Details for Cheilotrichia (Empeda) cinerascens
Name:Cheilotrichia (Empeda) cinerascens (Meigen, 1804)
Publication:Klass. Beschr. 1: 52 (as Erioptera)
Status:Recognized taxon, Synonym(s): nubila (Schummel, 1829); diluta (Zetterstedt, 1851); tenella (Walker, 1856).
Classification:Family Limoniidae
Subfamily Chioneinae
Keys:Stary and Freidberg, 2007 (Limoniidae Israel); Boardman, 2016 (craneflies Shropshire); Stubbs and Kramer, 2016i, 2016j; Stubbs, 2021 (British craneflies)
Cheilotrichia (Empeda) cinerascens : habitus - maleCheilotrichia (Empeda) cinerascens : habitus - maleCheilotrichia (Empeda) cinerascens : habitus - maleCheilotrichia (Empeda) cinerascens : habitus - maleCheilotrichia (Empeda) cinerascens : body part(s) - antenna
habitus
male
habitus
male
habitus
male
habitus
male
body part(s)
antenna
plus 21 more images of hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, wing, wing, wing, habitat, habitat, habitat, habitat, habitat and habitat.
Distribution:Andorra, Austria, Belarus, Belgium, Bulgaria, Czech Rep., Denmark (incl. Faroe Is), Estonia, Finland, France (incl. Corsica), Germany, Great Britain, Hungary, Ireland, Italy, Latvia, Lithuania, Malta, Netherlands, Norway, Poland, Portugal, Romania, Serbia, Slovakia, Slovenia, Spain (incl. Canary Is [Tenerife]), Sweden, Switzerland, Turkey (European part: Kirklareli), Ukraine; Russia: RUN, RUW, RUC, RUE, North Caucasus; Morocco; Georgia, Turkey (Asiatic part: see citations for provinces), Cyprus, Israel, Iran;; Kazakhstan (east).
Note: see the manual for abbreviations: present-day Russia.
Region(s):Westpalaearctic;; Eastpalaearctic
General note:Added: Corsica (Podenas et al., 1997). For Canary Is, incl. key, check de Jong, 1987b and citations below. For Italy check Stary and Oosterbroek, 1996, and citations below. For Finland check Salmela, 2012b, 2012c, and citations below. For Norway check Olsen et al., 2018, and citations below. For Spain check Eiroa and Carles-Tolra, 2019, and citations below. For Portugal check Oosterbroek et al., 2020, and citations below. For Great Britain and Ireland check Stubbs, 2021, and citations below. For Morocco check Kettani and Oosterbroek, 2022a, and citations below.
Map:Koc et al., 2006 (SW Turkey); Podenas et al., 2006 (Switzerland); Ashe et al., 2007b (Ireland); Boardman, 2007 (Shropshire)
Figure
references:
Habitus: Boardman, 2007
Head: Slipka and Stary, 1977b (antenna); Stary, 1987a (antenna)
Wing: Slipka and Stary, 1977b; Podenas et al., 2006
Hypopygium: Stary and Rozkosny, 1970a; Stary, 1987a; Podenas et al., 2006; Kramer and Morris, 2022a

Immature stages
Larva: Podeniene, 2003a; Krivosheina and Krivosheina, 2011c
Citationson biology (mainly from 2000 onward):
Czech Rep.
Habitat. Stary and Vonicka, 2018: 53 (on Limoniidae and Pediciidae of two localities in northern Bohemia, with list of localities, altitudes and habitats).
Denmark
Habitat. Nielsen and Nielsen, 2009: 121, 131 (emerging from a forest floor in a Beech [Fagus] stand)(in Danish).
Habitat. Byriel and Rojas, 2017: 118 (among the 101 species in a study on cranefly hotspots in unmanaged and managed forests) (in Danish).
Finland
Habitat. Salmela, 2001a: 144, 146, 150 (eurytopic species in relation to waterlevel, citing Brinkmann, 1991).
Habitat. Salmela and Autio, 2007b: 49 (eurytopic dweller of wetlands and moist biotopes).
Habitat. Salmela and Vartija, 2007: 10 (common in moist environments, citing Salmela, 2006).
Habitat. Autio and Salmela, 2010: 48 (habitats Aland Is).
Habitat. Rasimus, 2018: 17 (on saproxylic craneflies in the Tampere region) (in Finnish).
France
Biology. Quindroit, 2020a: 26, 45-48 (annotated list with information on abundancy and habitat of the Tipuloidea of Pays de la Loire: Mayenne (53), Sarthe (72), Loire-Atlantique (44), Maine-et-Loire (49) and Vendee (85)) (in French).
Habitat. Quindroit, 2020b: 134 (loc(s) Nord (59) and Pas-de-Calais (62) with notes on habitats and distr) (in French).
Habitat. Quindroit and Lemoine, 2021a: 24 (loc(s), resultats de deux campagnes de piegeage dans le Pas-de-Calais (62); see paper for details on habitat).
Habitat. Quindroit, 2021a: 36 (loc(s), resultats d une campagne de piegeage dans le Pas-de-Calais (62); see paper for details on habitat).
Germany
Habitat. Hovemeyer and Schauermann, 2003: 72 (found in a study on succession of Diptera on dead Beach [Fagus] wood near Gottingen; see paper for details).
Habitat. Reusch, 2006: 134 (Kalkquellmoor [carbonate fen complex]).
Habitat. Reusch and Hohmann, 2009: 129 (from spring brooks).
Great Britain
Habitat. Godfrey, 2000a: 21.
Habitat. Godfrey, 2001a: 21, 23.
Habitat. Boardman, 2007: 40 (encountered regularly by sweeping the lower branches of churchyard trees, holly bushes and other shrubs in woodland, in Shropshire found to be almost universal in all habitat types).
Habitat. Skidmore, 2010: 161 (reared as casuals from cow dung).
Habitat. Drake, 2011a: 24 (among the Diptera from a wet Alder [Alnus] wood in Devon).
Habitat. Kramer, 2011j: 13,025 (common woodland species).
Habitat. Brighton, 2017c: 25, 32 (overall regional checklist Lancashire and Cheshire (VC58, 59 and 60) with notes on habitats and/or regional occurrence and/or status; see paper for details).
Kazakhstan
Habitat. Devyatkov, 2017b: 212 (collected along a stream).
Latvia
Habitat. Salmela and Vartija, 2007: 10 (at a brook with relatively swift flow, sandy and stony bottom, mosses and riparian forest).
Lithuania
Habitat. Podeniene, 2003a: 1-295 (larva, descr, figs, habitat) (in Lithuanian).
Morocco
Habitat. Driauach and Belqat, 2016: 134 (loc(s) Rif Mts and Beni Snassen with list of habitats).
Norway
Habitat. Olsen and Andersen, 2021: 207 (loc(s) Innlandet, in former Hedmark, with habitats specified separately, for habitats see also Jonassen and Andersen, 2020).
Poland
Habitat. Chaniecka and Wiedenska, 2006: 148 (occurrence in spring-, springbrook-, and/or bogspring plant communities) (in Polish).
Habitat. Wiedenska, 2014: 13 (on the occurence of Limoniidae in six selected plant communities in the Kamienica Valley, Gorce National Park, Western Carpathians) (in Polish, habitat tables also in English).
Phenology. Wiedenska, 2015a: 52 (on the flight activity of Limoniidae in the Kamienica Valley, Gorce National Park, Western Carpathians) (in Polish, tables also in English).
Habitat. Wiedenska, 2017a: 16 (faunistic and habitat data of species from the Gorce NP) (in Polish).
Romania
Habitat. Ujvarosi, 2005a: 244 (brook, small river, swamp, moist soil, citing various sources).
Habitat. Kolcsar et al., 2013: 71 (loc(s) Cluj-Napoca and surroundings; list of habitats with altitudes).
Sweden
Habitat. Lindstrom and Fritz, 2015: 44, 47 (distributin and ecology in five fens in Hallands County) (in Schwedish).
on characters and taxonomy (mainly from 2000 onward):
Country not relevant
Characters. Frantsevich, 2004: 133-135 (structure of coxa).
Characters. Devyatkov, 2017b: 213-214 (comparison).
Phylogeny. Men et al., 2018a: 97 (comparison of female internal reproductive system, based on literure data).
Great Britain
Characters. Boardman, 2007: 40, 87 (characters, colourphoto).
Characters. Kramer and Morris, 2022a: 4 (diagnosis, fig(s)).
Israel
Key. Stary and Freidberg, 2007: 304, 314-315 (key).
Lithuania
Immatures. Podeniene, 2003a: 1-295 (larva, descr, figs, habitat) (in Lithuanian).
on distribution (mainly from 2000 onward):
Andorra
First record. Eiroa and Carles-Tolra, 2019: 130 (loc(s)).
Austria
Distribution. Reusch and Heiss, 2012: 314-322 (loc(s) Nat. Park Gesause, see PDF for months and altitudes).
Belgium
Distribution. Dek and Peeters, 2023a: 135 (loc(s) Botanic Garden Jean Massart, Brussels-Capital Region, common in Belgium).
Bulgaria
Distribution. Hubenov, 2021a: 32 (checklist with summary of distr in Bulgaria, references).
Cyprus
First record. Stary and Oosterbroek, 2008: 2 (loc(s), distr).
Czech Rep.
Distribution. Stary et al., 2005a: 25 (loc(s)).
Distribution. Stary, 2009i: webpage (checklist).
Distribution. Stary and Vonicka, 2018: 53 (on Limoniidae and Pediciidae of two localities in northern Bohemia, with list of localities, altitudes and habitats).
Denmark
Distribution. Petersen and de Jong, 2001b: 148 (checklist).
Distribution. Nielsen and Nielsen, 2009: 121 (loc(s)).
Estonia
Distribution. Stary, 2004f: (Fauna Europaea: added Estonia).
Finland
Distribution. Salmela, 2001a: 144, 146, 150 (loc(s)).
Distribution. Salmela, 2002: 30-32 (loc(s) lake Iidesjarvi) (in Finnish).
Distribution. Salmela and Autio, 2007b: 54 (loc(s)).
Distribution. Salmela et al., 2007b: 45 (loc(s)).
Distribution. Autio and Salmela, 2010: 48 (loc(s) Aland Is).
Distribution. Salmela, 2010a: 8, 12 (survey fauna Tahmela spring in city of Tampere) (in Finnish).
Distribution. Salmela, 2012b: 236 (annotated list of Finnish crane flies).
Distribution. Salmela and Petrasiunas, 2014: 24 (checklist Finnish Tipulomorpha).
Distribution. Salmela et al., 2015: 100 (list of localities in extensive report on malaise-trapping of insects in conservation areas in Lapland in 2012–2014) (in Finnish).
Distribution. Rasimus, 2018: 17 (on saproxylic craneflies in the Tampere region) (in Finnish).
France
Distribution. Kramer and Langlois, 2019a: 76 (record(s) Ravin de Valbois, Doubs (25)).
Distribution. Quindroit, 2020a: 26, 45-48 (annotated list with information on abundancy and habitat of the Tipuloidea of Pays de la Loire: Mayenne (53), Sarthe (72), Loire-Atlantique (44), Maine-et-Loire (49) and Vendee (85)) (in French).
Distribution. Quindroit, 2020b: 134 (loc(s) Nord (59) and Pas-de-Calais (62) with notes on habitats and distr) (in French).
Distribution. Quindroit and Lemoine, 2021a: 24 (loc(s), resultats de deux campagnes de piegeage dans le Pas-de-Calais (62); see paper for details on habitat).
Distribution. Quindroit, 2021a: 36 (loc(s), resultats d une campagne de piegeage dans le Pas-de-Calais (62); see paper for details on habitat).
Distribution. Tissot et al., 2021: 208 (the Diptera of the Reserves Naturelles Nationales du Doubs (25), results of Malaise trapping since 2009) (in French).
Germany
Distribution. Schacht, 1999: 134 (checklist Bayern).
Distribution. Reusch and Oosterbroek, 2000: 157 (checklist German Bundeslander).
Distribution. Reusch and Bellstedt, 2001b: 73 (checklist Thuringen).
Distribution. Hovemeyer and Schauermann, 2003: 72 (found in a study on succession of Diptera on dead Beach [Fagus] wood near Gottingen; see paper for details).
Distribution. Schacht, 2005: 4 (checklist Bayern).
Distribution. Reusch, 2006: 134 (loc(s)).
Distribution. Reusch and Hohmann, 2009: 129 (loc(s) Sachsen-Anhalt).
Distribution. Hable et al., 2010b: 90 (loc(s) Bayern).
Distribution. Schacht, 2010: 16 (checklist Diptera Bayern).
Distribution. Stuke, 2019: 141 (checklist Niedersachsen and Bremen with reference to original sources).
Great Britain
Distribution. Godfrey, 2000a: 21 (loc(s)).
Distribution. Godfrey, 2001a: 21, 23 (loc(s)).
Distribution. Boardman, 2007: 40, 87 (distr, map).
Distribution. Skidmore, 2009: 126 (review of occurrence on the Western Isles of Scotland, distr).
Distribution. Drake, 2011a: 24 (loc(s) Devon).
Distribution. Kramer, 2011j: 13 (loc(s) Leicestershire and Rutland (VC55)).
Distribution. Kramer, 2013i: 8 (checklist of species from Leigh Woods NNR, Bristol district).
Distribution. Hewitt, 2014: (provisional checklist Cumbrian Diptera).
Distribution. Kramer, 2014c: 4 (in table of cranefly species recorded at Shapwick Heath NNR, as given in H. Audcent, 1949 and/or collected by J. Kramer in 2013).
Distribution. Wolton et al., 2014: 26 (among Diptera associated with a hedge in Devon).
Distribution. Chandler, 2015: 89 (loc(s) Bushy Park, Middlesex).
Distribution. Brighton, 2017c: 25, 32 (overall regional checklist Lancashire and Cheshire (VC58, 59 and 60) with notes on habitats and/or regional occurrence and/or status; see paper for details).
Distribution. Kramer and Morris, 2022a: 4 (info on the species, on the distr and map of loc(s) Leicestershire and Rutland (VC55)).
Ireland
Distribution. Ashe et al., 2007b: 361, 364-365, 402 (review Irish records, map).
Distribution. Chandler et al., 2008: 9 (checklist).
Israel
First record. Stary and Freidberg, 2007: 304, 314-315 (loc(s), distr).
Distribution. Stary, 2014c: 150 (loc(s) Israel).
Italy
Distribution. Podenas and Podeniene, 2008: 347, 351 (loc(s), annotated checklist South Tyrol [Prov. Bolzano]).
Kazakhstan
First record. Devyatkov, 2017b: 212 (loc(s) East Kazakhstan, first record for Eastpalaearctic).
Distribution. Devyatkov, 2020b: 219 (loc(s) East Kazakhstan, distr) (in Russian).
Latvia
Distribution. Salmela and Vartija, 2007: 10 (loc(s)).
Lithuania
Distribution. Pakalniskis et al., 2006: 17 (checklist).
Malta
First record. Ebejer, 2015: 52 (loc(s) Malta).
Distribution. Ebejer and Gatt, 2021: 17 (checklist of the Diptera of Malta).
Morocco
First record. Driauach et al., 2013: 188 (review Limoniidae records of Morocco, loc(s)).
Distribution. Driauach and Belqat, 2016: 134 (loc(s) Rif Mts and Beni Snassen with list of habitats).
Distribution. Kettani and Oosterbroek, 2022a: 20 (annotated checklist, with review loc(s) and/or references).
Netherlands
Distribution. de Jong and Oosterbroek, 2002b: 28 (checklist).
Distribution. Stary, 2020a: 45 (Limoniidae Kaaistoep, Noord-Brabant).
Norway
Distribution. Olsen et al., 2018: 152-153 (annotated checklist Nordic countries and for Norway according to the Strand regions).
Distribution. Olsen and Andersen, 2021: 207 (loc(s) Innlandet, in former Hedmark, with habitats specified separately, for habitats see also Jonassen and Andersen, 2020).
Poland
Distribution. Chaniecka and Wiedenska, 2006: 148 (loc(s)).
Distribution. Wiedenska, 2007a: 74 (checklist).
Distribution. Wiedenska, 2010: 131 (loc(s) Parku Krajobrazowego Wzniesien Lodzkich, Lodz).
Distribution. Wiedenska, 2014: 13 (on the occurence of Limoniidae in six selected plant communities in the Kamienica Valley, Gorce National Park, Western Carpathians) (in Polish, habitat tables also in English).
Distribution. Wiedenska, 2017a: 16 (faunistic and habitat data of species from the Gorce NP) (in Polish).
Distribution. Palaczyk and Klasa, 2021: 173 (list of Diptera in Catalogue of the fauna of the Ojcow National Park).
Distribution. Wiedenska and Syratt, 2023: 136 (loc(s) Bieszczady Mountains, distr and remarks).
Portugal
First record. Stary, 2014b: 84 (loc(s) Faro).
Distribution. Oosterbroek et al., 2020: 320 (known from Faro only, distr).
Romania
Distribution. Ujvarosi, 2007: 228-231 (checklist Limoniidae).
Distribution. Kolcsar et al., 2013: 71 (loc(s) Cluj-Napoca and surroundings; list of habitats with altitudes).
Russia
Distribution. Paramonov and Pilipenko, 2016: 96 (loc(s) RUC: Tverskaya oblast) (in Russian).
Distribution. Paramonov and Korobkov, 2019: 91-92 (loc(s) RUC: Tverskaya oblast, distr) (in Russian).
Distribution. Kolcsar et al., 2021b: 23-24 (loc(s) RUE: Bashkortostan Rep.).
Serbia
Distribution. Oosterbroek and Simova-Tosic, 2004: 443 (review literature).
Slovakia
Distribution. Stary, 2009d: 27 (loc(s) Polana area).
Distribution. Stary, 2009i: webpage (checklist).
Slovenia
Distribution. Oosterbroek and Simova-Tosic, 2004: 443 (review literature).
Spain
Distribution. Eiroa and Baez, 2002a: 54 (checklist).
Distribution. Baez and Garcia, 2004: 270 (lista de especies silvestres de Canarias).
Distribution. Stary, 2014b: 84 (loc(s) Malaga).
Distribution. Mederos and Eiroa, 2015: 266 (loc(s) Lerida).
Distribution. Eiroa and Carles-Tolra, 2019: 130 (list of Spanish provinces, distr) (in Spanish).
Distribution. Keresztes et al., 2022: 192 (loc(s) Asturias and review distribution in Spain) (in Spanish).
Sweden
Distribution. Salmela, 2010b: 133 (loc(s) Malmo).
Distribution. Lindstrom and Fritz, 2015: 44, 47 (distributin and ecology in five fens in Hallands County) (in Schwedish).
Switzerland
Distribution. Podenas et al., 2006: 85, 203 (map, vertical distr per thermic level).
Distribution. Dufour and Merz, 2012: 353 (annotated checklist canton Geneva).
Turkey
First record. Koc et al., 2006: 19-21 (loc(s), distr, map).
Distribution. Ozgul et al., 2009: 63 (survey provinces SW Turkey).
Distribution. Koc et al., 2016: 10 (provinces Marmara region).
Distribution. Provinces Asiatic part, 2024: (known from Aydin, Balikesir, Bursa, Canakkale, Mugla, Sakarya).
on flight period (mainly from 2000 onward):
Andorra
Eiroa and Carles-Tolra, 2019: month(s): 4-5.
Belgium
Dek and Peeters, 2023a: month(s): 5, 9-11.
Cyprus
Stary and Oosterbroek, 2008: month(s): 10.
Czech Rep.
Stary et al., 2005a: month(s): 3-8.
Stary and Vonicka, 2018: month(s): 4-5, 7-8(9).
Finland
Salmela, 2001a: month(s): 5.
France
Quindroit, 2020a: month(s): 4, 10.
Quindroit, 2020b: month(s): 4.
Great Britain
Godfrey, 2000a: month(s): 6.
Godfrey, 2001a: month(s): 5.
Boardman, 2007: month(s): 4-11.
Kramer, 2011j: month(s): 5-6, 10.
Kramer, 2014c: month(s): 6.
Chandler, 2015: month(s): 11.
Ireland
Ashe et al., 2007b: month(s): 4-11.
Israel
Stary and Freidberg, 2007: month(s): 9-2.
Stary, 2014c: month(s): 11-12.
Italy
Podenas and Podeniene, 2008: month(s): 5-6, 8- 9.
Kazakhstan
Devyatkov, 2017b: month(s): 8-9.
Devyatkov, 2020b: month(s): 5, 9.
Latvia
Salmela and Vartija, 2007: month(s): 5.
Malta
Ebejer, 2015: month(s): 1.
Morocco
Driauach and Belqat, 2016: month(s): 4-5, 11-12.
Norway
Olsen and Andersen, 2021: month(s): 8.
Poland
Wiedenska, 2014: month(s): 5-9.
Wiedenska, 2015a: month(s): 5-8.
Wiedenska, 2017a: month(s): 6-8.
Wiedenska and Syratt, 2023: month(s): 8-9.
Portugal
Stary, 2014b: month(s): 4.
Romania
Kolcsar et al., 2013: month(s): 10.
Russia
Paramonov and Pilipenko, 2016: month(s): 8-9.
Paramonov and Korobkov, 2019: month(s): 5.
Kolcsar et al., 2021b: month(s): 8.
Slovakia
Stary, 2009d: month(s): 5-7, 9.
Spain
Stary, 2014b: month(s): 3-4.
Keresztes et al., 2022: month(s): 8.
Switzerland
Podenas et al., 2006: month(s): 4-11.
Turkey
Koc et al., 2006: month(s): 11.
on altitude (mainly from 2000 onward):
Bulgaria
Hubenov, 2021a: altitude: 230-300 m.
Cyprus
Stary and Oosterbroek, 2008: altitude: 500-700 m.
Germany
Reusch and Hohmann, 2009: altitude: 435 m.
Israel
Stary and Freidberg, 2007: altitude: 670-1600 m.
Italy
Podenas and Podeniene, 2008: altitude: 940-1630 m.
Kazakhstan
Devyatkov, 2020b: altitude: 465 m.
Morocco
Driauach et al., 2013: altitude: 680 m.
Driauach and Belqat, 2016: altitude: 13-1600 m.
Norway
Olsen and Andersen, 2021: altitude: 700 m.
Poland
Wiedenska, 2017a: altitude: 800-1010 m.
Wiedenska and Syratt, 2023: altitude: 700 m.
Portugal
Stary, 2014b: altitude: 75-500 m.
Russia
Kolcsar et al., 2021b: altitude: 607 m.
Slovakia
Stary, 2009d: altitude: 560-780 m.
Spain
Mederos and Eiroa, 2015: altitude: 1300 m.
Keresztes et al., 2022: altitude: 574 m.
Turkey
Koc et al., 2006: altitude: 235 m.

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