Details for Idioptera submarmorata

Name: Idioptera submarmorata (Verrall, 1887) Publication: Entomologist's Mon. Mag. 23: 264 (as Ephelia) Status: Recognized taxon, Synonym(s): bistriata (Edwards, 1938); eatoni (Edwards, 1938); oligosticta (Edwards, 1938); pentasticta (Edwards, 1938); suffumata (Edwards, 1938). Classification: Family Limoniidae Subfamily Limnophilinae Revision: Podenas, 2023 (as Eloeophila). Descriptive note: Type-material: Pont, 1995. Synonymy of Eloeophila with Idioptera after Santos and Ribeiro, 2024. Keys: Stary, 2009b (European Idioptera); Peeters en Oosterbroek, 2014c (craneflies Netherlands, Belgium and Luxembourg) (in Dutch); Boardman, 2016 (craneflies Shropshire); Stubbs and Kramer, 2016g; Stubbs, 2021 (British craneflies) (all as Eloeophila) habitus male habitus male habitus male habitus male habitus female plus 18 more images of habitus, habitus, body part(s), body part(s), hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, wing, wing, wing, wing, habitat, habitat, habitat, habitat and habitat. Distribution: Austria, Bulgaria, Czech Rep., Denmark, Finland, France, Germany, Great Britain, Hungary, Ireland, Italy (incl. Sicily), Lithuania, Netherlands, Norway, Poland, Romania, Serbia, Slovakia, Slovenia, Sweden, Switzerland, Turkey (European part: Kirklareli), Ukraine; Russia: RUC; Georgia, Turkey (Asiatic part: see citations for provinces), Iran. Note: see the manual for abbreviations: present-day Russia. Region(s): Westpalaearctic General note: Added: Slovenia (Lackschewitz, 1940b), Lithuania (Podenas, 1995). For Italy check Stary and Oosterbroek, 1996, and citations below. For Finland check Salmela, 2012b, 2012c, and citations below. For Norway check Olsen et al., 2018, and citations below. For Great Britain and Ireland check Stubbs, 2021, and citations below. Map: Ashe et al., 2005a (Ireland); Podenas et al., 2006 (Switzerland); Boardman, 2007 (Shropshire) Figure references: Habitus: Lundstrom and Fritz, 2015 (male); Podenas, 2023 (holotype male eatoni) Head: Podenas, 2023 (antenna) Wing: Podenas et al., 2006; Boardman, 2007; Podenas, 2023 Hypopygium: Podenas et al., 2006; Kramer and Morris, 2022b; Podenas, 2023 Other figures: Podenas, 2023 (thorax) Immature stages Larva: Podeniene, 2003a Miscellaneous Lundstrom and Fritz, 2015 (habitat) Citations on biology (mainly from 2000 onward): Bulgaria Habitat. Hubenov, 2025b: suppl: 3 (annotated list Diptera high Bulgarian mts, local distr, habitats and zoogeography). Country not relevant Habitat. Carles-Tolra, 2015: 119 (often found at the entrance of a cave) (in Spanish). Biology. Olsen et al., 2018: 143 (found in shaded woodland and dingle woodlands, often near springs and springbrooks, calcareous seepages in fens and carr, wooded streams (Boardman 2007, 2016, Reusch 2006, Salmela et al. 2007, Stubbs & Kramer 2016a), but also in marshy grounds (Kramer 2011, Ujvarosi 2005); larvae can be found in pools (Leskova 2009) and along margins of running water (Mendl 1978); the period of flight is may-august; see table 1 for habitat information of the Norwegian locality(ties)). Czech Rep. Habitat. Stary and Vonicka, 2018: 49 (on Limoniidae and Pediciidae of two localities in northern Bohemia, with list of localities, altitudes and habitats). Europe Habitat. Podenas, 2023: 182-183 (most associated with woodland seepages, especially those that are calcareous, can also feature in fen carr and various other types of wet woodland and is sometimes found along small woodland streams; there would appear to be a strong association with peat and other highly organic substrates (citing Stubbs, 2021); brooks, small rivers, swamps, moist soil (citing Ujvarosi, 2005); shaded woodland and dingle woodlands (citing Boardman, 2007)) (as Eloeophila). Finland Habitat. Salmela et al., 2007b: 39 (all 5 recent records are from springs and springbrooks). Habitat. Penttinen et al., 2010: 485 (red list status incl. criteria, habitats, causes of threat, threat factors). Habitat. Salmela et al., 2019: 522 (red list status, habitat types, threat factors) (as Eloeophila).. France Habitat. Quindroit and Lemoine, 2021b: 6 (loc(s) Nord (59); see paper for habitat description) (in French). Habitat. Quindroit and Lemoine, 2022a: 29 (loc(s) Aisne (02) and Somme (80) with notes on habitats and distr) (in French). Habitat. Quindroit, 2022a: 19, 27 (annotated list with information on abundancy and habitat of the Tipuloidea of the Pays de la Loire regoin, distr) (in French). Germany Habitat. Reusch, 2006: 134 (Kalkquellmoor [carbonate fen complex]). Great Britain Habitat. Godfrey, 2000a: 20, 22. Habitat. Godfrey, 2001a: 21, 23. Habitat. Boyce, 2002: 13 (review of seepage invertebrates in England, found at woodland seepages; see paper for details). Habitat. Boyce, 2002: 17 (review of seepage invertebrates in England, found at calcareous seepages; see paper for details). Habitat. Boardman, 2007: 62 (associated with seepages; found in the same habitats as E. maculata and E. mundata, shaded woodland and dingle woodlands). Habitat. Cranston and Drake, 2010: 175 (Eloeophila larvae can be found in marginal situations along flowing waters [no species mentioned]). Habitat. Drake, 2011a: 24 (among the Diptera from a wet Alder [Alnus] wood in Devon). Habitat. Kramer, 2011j: 18 (a common species of marshy ground). Habitat. Brighton, 2017c: 27, 36 (overall regional checklist Lancashire and Cheshire (VC58, 59 and 60) with notes on habitats and/or regional occurrence and/or status; see paper for details). Habitat. Kramer and Morris, 2022b: 6 (info on the species, on the distr and map of loc(s) Leicestershire and Rutland (VC55); associated with marshy locations). Lithuania Habitat. Podeniene, 2003a: 1-295 (larva, descr, figs, habitat) (in Lithuanian). Poland Habitat. Chaniecka and Wiedenska, 2006: 147 (occurrence in spring-, springbrook-, and/or bogspring plant communities) (in Polish). Habitat. Wiedenska, 2014: 12 (on the occurence of Limoniidae in six selected plant communities in the Kamienica Valley, Gorce National Park, Western Carpathians; data partly based on Chaniecka and Wiedenska, 2006) (in Polish, habitat tables also in English). Phenology. Wiedenska, 2015a: 52 (on the flight activity of Limoniidae in the Kamienica Valley, Gorce National Park, Western Carpathians) (in Polish, tables also in English). Habitat. Wiedenska, 2017a: 13 (faunistic and habitat data of species from the Gorce NP) (in Polish). Romania Habitat. Ujvarosi, 2005a: 245 (brook, small river, swamp, moist soil, citing various sources). Slovakia Habitat. Leskova, 2009: 116 ([larvae] occur in pools with dominance of psammal). Sweden Habitat. Lindstrom and Fritz, 2015: 14, 26, 39, 41, 44, 47 (on distribution, ecology, habitat preference and habitat conservation in a method study and species inventory in five fens in Hallands County) (in Schwedish). on characters and taxonomy (mainly from 2000 onward): Country not relevant Key. Stary, 2009b: 427, 432 (key, comparison). Characters. Podeniene and Gelhaus, 2010: 188-189 (comparison). Characters. Olsen et al., 2018: 143 (within Eloeophila recognized by the wing pattern: apart from the large spots along the front margin of the wing, there is a limited number of small spots on the longitudinal veins, mainly restricted to the top half of the wing; fig). Phylogeny. Santos and Ribeiro, 2024: 4-14 (phylogentic analysis of the Idioptera-Eloeophila complex, transferred to Idioptera with Eloeophila as synonym of Idioptera). Europe Description. Podenas, 2023: 180-183 (descr, figs) (as Eloeophila). France Characters. Quindroit in litt., 2025: (in Mayenne (53), I received some submarmorata of the form pentasticta Edwards 1938 mixed with submarmorata of the classic form with no intermediate specimens; this form gave me a lot of identification trouble). Great Britain Characters. Kramer and Morris, 2022b: 6 (diagnosis, figs). Lithuania Immatures. Podeniene, 2003a: 1-295 (larva, descr, figs, habitat) (in Lithuanian). on distribution (mainly from 2000 onward): Austria Distribution. Reusch and Heiss, 2012: 314-322 (loc(s) Nat. Park Gesause, see PDF for months and altitudes). Bulgaria Distribution. Hubenov, 2018a: 15 (on the Diptera of the Vitosha Mt; see paper for details on vegetation belts and local distr). Distribution. Hubenov, 2021a: 30 (checklist with summary of distr in Bulgaria, references). Distribution. Hubenov, 2025b: suppl: 3 (annotated list Diptera high Bulgarian mts, local distr, habitats and zoogeography). Czech Rep. Distribution. Stary et al., 2005a: 26 (loc(s)). Distribution. Stary, 2009i: webpage (checklist). Distribution. Stary and Vonicka, 2018: 49 (on Limoniidae and Pediciidae of two localities in northern Bohemia, with list of localities, altitudes and habitats). Denmark Distribution. Petersen and Jong, 2001b: 149 (checklist). Finland Distribution. Salmela et al., 2007b: 37-39, 45 (a red-listed species known from 6 sites, 5 of them collected between 2001 and 2006, loc(s), distr). Distribution. Salmela, 2012b: 237 (annotated list of Finnish crane flies). Distribution. Salmela and Petrasiunas, 2014: 27 (checklist Finnish Tipulomorpha). Distribution. Salmela et al., 2019: 522 (red list status, habitat types, threat factors) (as Eloeophila).. France Distribution. Quindroit and Lemoine, 2021b: 6 (loc(s) Nord (59); see paper for habitat description) (in French). Distribution. Quindroit and Lemoine, 2022a: 29 (loc(s) Aisne (02) and Somme (80) with notes on habitats and distr) (in French). Distribution. Quindroit and Lemoine, 2022a: 37 (table specifying the presence in the five departments of the Hauts-de-France region). Distribution. Quindroit, 2022a: 19, 27 (annotated list with information on abundancy and habitat of the Tipuloidea of the Pays de la Loire regoin, distr) (in French). Distribution. Podenas, 2023: 183 (loc(s) Haute-Garonne (31) and Pyrenees-Orientales (66)) (as Eloeophila). Distribution. Quindroit and Racine, 2025: 359 (table specifying the presence in the five departments of the Pays de la Loire region). Distribution. Quindroit in litt., 2025: (in Mayenne (53), I received some submarmorata of the form pentasticta Edwards 1938 mixed with submarmorata of the classic form with no intermediate specimens; this form gave me a lot of identification trouble). Germany Distribution. Schacht, 1999: 134 (checklist Bayern). Distribution. Reusch and Bellstedt, 2000: 260 (loc(s) Thuringen). Distribution. Reusch and Oosterbroek, 2000: 155 (checklist German Bundeslander). Distribution. Reusch and Bellstedt, 2001a: 268 (red list status Thuringen). Distribution. Reusch and Bellstedt, 2001b: 72 (checklist Thuringen). Distribution. Schacht, 2005: 5 (checklist Bayern). Distribution. Reusch, 2006: 134 (loc(s) Bayern). Distribution. Schacht, 2010: 16 (checklist Diptera Bayern). Distribution. Dunk and Kraus, 2014: 169 (loc(s) Bayern). Distribution. Stuke, 2019: 142 (checklist Niedersachsen and Bremen with reference to original sources). Great Britain Distribution. Godfrey, 2000a: 20, 22 (see paper for details of sites surveyed in Shropshire and into Wales). Distribution. Godfrey, 2001a: 21, 23 (recorded from sites in Shropshire and just over the country border in Montgmeryshire; see paper for details). Distribution. Boardman, 2007: 62 (loc(s), map). Distribution. Skidmore, 2009: 128 (review of occurrence on the Western Isles of Scotland, distr). Distribution. Drake, 2011a: 24 (loc(s) Devon). Distribution. Kramer, 2011j: 18 (loc(s) Leicestershire and Rutland (VC55)). Distribution. Kramer, 2013i: 8 (checklist of species from Leigh Woods NNR, Bristol district). Distribution. Hewitt, 2014: (provisional checklist Cumbrian Diptera). Distribution. Brighton, 2017c: 27, 36 (overall regional checklist Lancashire and Cheshire (VC58, 59 and 60) with notes on habitats and/or regional occurrence and/or status; see paper for details). Distribution. Kramer and Morris, 2022b: 6 (info on the species, on the distr and map of loc(s) Leicestershire and Rutland (VC55); associated with marshy locations). Hungary First record. Kolcsar et al., 2021b: 95 (loc(s) Baranya and Pest). Ireland Distribution. Ashe et al., 2005a: 317, 323 (review Irish records). Distribution. Ashe et al., 2008: 13 (additional Irish record(s), distr). Distribution. Chandler et al., 2008: 11 (checklist). Lithuania Distribution. Pakalniskis et al., 2006: 19 (checklist). Distribution. Podenas, 2023: 183 (loc(s)) (as Eloeophila). Netherlands First record. Theowald, 1971b: 225, 235 (XXX). Distribution. Jong and Oosterbroek, 2002b: 31 (checklist). Norway First record. Olsen et al., 2018: 143, 158-159 (loc(s) VE and TEY; distr; annotated checklist Nordic countries and for Norway according to the Strand regions). Poland Distribution. Chaniecka and Wiedenska, 2006: 147 (loc(s)). Distribution. Wiedenska, 2007a: 75 (checklist). Distribution. Wiedenska, 2010: 132 (loc(s) Parku Krajobrazowego Wzniesien Lodzkich, Lodz). Distribution. Wiedenska, 2014: 12 (on the occurence of Limoniidae in six selected plant communities in the Kamienica Valley, Gorce National Park, Western Carpathians; data partly based on Chaniecka and Wiedenska, 2006) (in Polish, habitat tables also in English). Distribution. Wiedenska, 2017a: 13 (faunistic and habitat data of species from the Gorce NP) (in Polish). Distribution. Wiedenska and Syratt, 2023: 131 (loc(s) Bieszczady Mountains, distr and remarks). Romania Distribution. Ujvarosi, 2007: 228-231 (checklist Limoniidae). Russia First record. Kolcsar et al., 2021b: 95-96 (loc(s) RUC: Moskovskaya oblast). Serbia First record. Kolcsar et al., 2021b: 96 (loc(s)). Slovakia Distribution. Leskova, 2009: 114 (frequency, loc(s)). Distribution. Stary, 2009d: 24 (loc(s) Polana area). Distribution. Stary, 2009i: webpage (checklist). Slovenia Distribution. Oosterbroek and Simova-Tosic, 2004: 445 (review literature). Sweden Distribution. Lindstrom and Fritz, 2015: 14, 26, 39, 41, 44, 47 (on distribution, ecology, habitat preference and habitat conservation in a method study and species inventory in five fens in Hallands County) (in Schwedish). Switzerland Distribution. Podenas et al., 2006: 137, 241 (map, vertical distr per thermic level). Distribution. Dufour and Merz, 2012: 354 (annotated checklist canton Geneva). Distribution. Podenas, 2023: 183 (loc(s) various cantons) (as Eloeophila). Turkey First record. Koc et al., 2016: 8 (loc(s) European and Asiatic part, Marmara region [the wing figure depicts I. maculata]). Distribution. Akpinar and Ozgul, 2023: 55 (loc(s) Aksaray). Distribution. Provinces Asiatic part, 2025: (known from Bilecik, Bursa, Canakkale). on flight period (mainly from 2000 onward): Czech Rep. Stary et al., 2005a: month(s): 5-6. Stary and Vonicka, 2018: month(s): 5-7. France Quindroit and Lemoine, 2021b: month(s): 5. Quindroit and Lemoine, 2022a: month(s): 5. Quindroit, 2022a: month(s): 6. Podenas, 2023: month(s): 6. Germany Reusch and Bellstedt, 2000: month(s): 6. Great Britain Godfrey, 2000a: month(s): 6. Godfrey, 2001a: month(s): 5. Kramer, 2011j: month(s): 5. Kramer and Morris, 2022b: month(s): 4-8. Hungary Kolcsar et al., 2021b: month(s): 5. Ireland Ashe et al., 2005a: month(s): 5, 7. Ashe et al., 2008: month(s): 6. Lithuania Podenas, 2023: month(s): 6-7. Norway Olsen et al., 2018: month(s): 5-6. Poland Wiedenska, 2014: month(s): 7. Wiedenska, 2015a: month(s): 6-8. Wiedenska, 2017a: month(s): 5-8. Wiedenska and Syratt, 2023: month(s): 8. Russia Kolcsar et al., 2021b: month(s): 5-6. Serbia Kolcsar et al., 2021b: month(s): 6. Slovakia Stary, 2009d: month(s): 5-7. Switzerland Podenas et al., 2006: month(s): 5-8. Podenas, 2023: month(s): 6-8. Turkey Koc et al., 2016: month(s): 6-8. on altitude (mainly from 2000 onward): Bulgaria Hubenov, 2025b: altitude: 700-1300 m. France Podenas, 2023: altitude: 1450 m. Germany Reusch and Bellstedt, 2000: altitude: 730 m. Poland Wiedenska, 2017a: altitude: 720-1210 m. Wiedenska and Syratt, 2023: altitude: 700 m. Russia Kolcsar et al., 2021b: altitude: 220 m. Serbia Kolcsar et al., 2021b: altitude: 1600 m. Slovakia Stary, 2009d: altitude: 630-780 m. Switzerland Podenas, 2023: altitude: 540-970 m. Turkey Koc et al., 2016: altitude: 18-1185 m.