Details for Idioptera maculata

Name: Idioptera maculata (Meigen, 1804) Publication: Klass. Beschr. 1: 61 (as Limonia) Status: Recognized taxon, Synonym(s): marmorata (Meigen, 1818); decora (Haliday, 1833); ?guttata (Macquart, 1834); aegle (Haliday, 1837) (nomen nudum); spoliata (Loew, 1871); aegle Edwards, 1938; melanoptera (Lackschewitz, 1940). Classification: Family Limoniidae Subfamily Limnophilinae Revision: Podenas, 2023 (as Eloeophila). Descriptive note: Synonymy: Ashe et al., 2005a; Podenas, 2023. Synonymy of Eloeophila with Idioptera after Santos and Ribeiro, 2024. Keys: Stary, 2009b (European Idioptera); Peeters en Oosterbroek, 2014c (craneflies Netherlands, Belgium and Luxembourg) (in Dutch); Stubbs and Kramer, 2016g; Stubbs, 2021 (British craneflies) (all as Eloeophila) habitus male habitus male habitus male habitus male habitus male plus 32 more images of habitus, habitus, habitus, body part(s), body part(s), body part(s), body part(s), hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, ovipositor, wing, wing, wing, wing, wing, wing, wing, wing, habitat, habitat, habitat, habitat, habitat, habitat, habitat and habitat. Distribution: Austria, Belgium, Bosnia-Herzegovina, Bulgaria, Croatia, Czech Rep., Denmark, Estonia, Finland, France (incl. Corsica), Germany, Great Britain, Greece (Kriti [Crete]), Hungary, Ireland, Italy, Latvia, Lithuania, Luxembourg, Montenegro, Netherlands, North Macedonia, Norway, Poland, Romania, Serbia, Slovakia, Slovenia, Spain, Sweden, Switzerland, Turkey (European part: Istanbul, Kirklareli), Ukraine; Russia: RUN, RUW, RUC, North Caucasus; Georgia, Azerbaijan, Turkey (Asiatic part: see citations for provinces), Iran;; Kazakhstan (northwest, east). Note: see the manual for abbreviations: present-day Russia. Region(s): Westpalaearctic;; Eastpalaearctic General note: For Italy check Stary and Oosterbroek, 1996, and citations below. For Finland check Salmela, 2012b, 2012c, and citations below. For Norway check Olsen et al., 2018, and citations below. For Spain check Eiroa and Carles-Tolra, 2019, and citations below. For Great Britain see Stubbs, 2021, and citations below. To be confirmed for Russia: RUE (Middle Volga region) (in Chuzhekova, 2015, based on larvae only). Removed, as found in earlier CCW versions and in Podenas, 2023: Armenia (excluded for Armenia by Hakobyan and Jenderedjian, 2023). Map: Ashe et al., 2005a (Ireland); Koc, 2004 (Mugla); Podenas et al., 2006 (Switzerland); Boardman, 2007 (Shropshire); Reusch and Weber, 2013 (Luxembourg) Figure references: Habitus: Quindroit, 2020a (female); Podenas, 2023 (male) Head: Podenas, 2023 Wing: Slipka and Stary, 1977b; Podenas et al., 2006; Kramer and Morris, 2022b; Podenas, 2023 Hypopygium: Podenas et al., 2006; Ribeiro, 2008; Kramer and Morris, 2022b; Podenas, 2023 Ovipositor: Podenas, 2023 Other figures: Frantsevich, 2004 (mid coxa) (as marmorata); Podenas, 2023 (thorax) Immature stages Larva: Podeniene, 2003a; Krivosheina and Krivosheina, 2011c; Podenas, 2023 Citations on biology (mainly from 2000 onward): Belgium Habitat. Tack et al., 2021: 193 (loc(s) Antwerpen, alle taxa biodiversiteit inventarisatie Bos t Ename, nat bos en stromend open water, verlanding en moeras (in Dutch), see paper for description). Bulgaria Habitat. Hubenov, 2025b: suppl: 3 (annotated list Diptera high Bulgarian mts, local distr, habitats and zoogeography). Czech Rep. Habitat. Stary and Vonicka, 2018: 49 (on Limoniidae and Pediciidae of two localities in northern Bohemia, with list of localities, altitudes and habitats). Denmark Habitat. Byriel and Rojas, 2017: 118 (among the 101 species in a study on cranefly hotspots in unmanaged and managed forests) (in Danish). Finland Habitat. Salmela, 2001a: 144, 146, 150 (aquatic, in general running water species, citing Brinkmann, 1991). Habitat. Salmela, 2004: 9 (springs, spring fens, brooks and small rivers). Biology. Salmela, 2008: 10, 27, 50 (49) (ecology or habitat: lotic; phenology). Habitat. Rasimus, 2018: 17 (on saproxylic craneflies in the Tampere region) (in Finnish). France Habitat. Breil-Moubayed and Garrigue, 2014: 68 (in: Biocenoses aquatiques et subaquatiques de la vallee fluviale de la Massane. Etude diachronique; Pyrenees-Orientales (66)). Biology. Quindroit, 2020a: 30, 45-48 (annotated list with information on abundancy and habitat of the Tipuloidea of the Pays de la Loire regoin, distr) (in French). Habitat. Quindroit, 2020b: 135 (loc(s) Nord (59) and Pas-de-Calais (62) with notes on habitats and distr) (in French). Habitat. Quindroit and Lemoine, 2022a: 29 (loc(s) Aisne (02) and Somme (80) with notes on habitats and distr) (in French). Germany Habitat. Ruckert, 2005: 47, 53 (found in a study on the association with the type of source: limnokrene, rheokrene, or helokrene in Schlesig-Holstein; see paper for details, as well as Anhang A2-A10). Biology. Wittrock, 2005: 84 (summer emergence; helokrene (subdominant), rheokrene and limnokrene). Habitat. Reusch and Schrankel, 2006: 178-179 (Quellbereichen; Larven aquatisch, in Fliessgewassern und Quellen; Larven zoophag). Habitat. Reusch, 2006: 134 (Kalkquellmoor [carbonate fen complex]). Habitat. Ruckert et al., 2007: 56 (Rheophil) (see also Ruckert et al., 2006). Habitat. Lehmann and Reusch, 2009: 122 (from around lowland springs). Habitat. Reusch and Hohmann, 2009: 129 (from spring brooks). Great Britain Habitat. Godfrey, 2000a: 20, 22. Habitat. Godfrey, 2001a: 23. Habitat. Heaver, 2006: 78 (tufa flushes). Habitat. Boardman, 2007: 62 (found in the same habitats as E. mundata and E. submarmorata, shaded woodland and dingle woodlands). Habitat. Cranston and Drake, 2010: 175 (Eloeophila larvae can be found in marginal situations along flowing waters [no species mentioned]). Habitat. Drake, 2011a: 24 (among the Diptera from a wet Alder [Alnus] wood in Devon). Habitat. Kramer, 2011j: 18 (a common species of marshy ground). Habitat. Brighton, 2017c: 27, 36 (overall regional checklist Lancashire and Cheshire (VC58, 59 and 60) with notes on habitats and/or regional occurrence and/or status; see paper for details). Habitat. Kramer and Morris, 2022b: 5 (info on the species, on the distr and map of loc(s) Leicestershire and Rutland (VC55); tend to be found in open situations throughout Britain although some VC55 records seem to come from wooded areas). Lithuania Habitat. Podeniene and Gelhaus, 2002: 68 (larvae in the riparian zone of river, in sand and silt covered with vegetation). Habitat. Podeniene, 2002: 299, 306 (larvae were found in riparian zones of small river, and in mud of low-laying marsh (Alnetalia glutinosae) on boggy lakeshore). Habitat. Podeniene, 2003a: 1-295 (larva, descr, figs, habitat) (in Lithuanian). Habitat. Podeniene, 2003b: 10, 20, 21, 29 (larval habitat: at the bottom of small rivers, sand and silt in riparian zone of small rivulets, mud in springy slopes of lakes and rivers). Luxembourg Habitat. Reusch and Weber, 2013: 271 (found in a cave, as eutrogloxeen [as by change], summary of biology). Netherlands Habitat. Oosterbroek and Dek, 2020: 16, 19 (Limoniidae and Tipulidae Nationaal Park Hollandse Duinen in 2018, Zuid-Holland). North Macedonia Habitat. Kolcsar et al., 2021b: 91 (brook). Poland Habitat. Chaniecka and Wiedenska, 2006: 147 (occurrence in spring-, springbrook-, and/or bogspring plant communities) (in Polish). Habitat. Wiedenska, 2014: 12 (on the occurence of Limoniidae in six selected plant communities in the Kamienica Valley, Gorce National Park, Western Carpathians, found only in bog spring with Caltha-Chaerophyllum) (in Polish, habitat tables also in English). Phenology. Wiedenska, 2015a: 52 (on the flight activity of Limoniidae in the Kamienica Valley, Gorce National Park, Western Carpathians) (in Polish, tables also in English). Habitat. Wiedenska, 2017a: 12 (faunistic and habitat data of species from the Gorce NP) (in Polish). Romania Habitat. Parvu, 2003: 233. Habitat. Ujvarosi, 2005a: 244 (brook, small river, swamp, moist soil, citing various sources). Habitat. Ujvarosi and Poti, 2006: 256, 260 (habitat in Dupa Lunca marshland: predominantly shrubs along rivers and streams). Habitat. Ujvarosi et al., 2011b: 107 (loc(s) Dupa Lanca marshy area near Voslobeni) (in Hungarian, species list with habitat information in English). Habitat. Kolcsar et al., 2013: 71 (loc(s) Cluj-Napoca and surroundings; list of habitats with altitudes). Slovakia Habitat. Leskova, 2009: 116 ([larvae] occur in pools with dominance of psammal). Sweden Habitat. Lindstrom and Fritz, 2015: 44, 47 (distributin and ecology in five fens in Hallands County) (in Schwedish). Turkey Biology. Koc, 2004: 21, 86, 116-118, 150 (habitat; phenology) (in Turkish). Westpalaearctic Biology. Podenas, 2023: 105 (larvae develop in sandy bottoms of small streams and springs; adults are typically found in lowlands by shaded ditches and woodland streams, under carr and at seepages, even in open situations, in the uplands, it is largely a species of valleys and shaded ravines, brooks, small rivers, swamps, moist soil, citing Ujvarosi, 2005and Stubbs, 2021); adults are attracted to light) (as Eloeophila). on characters and taxonomy (mainly from 2000 onward): Country not relevant Characters. Frantsevich, 2004: 133-134 (structure of coxa, figs) (as marmorata). Synonymy. Ashe et al., 2005a: 313, 317, 321-322, 335 (synonymy). Phylogeny. Ahonen, 2007: 1-28 (included in phylogenetic analysis of Tipuloidea subfamilies relationships based on molecular data). Phylogeny. Ribeiro, 2008: 627-694 (included in phylogenetic analysis of Limnophilinae and Tipulomorpha, figs). Key. Stary, 2009b: 426-429, 432-434 (characters, key, comparison). Phylogeny. Santos and Ribeiro, 2024: 4-14 (phylogentic analysis of the Idioptera-Eloeophila complex, transferred to Idioptera with Eloeophila as synonym of Idioptera). Great Britain Characters. Kramer and Morris, 2022b: 5 (diagnosis, figs). Lithuania Immatures. Podeniene, 2003a: 1-295 (larva, descr, figs, habitat) (in Lithuanian). Westpalaearctic Description. Podenas, 2023: 102-107, 212 (descr, remark on wrong identifications (see page 212), figs) (as Eloeophila). on distribution (mainly from 2000 onward): Armenia Misinterpretation. Obona et al., 2016: 138 (checklist but excluded for Armenia by Hakobyan and Jenderedjian, 2023). Austria Distribution. Reusch and Heiss, 2012: 314-322 (loc(s) Nat. Park Gesause, see PDF for months and altitudes). Distribution. Vogtenhuber and Kofler, 2017: 138 (loc(s) Osttirol). Azerbaijan Distribution. Snegovaya, 2021: 1145-1146 (checklist with details on distr in Azerbaijan). Belgium Distribution. Peeters in litt., 2016: (among Malaisetrap material from Jardin Massart, Brussels). Distribution. Tack et al., 2021: 193 (loc(s) Antwerpen, alle taxa biodiversiteit inventarisatie Bos t Ename, nat bos en stromend open water, verlanding en moeras (in Dutch), see paper for description). Distribution. Dek and Peeters, 2023a: 136 (loc(s) Botanic Garden Jean Massart, Brussels-Capital Region, rare in Belgium, in the eastern and southern part of the country). Bosnia-Herzegowina Distribution. Oosterbroek and Simova-Tosic, 2004: 445 (review literature). Bulgaria Distribution. Hubenov, 2015: 220 (annotated list Diptera Pirin Mt, loc(s), faunistics and zoogeography). Distribution. Hubenov, 2016: 47 (annotated checklist Diptera of the Rila Mountains based on data from the literature). Distribution. Hubenov, 2017: 67 (on vertical distribution in the Pirin and Rila Mts; see paper for details on forests and subalpine-alpine zones, distr). Distribution. Hubenov, 2018a: 15 (on the Diptera of the Vitosha Mt; see paper for details on vegetation belts and local distr). Distribution. Hubenov, 2021a: 30 (checklist with summary of distr in Bulgaria, references). Distribution. Hubenov, 2025b: suppl: 3 (annotated list Diptera high Bulgarian mts, local distr, habitats and zoogeography). Channel Is Distribution. Stubbs, 2022b: 5 (on a collection of craneflies from Jersey, collected between 1985 and 2002 by Tony Warne). Croatia Distribution. Oosterbroek and Simova-Tosic, 2004: 445 (review literature). Distribution. Kolcsar et al., 2015a: 30 (loc(s), distr). Czech Rep. Distribution. Stary et al., 2005a: 26 (loc(s)). Distribution. Stary, 2009i: webpage (checklist). Distribution. Stary and Vonicka, 2018: 49 (on Limoniidae and Pediciidae of two localities in northern Bohemia, with list of localities, altitudes and habitats). Denmark Distribution. Petersen and Jong, 2001b: 149 (checklist). Distribution. Ribeiro, 2008: 691 (loc(s)). Estonia First record. Salmela in litt., 2003: (new for Estonia). Finland Distribution. Salmela, 2001a: 144, 146, 150 (loc(s)). Distribution. Salmela, 2004: 9 (loc(s)). Distribution. Salmela et al., 2007b: 45 (loc(s)). Distribution. Salmela, 2008: 10, 50 (49) (loc(s), known from all Finnish ecoregions). Distribution. Salmela, 2010a: 12 (survey fauna Tahmela spring in city of Tampere) (in Finnish). Distribution. Salmela, 2012b: 237 (annotated list of Finnish crane flies). Distribution. Salmela and Petrasiunas, 2014: 27 (checklist Finnish Tipulomorpha). Distribution. Salmela et al., 2015: 101 (list of localities in extensive report on malaise-trapping of insects in conservation areas in Lapland in 2012–2014) (in Finnish). Distribution. Rasimus, 2018: 17 (on saproxylic craneflies in the Tampere region) (in Finnish). France Distribution. Breil-Moubayed and Garrigue, 2014: 68 (in: Biocenoses aquatiques et subaquatiques de la vallee fluviale de la Massane. Etude diachronique; Pyrenees-Orientales (66)). Distribution. Quindroit, 2020a: 30, 45-48 (annotated list with information on abundancy and habitat of the Tipuloidea of the Pays de la Loire regoin, distr) (in French). Distribution. Quindroit, 2020b: 135 (loc(s) Nord (59) and Pas-de-Calais (62) with notes on habitats and distr) (in French). Distribution. Quindroit and Lemoine, 2022a: 29 (loc(s) Aisne (02) and Somme (80) with notes on habitats and distr) (in French). Distribution. Quindroit and Lemoine, 2022a: 37 (table specifying the presence in the five departments of the Hauts-de-France region). Distribution. Podenas, 2023: 105 (loc(s) Alpes-Maritimes (06), Pyrenees-Orientales (66)) (as Eloeophila). Distribution. Quindroit and Racine, 2025: 359 (table specifying the presence in the five departments of the Pays de la Loire region). Germany Distribution. Schacht, 1999: 134 (checklist Bayern). Distribution. Reusch and Oosterbroek, 2000: 155 (checklist German Bundeslander). Distribution. Reusch and Bellstedt, 2001b: 72 (checklist Thuringen). Distribution. Ruckert, 2005: 47, 53 (found in a study on the association with the type of source: limnokrene, rheokrene, or helokrene in Schlesig-Holstein; see paper for details, as well as Anhang A2-A10). Distribution. Schacht, 2005: 5 (checklist Bayern). Distribution. Wittrock, 2005: 84 (loc(s) Schleswig-Holstein). Distribution. Reusch and Schrankel, 2006: 178-179 (loc(s) Bayern, Nationalpark Berchtesgaden, distr). Distribution. Reusch, 2006: 134 (loc(s) Bayern). Distribution. Ruckert et al., 2007: 56 (loc(s) Schleswig-Holstein). Distribution. Lehmann and Reusch, 2009: 122 (loc(s) Niedersachsens). Distribution. Reusch and Hohmann, 2009: 129 (loc(s) Sachsen-Anhalt). Distribution. Hable et al., 2010b: 91 (loc(s) Bayern). Distribution. Schacht, 2010: 16 (checklist Diptera Bayern). Distribution. Stuke, 2019: 142 (checklist Niedersachsen and Bremen with reference to original sources). Distribution. Stuke et al., 2020: 283 (loc(s) Niedersachsen). Distribution. Stuke et al., 2024: 161 (loc(s) Niedersachsen). Great Britain Distribution. Godfrey, 2000a: 20, 22 (see paper for details of sites surveyed in Shropshire and into Wales). Distribution. Godfrey, 2001a: 23 (recorded from sites in Shropshire and just over the country border in Montgmeryshire; see paper for details). Distribution. Hewitt et al., 2005: 38 (loc(s) Cumbria) (as Limnophila). Distribution. Heaver, 2006: 78 (loc(s) Yorkshire). Distribution. Boardman, 2007: 62 (loc(s), map). Distribution. Skidmore, 2009: 128 (review of occurrence on the Western Isles of Scotland, distr). Distribution. Drake, 2011a: 24 (loc(s) Devon). Distribution. Kramer, 2011j: 18 (loc(s) Leicestershire and Rutland (VC55)). Distribution. Hewitt, 2014: (provisional checklist Cumbrian Diptera). Distribution. Brighton, 2017c: 27, 36 (overall regional checklist Lancashire and Cheshire (VC58, 59 and 60) with notes on habitats and/or regional occurrence and/or status; see paper for details). Distribution. Kramer and Morris, 2022b: 5 (info on the species, on the distr and map of loc(s) Leicestershire and Rutland (VC55); tend to be found in open situations throughout Britain although some VC55 records seem to come from wooded areas). Iran First record. Podenas, 2023: 105 (loc(s) Mazandaran, Teheran) (as Eloeophila). Ireland Distribution. Ashe et al., 2005a: 313, 317, 321-322, 335 (review Irish records). Distribution. Ashe et al., 2008: 13 (additional Irish record(s), distr). Distribution. Chandler et al., 2008: 11 (checklist). Italy Distribution. Podenas and Podeniene, 2008: 347, 351-352 (loc(s), annotated checklist South Tyrol [Prov. Bolzano]). Kazakhstan Distribution. Devyatkov, 2020d: 269-270 (loc(s) East Kazakhstan, distr) (in Russian). Latvia Distribution. Stary, 2004f: (Fauna Europaea: added Latvia). Lithuania Distribution. Podeniene and Gelhaus, 2002: 68 (loc(s)). Distribution. Podeniene, 2002: 299, 306 (loc(s)). Distribution. Pakalniskis et al., 2006: 19 (checklist). Distribution. Podenas, 2023: 105 (loc(s)) (as Eloeophila). Luxembourg First record. Reusch and Weber, 2013: 271 (loc(s) canton Wiltz, map, distr) (as Eloeophila). Montenegro Distribution. Oosterbroek and Simova-Tosic, 2004: 445 (review literature). Netherlands Distribution. Jong and Oosterbroek, 2002b: 31 (checklist). Distribution. Oosterbroek and Dek, 2020: 16, 19 (Limoniidae and Tipulidae Nationaal Park Hollandse Duinen in 2018, Zuid-Holland). Distribution. Podenas, 2023: 105 (loc(s) Limburg) (as Eloeophila). Distribution. Hoven, 2024: 13 (loc(s) Limburg). North Macedonia First record. Kolcsar et al., 2021b: 91 (loc(s)). Norway Distribution. Olsen et al., 2018: 158-159 (annotated checklist Nordic countries and for Norway according to the Strand regions). Poland Distribution. Chaniecka and Wiedenska, 2006: 147 (loc(s)). Distribution. Wiedenska, 2007a: 75 (checklist). Distribution. Wiedenska, 2010: 132 (loc(s) Parku Krajobrazowego Wzniesien Lodzkich, Lodz). Distribution. Wiedenska, 2014: 12 (on the occurence of Limoniidae in six selected plant communities in the Kamienica Valley, Gorce National Park, Western Carpathians, found only in bog spring with Caltha-Chaerophyllum) (in Polish, habitat tables also in English). Distribution. Wiedenska, 2017a: 12 (faunistic and habitat data of species from the Gorce NP) (in Polish). Distribution. Wiedenska and Syratt, 2023: 130 (loc(s) Bieszczady Mountains, distr and remarks). Romania Distribution. Parvu, 2003: 233 (loc(s)). Distribution. Parvu, 2004: 184 (loc(s)). Distribution. Parvu, 2006: 260 (loc(s)). Distribution. Ujvarosi and Poti, 2006: 256, 260 (loc(s) eastern Carpathians). Distribution. Ujvarosi, 2007: 228-231 (checklist Limoniidae). Distribution. Ujvarosi et al., 2011b: 107 (loc(s) Dupa Lanca marshy area near Voslobeni) (in Hungarian, species list with habitat information in English). Distribution. Keresztes, 2012: 30 (loc(s) Rimetea area [Alba County]). Distribution. Kolcsar et al., 2013: 71 (loc(s) Cluj-Napoca and surroundings; list of habitats with altitudes). Russia Distribution. Polevoi, 2006: 96 (loc(s) RUN: Kareliya) (in Russian) (as marmorata). Distribution not confirmed. Chuzhekova, 2015: 211, 240 (on macrozoobenthos communities in spring brooks of Middle Volga) (PhD thesis, in Russian, 242 pages; for English summary see Chuzhekova, 2016, 37 pages). Distribution. Lantsov and Pilipenko, 2018: 79 (first information about limoniids crane flies of the North Caucasus: Krasnodarskiy kray: the Utrish reserve and adjacent areas) (in Russian). Distribution. Pilipenko et al., 2020: 357 (loc(s) RUC: Mordoviya Rep., distr). Serbia Distribution. Oosterbroek and Simova-Tosic, 2004: 445 (review literature). Distribution. Gavryushin in litt., 2015b: (loc(s) Serbia). Slovakia Distribution. Leskova, 2009: 114 (frequency, loc(s)). Distribution. Stary, 2009d: 24 (loc(s) Polana area). Distribution. Stary, 2009i: webpage (checklist). Slovenia Distribution. Oosterbroek and Simova-Tosic, 2004: 445 (review literature). Spain Distribution. Eiroa and Baez, 2002a: 54 (checklist). Distribution. Mederos and Eiroa, 2015: 266 (loc(s) Lerida). Distribution. Mederos and Eiroa, 2018: 27 (loc(s) Lerida, distr). Distribution. Eiroa and Carles-Tolra, 2019: 131 (loc(s) Gerona, list of Spanish provinces, distr) (in Spanish). Distribution. Hancock, 2020: 97 (loc(s) Jaen, Cazorla National Park). Distribution. Podenas, 2023: 105 (loc(s) Lerida) (as Eloeophila). Sweden Distribution. Fritz and Lindstrom, 2013: 62 (loc(s) Hallands County) (in Schwedish). Distribution. Lindstrom and Fritz, 2015: 44, 47 (distributin and ecology in five fens in Hallands County) (in Schwedish). Switzerland Distribution. Podenas et al., 2006: 136, 240 (map, vertical distr per thermic level). Distribution. Dufour and Merz, 2012: 354 (annotated checklist canton Geneva). Distribution. Podenas, 2023: 105 (loc(s) various cantons) (as Eloeophila). Turkey Distribution. Koc, 2004: 21, 86, 116-118, 150 (loc(s), distr, map) (in Turkish). Distribution. Koc et al., 2005b: 616 (checklist). Distribution. Ozgul et al., 2009: 63 (survey provinces SW Turkey). Distribution. Bilgin et al., 2015: 6 (loc(s) Eskisehir, distr) (in Turkish). Distribution. Koc et al., 2016: 11 (provinces Marmara region). Distribution. Provinces Asiatic part, 2025: (known from Balikesir, Bursa, Eskisehir, Mugla, Sakarya). on flight period (mainly from 2000 onward): Austria Vogtenhuber and Kofler, 2017: month(s): 7. Bulgaria Hubenov, 2015: month(s): 6. Hubenov, 2016: month(s): 6. Channel Is Stubbs, 2022b: month(s): 10. Croatia Kolcsar et al., 2015a: month(s): 8, 10. Czech Rep. Stary et al., 2005a: month(s): 4-8. Stary and Vonicka, 2018: month(s): 5-8. Denmark Ribeiro, 2008: month(s): 8. Finland Salmela, 2001a: month(s): 6. Salmela, 2008: month(s): 7. France Quindroit, 2020a: month(s): 5-6. Quindroit, 2020b: month(s): 5, 7. Quindroit and Lemoine, 2022a: month(s): 5, 7. Podenas, 2023: month(s): 6-7. Germany Reusch and Schrankel, 2006: month(s): 7-8. Stuke et al., 2024: month(s): 6. Great Britain Godfrey, 2000a: month(s): 6. Godfrey, 2001a: month(s): 5. Kramer, 2011j: month(s): 5-6, 10. Kramer and Morris, 2022b: month(s): 5-9. Iran Podenas, 2023: month(s): 7, 9. Ireland Ashe et al., 2005a: month(s): 4-9. Ashe et al., 2008: month(s): 8. Italy Podenas and Podeniene, 2008: month(s): 5-7. Kazakhstan Devyatkov, 2020d: month(s): 6-8. Lithuania Podenas, 2023: month(s): 5-8. Netherlands Oosterbroek and Dek, 2020: month(s): 5. Podenas, 2023: month(s): 6. Hoven, 2024: month(s): 5. North Macedonia Kolcsar et al., 2021b: month(s): 5-6. Poland Wiedenska, 2015a: month(s): 6-8. Wiedenska, 2017a: month(s): 8. Wiedenska and Syratt, 2023: month(s): 7-8. Romania Parvu, 2003: month(s): 7-8. Parvu, 2004: month(s): 6-7. Parvu, 2006: month(s): 6. Ujvarosi et al., 2011b: month(s): 5-6. Kolcsar et al., 2013: month(s): 5, 8. Russia Lantsov and Pilipenko, 2018: month(s): 5. Pilipenko et al., 2020: month(s): 5. Serbia Gavryushin in litt., 2015b: month(s): 7. Slovakia Stary, 2009d: month(s): 5-6, 9. Spain Mederos and Eiroa, 2018: month(s): 6. Hancock, 2020: month(s): 6. Podenas, 2023: month(s): 6. Switzerland Podenas et al., 2006: month(s): 5-9. Podenas, 2023: month(s): 5-9. Turkey Koc, 2004: month(s): 5-6. on altitude (mainly from 2000 onward): Austria Vogtenhuber and Kofler, 2017: altitude: 1920 m. Bulgaria Hubenov, 2015: altitude: 1142 m. Hubenov, 2016: altitude: 1147 m. Hubenov, 2017: altitude: 1142-1147 m. Hubenov, 2018a: altitude: 870-880 m. Hubenov, 2021a: altitude: 0-1700 m. Hubenov, 2025b: altitude: 0-1700 m. Croatia Kolcsar et al., 2015a: altitude: 665-715 m. France Podenas, 2023: altitude: 1120-1985 m. Germany Reusch and Schrankel, 2006: altitude: 1100-1250 m. Reusch and Hohmann, 2009: altitude: 375-825 m. Stuke et al., 2024: altitude: 200-270 m. Iran Podenas, 2023: altitude: 2000 m. Italy Podenas and Podeniene, 2008: altitude: 940-1220 m. Kazakhstan Devyatkov, 2020d: altitude: 353-1030 m. North Macedonia Kolcsar et al., 2021b: altitude: 755-875 m. Poland Wiedenska, 2017a: altitude: 760 m. Wiedenska and Syratt, 2023: altitude: 470-750 m. Romania Ujvarosi and Poti, 2006: altitude: 670 m. Serbia Gavryushin in litt., 2015b: altitude: 1500-1550 m. Slovakia Stary, 2009d: altitude: 630-870 m. Spain Mederos and Eiroa, 2015: altitude: 1450 m. Mederos and Eiroa, 2018: altitude: 960 m. Switzerland Podenas, 2023: altitude: 450-970 m. Turkey Koc, 2004: altitude: 700-1025 m.