Details for Dicranophragma (Brachylimnophila) separatum

Name: Dicranophragma (Brachylimnophila) separatum (Walker, 1848) Publication: List Dipt. Colln Brit. Mus. 1: 56 (as Limnobia) Status: Recognized taxon. Classification: Family Limoniidae Subfamily Limnophilinae Revision: Stary and Reusch, 2009. Descriptive note: Species status after Stubbs, 1995b, 1997c; Stary and Reusch, 2009. Lectotype designation: Stary and Reusch, 2009. Keys: Boardman, 2016 (craneflies Shropshire); Peeters en Oosterbroek, 2016c (craneflies Netherlands, Belgium and Luxembourg) (in Dutch); Stubbs and Kramer, 2016g; Stubbs, 2021 (British craneflies) Variant spelling(s): separata habitus male habitus male habitus male hypopygium hypopygium plus 26 more images of hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, ovipositor, wing, wing, wing, habitat, habitat, habitat, habitat, habitat, habitat, habitat, habitat, habitat, habitat, habitat, habitat, habitat, habitat, habitat, habitat and habitat. Distribution: Austria, Belarus, Bulgaria, Czech Rep., Denmark, Finland, France, Germany, Great Britain, Hungary, Latvia, Netherlands, Norway, Poland, Romania, Serbia, Slovakia, Sweden (south), Switzerland, Ukraine; Russia: RUN, RUC, RUE; North Caucasus, FE (Khabarovskiy kray); Georgia, Azerbaijan;; Kazakhstan (east), Tajikistan. Note: see the manual for abbreviations: present-day Russia. Region(s): Westpalaearctic;; Eastpalaearctic General note: For Finland check Salmela, 2012b, 2012c, and citations below. For Norway check Olsen et al., 2018, and citations below. For Great Britain check Stubbs, 2021, and citations below. Map: Boardman, 2007 (Shropshire) Figure references: Habitus: Tillier, 2021b, 2025d (male) Hypopygium: Stary and Reusch, 2009; Tillier, 2021b; Kramer and Morris, 2022b Citations on biology (mainly from 2000 onward): Austria Habitat. Haseke and Remschak, 2021: 22 (from a tufa spring). Habitat. Aistleitner et al., 2024: 32 (loc(s) Vorarlberg, see habitat photo above). Azerbaijan Habitat. Stary and Obona, 2020: 131 (loc(s); with list of localities, habitats and altitudes). Bulgaria Habitat. Hubenov, 2025b: suppl: 3 (annotated list Diptera high Bulgarian mts, local distr, habitats and zoogeography). Czech Rep. Habitat. Stary and Vonicka, 2018: 49 (on Limoniidae and Pediciidae of two localities in northern Bohemia, with list of localities, altitudes and habitats). Finland Habitat. Salmela, 2001a: 143-146, 150 (a common species around springs in southern Finland (species with a wide range of habitats, both terrestrial and aquatic, can attain relatively high densities in spring habitats, citing various sources)) (as nemoralis [= nemorale], misidentification, see Salmela, 2012b: 225, 228). Biology. Salmela, 2008: 10, 24, 27, 33, 50 (49) (ecology or habitat: eurytopic, terrestrial; phenology) (as nemoralis [= nemorale], misidentification, see Salmela, 2012b: 225, 228). Habitat. Autio and Salmela, 2010: 48, 51 (habitats Aland Is, the larvae of this typical brook- and ditch-inhabiting species can probably tolerate long dry periods). Habitat. Salmela, 2011a: 1-28 (study on three wetland habitats: aapa mires, springs and headwater streams; found in headwater streams only). France Habitat. Quindroit, 2020b: 135 (loc(s) Nord (59) and Pas-de-Calais (62) with notes on habitats and distr) (in French) (as nemorale, see Quindroit and Lemoine, 2023). Habitat. Quindroit and Lemoine, 2021a: 23, 25 (loc(s), resultats de deux campagnes de piegeage dans le Pas-de-Calais (62); see paper for details on habitat) (as nemorale, see Quindroit and Lemoine, 2023). Habitat. Quindroit, 2021a: 36 (loc(s), resultats d une campagne de piegeage dans le Pas-de-Calais (62); see paper for details on habitat) (as nemorale, see Quindroit and Lemoine, 2023). Habitat. Quindroit, 2021c: 424, 426 (loc(s) Mercantour National Park, Alpes-Maritimes (06), common species; from woodland, often humid). Habitat. Tillier, 2021b: 282 (dans une zone marecageuse en sous-bois; au niveau d une zone de sources en sous-bois, presentant de nombreuses zones boueuses). Phenology. Tillier, 2021b: 282 (discussion on date of collecting in april, which is early compared to the known period of flight) (in French). Habitat. Quindroit and Lemoine, 2022a: 29 (loc(s) Oise (60) and Somme (80) with notes on habitats and distr) (in French). Habitat. Quindroit, 2022a: 14 (in relation to D. nemorale), 19, 27 (annotated list with information on abundancy and habitat of the Tipuloidea of the Pays de la Loire regoin, distr) (in French). Habitat. Quindroit, 2022d: 417 (loc(s) Ariege (09); prairie avec une zone suintante ouverte; loc(s) Pyrenees-Orientales (66); sous-bois berge de ruisseau; remarks on distribution in France, also in relation to D. nemorale). Georgia Habitat. Stary and Obona, 2020: 131 (loc(s); with list of localities, habitats and altitudes). Germany Habitat. Reusch and Oosterbroek, 2000: 151, 156 (Quellbereich). Habitat. Ruckert, 2005: 47, 53 (found in a study on the association with the type of source: limnokrene, rheokrene, or helokrene in Schlesig-Holstein; restricted to limnokrene sources; see paper for details, as well as Anhang A2-A10). Habitat. Wittrock, 2005: 84 (summer emergence; limnokrene). Habitat. Reusch, 2006: 133-135 (Kalkquellmoor [carbonate fen complex]). Habitat. Reusch and Hohmann, 2009: 128, 129 (from spring brooks). Great Britain Habitat. Boardman, 2007: 68 (most typically on upland peaty moorland, though lower peaty soils and boggy ground also offer habitat). Habitat. Kramer, 2011f: 2 (at a chalk-pit stream and marsh). Habitat. Brighton, 2017c: 27, 36 (overall regional checklist Lancashire and Cheshire (VC58, 59 and 60) with notes on habitats and/or regional occurrence and/or status; see paper for details). Hungary Habitat. Kolcsar et al., 2021b: 76 (Hornbeam-Oak [Carpinus-Quercus] forest). Latvia Habitat. Kolcsar et al., 2021b: 77 (Birch-Spruce [Betula-Picea] forest, small stream). Norway Habitat. Olsen and Andersen, 2021: 210-211 (loc(s) Innlandet, in former Hedmark, with habitats specified separately, for habitats see also Jonassen and Andersen, 2020). Romania Habitat. Kolcsar et al., 2013: 71 (loc(s) Cluj-Napoca and surroundings; list of habitats with altitudes). Russia Habitat. Gavryushin in litt., 2011b: (loc(s) RUC: Moskovskaya oblast; collected in a herbaceous vegetation with Urtica, Chamerion, umbellifers and grasses near a ditch). Habitat. Lantsov, 2017b: 107 (on dominant species of the crane flies (Diptera, Tipuloidea) of the North Caucasus as biological indicators of habitats) (in Russian). Habitat. Paramonov and Korobkov, 2019: 96-97 (loc(s) RUC: Tverskaya oblast, distr, habitat, citing various authors) (in Russian). Sweden Habitat. Lindstrom and Fritz, 2015: 44, 47 (distributin and ecology in five fens in Hallands County) (in Schwedish). on characters and taxonomy (mainly from 2000 onward): Country not relevant Characters. Kato and Tachi, 2018: 33 (comparison). Characters. Quindroit and Lemoine, 2022a: 29 (comparison with nemorale) (in French). France Characters. Quindroit and Lemoine, 2023: 13 (detailed discussion on the discrimating characters between nemorale and separatum) (in French). Great Britain Characters. Kramer and Morris, 2022b: 5 (diagnosis, figs). Norway Revision. Stary and Reusch, 2009: 207-216 (revision, type-mat, lectotype des, systematic position, descr, comparison, figs). on distribution (mainly from 2000 onward): Austria First record. Reusch and Heiss, 2012: 168-169, 314-322 (loc(s) Nat. Park Gesause, Steiermark). Distribution. Vogtenhuber and Kofler, 2017: 138 (loc(s) Osttirol). Distribution. Wagner et al., 2018: 118 (report on the fourth insect camp of the Entomological Society of Austria (ESA) from the Biosphere Reserve Nockberge, Carinthia (in German). Distribution. Haseke and Remschak, 2021: 22 (loc(s) Nat. Park Gesause, Steiermark). Distribution. Aistleitner et al., 2024: 32 (loc(s) Vorarlberg, see habitat photo above). Azerbaijan First record. Stary and Obona, 2020: 131 (loc(s); with list of localities, habitats and altitudes). Distribution. Snegovaya, 2021: 1145 (checklist with details on distr in Azerbaijan). Belarus First record. Kolcsar et al., 2021b: 75-76 (loc(s) Minsk and Gomel regions). Bulgaria First record. Stary and Reusch, 2009: 213 (loc(s), distr). Distribution. Hubenov, 2021a: 30 (checklist with summary of distr in Bulgaria, references). Distribution. Hubenov, 2025b: suppl: 3 (annotated list Diptera high Bulgarian mts, local distr, habitats and zoogeography). Czech Rep. First record. Stary and Reusch, 2009: 213 (loc(s), distr). Distribution. Stary, 2009i: webpage (checklist). Distribution. Stary and Vonicka, 2018: 49 (on Limoniidae and Pediciidae of two localities in northern Bohemia, with list of localities, altitudes and habitats). Denmark First record. Salmela, 2012d: 121 (loc(s)). Finland Distribution. Salmela, 2001a: 143-146, 150 (loc(s)) (as nemoralis [= nemorale], misidentification, see Salmela, 2012b: 225, 228). Distribution. Salmela, 2002: 30-32 (loc(s) lake Iidesjarvi) (as nemoralis [= nemorale], misidentification, see Salmela, 2012b: 225, 228). Distribution. Salmela et al., 2007b: 37, 45 (abundance, loc(s)) (as nemoralis [= nemorale], misidentification, see Salmela, 2012b: 225, 228). Distribution. Salmela, 2008: 10, 50 (49) (loc(s), known from all Finnish ecoregions) (as nemoralis [= nemorale], misidentification, see Salmela, 2012b: 225, 228). Distribution. Autio and Salmela, 2010: 48 (loc(s) Aland Is). First record. Salmela, 2010a: 12 (survey fauna Tahmela spring in city of Tampere) (in Finnish). Distribution. Salmela, 2011a: 24 (loc(s) subalpine northern boreal Finland). Distribution. Salmela, 2012b: 225, 237 (annotated list of Finnish crane flies; one of the commonest crane flies, known from all Finnish biogeographical regions). Distribution. Salmela, 2013: 18 (belongs to the most abundant and frequent cranefly species of Finland, see table above under Habitat). Distribution. Salmela and Petrasiunas, 2014: 27 (checklist Finnish Tipulomorpha). Distribution. Salmela et al., 2015: 101 (list of localities in extensive report on malaise-trapping of insects in conservation areas in Lapland in 2012–2014) (in Finnish). France Distribution. Quindroit, 2020b: 135 (loc(s) Nord (59) and Pas-de-Calais (62) with notes on habitats and distr) (in French) (as nemorale, see Quindroit and Lemoine, 2023). Distribution. Quindroit and Lemoine, 2021a: 23, 25 (loc(s), resultats de deux campagnes de piegeage dans le Pas-de-Calais (62); see paper for details on habitat) (as nemorale, see Quindroit and Lemoine, 2023). Distribution. Quindroit, 2021a: 36 (loc(s), resultats d une campagne de piegeage dans le Pas-de-Calais (62); see paper for details on habitat) (as nemorale, see Quindroit and Lemoine, 2023). Distribution. Quindroit, 2021c: 424, 426 (loc(s) Mercantour National Park, Alpes-Maritimes (06), common species; from woodland, often humid). First record. Tillier, 2021b: 281-282 (loc(s) Valle-d’Oise (95), distr). Distribution. Quindroit and Lemoine, 2022a: 29 (loc(s) Oise (60) and Somme (80) with notes on habitats and distr) (in French). Distribution. Quindroit and Lemoine, 2022a: 37 (table specifying the presence in the five departments of the Hauts-de-France region). Distribution. Quindroit, 2022a: 14 (in relation to D. nemorale), 19, 27 (annotated list with information on abundancy and habitat of the Tipuloidea of the Pays de la Loire regoin, distr) (in French). Distribution. Quindroit, 2022d: 417 (loc(s) Ariege (09); prairie avec une zone suintante ouverte; loc(s) Pyrenees-Orientales (66); sous-bois berge de ruisseau; remarks on distribution in France, also in relation to D. nemorale). Distribution. Quindroit and Racine, 2025: 359 (table specifying the presence in the five departments of the Pays de la Loire region). Georgia First record. Stary and Obona, 2020: 131 (loc(s); with list of localities, habitats and altitudes). Germany First record. Reusch and Oosterbroek, 2000: 151, 156 (checklist German Bundeslander). Distribution. Schumann, 2003: 438 (1e addition checklist DE). Distribution. Reusch et al., 2004: 111-112 (loc(s) four Bundeslander). Distribution. Ruckert, 2005: 47, 53 (found in a study on the association with the type of source: limnokrene, rheokrene, or helokrene in Schlesig-Holstein; restricted to limnokrene sources; see paper for details, as well as Anhang A2-A10). Distribution. Wittrock, 2005: 84 (loc(s) Schleswig-Holstein). Distribution. Reusch, 2006: 133-135 (loc(s) Bayern). Distribution. Ruckert et al., 2007: 56 (loc(s) Schleswig-Holstein). Distribution. Reusch and Hohmann, 2009: 128, 129 (loc(s) Sachsen-Anhalt). Distribution. Stary and Reusch, 2009: 213-214 (loc(s), distr). Great Britain Distribution. Boardman, 2007: 68 (loc(s), map). Distribution. Phillips, 2007: 150 (loc(s) Isle of Rum) (as Limnophila (Brachylimnophila) separata sensu Stubbs). Distribution. Skidmore, 2009: 127 (review of occurrence on the Western Isles of Scotland, distr). Distribution. Stary and Reusch, 2009: 213-214 (loc(s), distr). Distribution. Kramer, 2011f: 2 (loc(s) Devon). Distribution. Robinson et al., 2016: 80 (list of terrestrial invertebrates of Mingulay Is, Outer Hebrides). Distribution. Brighton, 2017c: 27, 36 (overall regional checklist Lancashire and Cheshire (VC58, 59 and 60) with notes on habitats and/or regional occurrence and/or status; see paper for details). Distribution. Kramer and Morris, 2022b: 5 (info on the species, on the distr and map of loc(s) Leicestershire and Rutland (VC55)). Hungary First record. Kolcsar et al., 2021b: 76 (loc(s) Baranya). Kazakhstan First record. Devyatkov, 2019b: 245 (loc(s) Pavlodarskaya oblast, distr) (in Russian). Distribution. Devyatkov, 2020d: 269 (loc(s) East Kazakhstan, distr) (in Russian). Latvia First record. Kolcsar et al., 2021b: 76-77 (loc(s)). Netherlands First record. Beuk in litt., 2018: (collected in De Meinweg in Limburg, corroborated by study of the male genitalia). Distribution. Beuk in litt., 2021: (collected at Hazelbekke near Ootmarsum in Overijssel). Norway Distribution. Stary and Reusch, 2009: 212-213 (type-loc(s), distr). Distribution. Olsen et al., 2018: 158-159 (annotated checklist Nordic countries and for Norway according to the Strand regions). Distribution. Olsen and Andersen, 2021: 210-211 (loc(s) Innlandet, in former Hedmark, with habitats specified separately, for habitats see also Jonassen and Andersen, 2020). Poland First record. Kolcsar et al., 2021b: 77 (loc(s) Lower Silesia). Distribution. Wiedenska and Syratt, 2023: 132 (loc(s) Bieszczady Mountains, distr and remarks). Romania First record. Kolcsar et al., 2013: 71 (loc(s) Cluj-Napoca and surroundings; list of habitats with altitudes). Russia First record. Gavryushin in litt., 2011b: (loc(s) RUC: Moskovskaya oblast; collected in a herbaceous vegetation with Urtica, Chamerion, umbellifers and grasses near a ditch). Distribution. Lantsov, 2016: 165-170 (on craneflies from Arkhyz, Karachay-Cherkessia Rep., North Caucasus) (in Russian). Distribution. Lantsov, 2017b: 107 (on dominant species of the crane flies (Diptera, Tipuloidea) of the North Caucasus as biological indicators of habitats) (in Russian). Distribution. Paramonov, 2017a: 172 (loc(s) RUC: Chuvash Rep.) (in Russian). Distribution. Paramonov and Korobkov, 2019: 96-97 (loc(s) RUC: Tverskaya oblast, distr, habitat, citing various authors) (in Russian). Distribution. Kolcsar et al., 2021b: 77-80 (loc(s) RUN and RUE). Distribution. Devyatkov and Dubatolov, 2025: 32 (loc(s) Khabarovskiy kray, Bolshekhekhtsirsky Nature Reserve, distr) (in Russian). Serbia First record. Kolcsar et al., 2021b: 80 (loc(s)). Slovakia First record. Stary and Reusch, 2009: 214 (loc(s), distr). Distribution. Stary, 2009d: 24 (loc(s) Polana area). Distribution. Stary, 2009i: webpage (checklist). Sweden First record. Salmela, 2010b: 129, 133 (loc(s) Malmo, distr; additional records from south Sweden in Artportalen). Distribution. Fritz and Lindstrom, 2013: 62 (loc(s) Hallands County) (in Schwedish). Distribution. Lindstrom and Fritz, 2015: 44, 47 (distributin and ecology in five fens in Hallands County) (in Schwedish). Switzerland First record. Stary and Reusch, 2009: 214 (loc(s), distr). Tajikistan First record. Devyatkov, 2020c: 76 (loc(s), distr) (in Russian). on flight period (mainly from 2000 onward): Austria Reusch and Heiss, 2012: month(s): 6-8. Vogtenhuber and Kofler, 2017: month(s): 7. Wagner et al., 2018: month(s): 7. Haseke and Remschak, 2021: month(s): 7. Aistleitner et al., 2024: month(s): 7. Azerbaijan Stary and Obona, 2020: month(s): 5. Belarus Kolcsar et al., 2021b: month(s): 6-7. Bulgaria Stary and Reusch, 2009: month(s): 7. Czech Rep. Stary and Reusch, 2009: month(s): 5-8. Stary and Vonicka, 2018: month(s): (5)6-8. Denmark Salmela, 2012d: month(s): 7. Finland Salmela, 2001a: month(s): 6-7. Salmela, 2008: month(s): 6-8. France Quindroit, 2020b: month(s): 5-7. Quindroit and Lemoine, 2021a: month(s): 5-7. Quindroit, 2021a: month(s): 5-7. Tillier, 2021b: month(s): 4. Quindroit and Lemoine, 2022a: month(s): 5-7. Quindroit, 2022a: month(s): 5-7. Quindroit, 2022d: month(s): 7. Georgia Stary and Obona, 2020: month(s): 5, 7. Germany Reusch and Oosterbroek, 2000: month(s): 8. Reusch et al., 2004: month(s): 6-8. Stary and Reusch, 2009: month(s): 6-8. Great Britain Stary and Reusch, 2009: month(s): 7. Kramer, 2011f: month(s): 7. Hungary Kolcsar et al., 2021b: month(s): 6-7. Kazakhstan Devyatkov, 2019b: month(s): 6. Devyatkov, 2020d: month(s): 6-9. Latvia Kolcsar et al., 2021b: month(s): 7. Norway Olsen and Andersen, 2021: month(s): 6-8(9). Poland Kolcsar et al., 2021b: month(s): 5. Wiedenska and Syratt, 2023: month(s): 6-7. Romania Kolcsar et al., 2013: month(s): 6. Russia Gavryushin in litt., 2011b: month(s): 6-7. Paramonov, 2017a: month(s): 6. Paramonov and Korobkov, 2019: month(s): 7. Kolcsar et al., 2021b: month(s): 7-8. Devyatkov and Dubatolov, 2025: month(s): 6, 8-9. Serbia Kolcsar et al., 2021b: month(s): 6. Slovakia Stary and Reusch, 2009: month(s): 5-7, 9. Stary, 2009d: month(s): 5-7. Switzerland Stary and Reusch, 2009: month(s): 7. Tajikistan Devyatkov, 2020c: month(s): 6. on altitude (mainly from 2000 onward): Austria Reusch and Heiss, 2012: altitude: 845-2015 m. Vogtenhuber and Kofler, 2017: altitude: 1800-2120 m. Haseke and Remschak, 2021: altitude: 1095 m. Aistleitner et al., 2024: altitude: 2020 m. Azerbaijan Stary and Obona, 2020: altitude: 595 m. Belarus Kolcsar et al., 2021b: altitude: 155-161 m. Bulgaria Stary and Reusch, 2009: altitude: over 2000 m. Hubenov, 2021a: altitude: 2000 m. Hubenov, 2025b: altitude: 2000 m. Czech Rep. Stary and Reusch, 2009: altitude: 600-1050 m. France Quindroit, 2021c: altitude: 2000 m. Tillier, 2021b: altitude: 40-45 m. Quindroit, 2022d: altitude: 1900 m. Germany Reusch et al., 2004: altitude: 20-700 m. Reusch and Hohmann, 2009: altitude: 375-842 m. Stary and Reusch, 2009: altitude: 7-1100 m. Kazakhstan Devyatkov, 2019b: altitude: 384-438 m. Devyatkov, 2020d: altitude: 300-1654 m. Latvia Kolcsar et al., 2021b: altitude: 3-12 m. Poland Kolcsar et al., 2021b: altitude: 103-105 m. Wiedenska and Syratt, 2023: altitude: 420-650 m. Russia Kolcsar et al., 2021b: altitude: 25-777 m. Devyatkov and Dubatolov, 2025: altitude: 70-100 m. Serbia Kolcsar et al., 2021b: altitude: 1556 m. Slovakia Stary and Reusch, 2009: altitude: 700-1665 m. Switzerland Stary and Reusch, 2009: altitude: 1500 m. Tajikistan Devyatkov, 2020c: altitude: 2440 m.