of low lying woody swamps, on swampy riversides, in littoral zone of swampy lakes (Noll 1985, Podeniene 2003); the period of flight is may-october, but in northern areas confined to late summer or autumn; see table 1 for habitat information of the Norwegian locality(ties)). | | Czech Rep. Habitat. Stary and Vonicka, 2018: 51 (on Limoniidae and Pediciidae of two localities in northern Bohemia, with list of localities, altitudes and habitats). | | Denmark Habitat. Byriel and Rojas, 2017: 118 (among the 101 species in a study on cranefly hotspots in unmanaged and managed forests) (in Danish). | | Finland Habitat. Salmela, 2001a: 143-146, 150 (a common species around springs in southern Finland (a common shredder in central european springs, citing Wagner et al., 1998; occurs also in small brooks, citing Mendl, 1978b); phenology: maximum occurence in august). Habitat. Salmela, 2004: 3, 6, 9 (species restricted to spring areas (crenobiont), probably most common in rheocrene and helocrene springs). | | France Habitat. Quindroit, 2023: 65 (capture en berge de ruisseau forestier mixte, dans une zone boueuse). | | Germany Habitat. Reusch and Hohmann, 2009: 129 (from spring brooks). Habitat. Reiff et al., 2015: 234 (loc(s) Baden-Wurttemberg with list of habitats). | | Great Britain Habitat. Godfrey, 2000a: 13, 20-22 (shaded streams). Habitat. Stubbs, 2003: 101, 110-111 (occurs very locally in wet both lowland and upland woods, usually at tiny seepages where bare wet humic mud is exposed; non-calcareous conditions are usually evident; larvae are assumed to be saprophagous, semiaquatic in wet mud; the paper includes further details on habitat management). Habitat. Hewitt et al., 2005: 38, 48 (associated with small shaded streams, larvae probably develop in shaded mud beside such streams). Habitat. Boardman, 2007: 70 (acid seepages in woodland, in Shropshire also from habitats with calcareous influences and collected together with P. ecalcarata, a limestone specialist). Habitat. Kramer, 2015g: 52 (a wet woodland species). Habitat. Brighton, 2017c: 27, 37 (overall regional checklist Lancashire and Cheshire (VC58, 59 and 60) with notes on habitats and/or regional occurrence and/or status; see paper for details). Habitat. Drake, 2022: 3 (collected in Devon: Knapp Copse, SY156953, 11 october 2021 in a small patch of possibly slightly acidic hillside seepage under sparse Sallow [Salix cinerea] woodland). | | Hungary Habitat. Kolcsar et al., 2021b: 200 (Oak [Quercus] forest). | | Lithuania Habitat. Podeniene and Gelhaus, 2002: 69 (larvae in the riparian zone of small rivulet, between sand, silt and leaf litter). Habitat. Podeniene, 2002: 301-306 (larvae were found in mud among roots of Phragmites and other plants on lake shore (littoral zone habitat Phragmitetalia), in mud of low-laying marsh (Alnetalia glutinosae) on boggy lakeshore, in the riparian zone of a small rivulet, (and among moss in damp areas, citing Podenas); adults on the banks of water bodies and wet Alnus groves). Habitat. Podeniene, 2003a: 1-295 (larva, descr, figs, habitat) (in Lithuanian). Habitat. Podeniene, 2003b: 10, 21, 22, 28 (larval habitat: sand and silt in riparian zone of small rivulets and littoral zone of lakes, mud (in springy slopes of lakes and rivers, in bogs of low lying woody swamps, on swampy riversides, in littoral zone of swampy lakes)). | | Poland Habitat. Chaniecka and Wiedenska, 2006: 145, 147 (occurrence in spring-, springbrook-, and/or bogspring plant communities) (in Polish). Biology. Wiedenska, 2014: 12 (on the occurence of Limoniidae in six selected plant communities in the Kamienica Valley, Gorce National Park, Western Carpathians; observed emergence of adults in august) (in Polish, habitat tables also in English). Biology. Wiedenska, 2015a: 51, 52 (on the flight activity of Limoniidae in the Kamienica Valley, Gorce National Park, Western Carpathians, with flight diagrams; observed emergence of adults in august; new correct flight diagrams are given in Wiedenska, 2017b) (in Polish, tables also in English). Habitat. Wiedenska, 2017a: 12 (faunistic and habitat data of species from the Gorce NP) (in Polish). | | Romania Habitat. Ujvarosi, 2005a: 247 (brooks, hygropetric zone, swamps, citing various sources). | | Russia Habitat. Kolcsar et al., 2021b: 201 (Tatarstan Rep.: ravine, wetland). | | Sweden Habitat. Lindstrom and Fritz, 2015: 39, 44 48 (on distribution, ecology, habitat preference and habitat conservation in a method study and species inventory in five fens in Hallands County) (in Schwedish). | | | | Country not relevant Phylogeny. Ahonen, 2007: 1-28 (included in phylogenetic analysis of Tipuloidea subfamilies relationships based on molecular data). Characters. Olsen et al., 2018: 143 (to be identified by a combination of wing and male genital characters as mentioned in the keys by Kramer (2015) and Stubbs & Kramer (2016a), most distinctly by the male apodeme having elongated longitudinal processes without lateral arms; fig). | | Great Britain Revision. Kramer, 2015g: 43-55 (review British species, identification, comparison, key, figs). Characters. Kramer and Morris, 2022b: 10 (diagnosis, figs). | | Lithuania Immatures. Podeniene, 2002: 301-306 (descr larva, comparison, figs). Immatures. Podeniene, 2003a: 1-295 (larva, descr, figs, habitat) (in Lithuanian). | | | | Armenia First record. Obona et al., 2016: 128 (loc(s)). Distribution. Hakobyan and Jenderedjian, 2023: 1511 (annotated checklist of the craneflies of Armenia with refs, distr and flight period in Caucasus Ecoregion with refs). | | Bulgaria Distribution. Hubenov, 2016: 47 (annotated checklist Diptera of the Rila Mountains based on data from the literature). Distribution. Hubenov, 2017: 67 (on vertical distribution in the Pirin and Rila Mts; see paper for details on forests and subalpine-alpine zones, distr). Distribution. Hubenov, 2021a: 29 (checklist with summary of distr in Bulgaria, references). Distribution. Hubenov, 2025b: suppl: 2 (annotated list Diptera high Bulgarian mts, local distr, habitats and zoogeography). | | Croatia First record. Kolcsar et al., 2023b: 28 (loc(s)). | | Czech Rep. Distribution. Stary, 2009i: webpage (checklist). Distribution. Stary and Vonicka, 2018: 51 (on Limoniidae and Pediciidae of two localities in northern Bohemia, with list of localities, altitudes and habitats). | | Denmark Distribution. Petersen and Jong, 2001b: 151 (checklist). | | Estonia First record. Salmela in litt., 2003: (new for Estonia). | | Finland Distribution. Salmela, 2001a: 143-146, 150 (loc(s)). Distribution. Salmela, 2004: 3, 6, 9 (loc(s), distr). Distribution. Salmela et al., 2007b: 45 (loc(s)). Distribution. Salmela, 2010a: 8, 12 (survey fauna Tahmela spring in city of Tampere) (in Finnish). Distribution. Salmela, 2012b: 239 (annotated list of Finnish crane flies). Distribution. Salmela and Petrasiunas, 2014: 28 (checklist Finnish Tipulomorpha). | | France First record. Quindroit, 2023: 65 (loc(s) Isere (38), distr). | | Germany Distribution. Schacht, 1999: 134 (checklist Bayern). Distribution. Reusch and Oosterbroek, 2000: 156 (checklist German Bundeslander). Distribution. Reusch and Bellstedt, 2001b: 72 (checklist Thuringen). Distribution. Schacht, 2005: 6 (checklist Bayern). Distribution. Reusch and Hohmann, 2009: 129 (loc(s) Sachsen-Anhalt). Distribution. Schacht, 2010: 17 (checklist Diptera Bayern). Distribution. Reiff et al., 2015: 234 (loc(s) Baden-Wurttemberg with list of habitats). Distribution. Stuke, 2019: 145 (checklist Niedersachsen and Bremen with reference to original sources). | | Great Britain Distribution. Godfrey, 2000a: 13, 20-22 (see paper for details of sites surveyed in Shropshire and into Wales). Distribution. Stubbs, 2003: 110 (distr in GB). Distribution. Hewitt et al., 2005: 38, 48 (loc(s) Cumbria). Distribution. Boardman, 2007: 70 (loc(s), map). Distribution. Drake and Godfrey, 2010: 15 (comparison). Distribution. Morris, 2010a: 15 (loc(s) Scottish Borders). Distribution. Whiteley, 2010: 4 (loc(s) Derbyshire and/or Nottinghamshire). Distribution. Kramer, 2011j: 19 (loc(s) Leicestershire and Rutland (VC55)). Distribution. Hewitt, 2014: (provisional checklist Cumbrian Diptera). Distribution. Kramer, 2015g: 51-52, 57 (review British distr, loc(s), map). Distribution. Brighton, 2017c: 27, 37 (overall regional checklist Lancashire and Cheshire (VC58, 59 and 60) with notes on habitats and/or regional occurrence and/or status; see paper for details). Distribution. Cunningham, 2022: 30 (loc(s) Knapp Copse, Devon). Distribution. Drake, 2022: 3 (collected in Devon: Knapp Copse, SY156953, 11 october 2021 in a small patch of possibly slightly acidic hillside seepage under sparse Sallow [Salix cinerea] woodland). Distribution. Kramer and Morris, 2022b: 10 (info on the species, on the distr and map of loc(s) Leicestershire and Rutland (VC55)). | | Hungary First record. Kolcsar et al., 2021b: 200 (loc(s) Heves and Pest). | | Ireland Distribution. Ashe et al., 2005a: 317, 330, 336 (review Irish records). Distribution. Chandler et al., 2008: 12 (checklist). | | Lithuania Distribution. Podeniene and Gelhaus, 2002: 69 (loc(s)). Distribution. Podeniene, 2002: 301-306 (loc(s)). Distribution. Pakalniskis et al., 2006: 23 (checklist). | | Norway First record. Olsen et al., 2018: 143, 158-159 (loc(s) in four regions; distr; annotated checklist Nordic countries and for Norway according to the Strand regions). | | Poland Distribution. Chaniecka and Wiedenska, 2006: 145, 147 (loc(s)). Distribution. Wiedenska, 2007a: 76 (checklist). Distribution. Wiedenska, 2010: 135 (loc(s) Parku Krajobrazowego Wzniesien Lodzkich, Lodz). Distribution. Wiedenska, 2014: 12 (on the occurence of Limoniidae in six selected plant communities in the Kamienica Valley, Gorce National Park, Western Carpathians) (in Polish, habitat tables also in English). Distribution. Wiedenska, 2017a: 12 (faunistic and habitat data of species from the Gorce NP) (in Polish). | | Romania Distribution. Ujvarosi, 2007: 228-231 (checklist Limoniidae). Distribution. Keresztes, 2012: 31 (loc(s) Rimetea area [Alba County]). | | Russia Distribution. Kolcsar et al., 2021b: 200-201 (loc(s) RUC an RUE). | | Slovakia Distribution. Stary, 2009d: 25 (loc(s) Polana area). Distribution. Stary, 2009i: webpage (checklist). | | Sweden Distribution. Lindstrom and Fritz, 2015: 39, 44 48 (on distribution, ecology, habitat preference and habitat conservation in a method study and species inventory in five fens in Hallands County) (in Schwedish). | | Switzerland Distribution. Podenas et al., 2006: 145, 320 (map, vertical distr per thermic level). | | | | Armenia Obona et al., 2016: month(s): 8. | | Bulgaria Hubenov, 2016: month(s): 9. | | Croatia Kolcsar et al., 2023b: month(s): 6. | | Czech Rep. Stary and Vonicka, 2018: month(s): 8-9(10). | | Finland Salmela, 2001a: month(s): 7-8, 10. | | France Quindroit, 2023: month(s): 9. | | Germany Reiff et al., 2015: month(s): 9. | | Great Britain Godfrey, 2000a: month(s): 6. Stubbs, 2003: month(s): 7-10. Boardman, 2007: month(s): 8-10. Morris, 2010a: month(s): 9. Whiteley, 2010: month(s): 7. Kramer, 2011j: month(s): 6. Kramer, 2015g: month(s): 5-9. Cunningham, 2022: month(s): 10. Kramer and Morris, 2022b: month(s): 8-10. | | Hungary Kolcsar et al., 2021b: month(s): 7, 9. | | Ireland Ashe et al., 2005a: month(s): 7, 9. | | Lithuania Podeniene, 2002: month(s): 6-10. | | Norway Olsen et al., 2018: month(s): 7-9. | | Poland Wiedenska, 2014: month(s): 8-10. Wiedenska, 2015a: month(s): 8-9. Wiedenska, 2017a: month(s): 8-9. | | Russia Kolcsar et al., 2021b: month(s): 8. | | Slovakia Stary, 2009d: month(s): 9. | | Switzerland Podenas et al., 2006: month(s): 6-9. | | | | Armenia Obona et al., 2016: altitude: 1872 m. | | Bulgaria Hubenov, 2016: altitude: 1147 m. Hubenov, 2017: altitude: 1147 m. Hubenov, 2021a: altitude: 1147 m. Hubenov, 2025b: altitude: 1147 m. | | Croatia Kolcsar et al., 2023b: altitude: 630 m. | | France Quindroit, 2023: altitude: 1300 m. | | Germany Reusch and Hohmann, 2009: altitude: 375-550 m. Reiff et al., 2015: altitude: 666 m. | | Poland Wiedenska, 2017a: altitude: 800-1230 m. | | Russia Kolcsar et al., 2021b: altitude: 80-220 m. |
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