Details for Symplecta (Psiloconopa) stictica stictica
Name:Symplecta (Psiloconopa) stictica stictica (Meigen, 1818)
Publication:Syst. Beschr. 1: 148 (as Limnobia)
Status:Recognized taxon, Synonym(s): similis (Schummel, 1829).
Classification:Family Limoniidae
Subfamily Chioneinae
Keys:Stary and Freidberg, 2007 (Limoniidae Israel); Peeters en Oosterbroek, 2014c (craneflies Netherlands, Belgium and Luxembourg) (in Dutch); Boardman, 2016 (craneflies Shropshire); Stubbs and Kramer, 2016i; Stubbs, 2021 (British craneflies)
Symplecta (Psiloconopa) stictica stictica: habitus - maleSymplecta (Psiloconopa) stictica stictica: habitus - maleSymplecta (Psiloconopa) stictica stictica: habitus - maleSymplecta (Psiloconopa) stictica stictica: habitus - maleSymplecta (Psiloconopa) stictica stictica: habitus - female
habitus
male
habitus
male
habitus
male
habitus
male
habitus
female
plus 21 more images of habitus, habitus, habitus, body part(s), hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, wing, wing, wing, wing, habitat, habitat, habitat, habitat, habitat, habitat and habitat.
Distribution:Albania, Austria, Belgium, Bosnia-Herzegovina, Bulgaria, Croatia, Czech Rep., Denmark, Estonia, Finland, France (incl. Corsica), Germany, Great Britain, Greece (incl. Lesvos, Mykonos, Samos), Hungary, Iceland, Ireland, Italy (incl. Sicily), Latvia, Lithuania, Malta, Netherlands, North Macedonia, Norway, Poland, Portugal, Romania, Serbia, Slovakia, Slovenia, Spain (incl. Balearic Is [Mallorca, Menorca]), Sweden, Switzerland, Turkey (European part: widespread), Ukraine; Russia: RUN, RUW, RUC, North Caucasus; Georgia, Azerbaijan, Turkey (Asiatic part: see citations for provinces), Israel, Iran;; Kazakhstan (east), Turkmenistan (Kopet Dag), Afghanistan.
Note: see the manual for abbreviations: present-day Russia.
Region(s):Westpalaearctic;; Eastpalaearctic
General note:Added: Ankara (Lackschewitz, 1940a), Lesvos, Mykonos (Museum Naturalis, Leiden). For Italy check Stary and Oosterbroek, 1996, and citations below. For Finland check Salmela, 2012b, 2012c, and citations below. For Norway check Olsen et al., 2018, and citations below. For Spain check Eiroa and Carles-Tolra, 2019, and citations below. For Portugal check Oosterbroek et al., 2020, and citations below. For Great Britain and Ireland check Stubbs, 2021, and citations below. Nearctic Region in Palaearctic Catalogue (Savchenko et al., 1992) refers to subspecies angularis. Removed, as found in earlier CCW versions: Armenia (excluded for Armenia by Hakobyan and Jenderedjian, 2023).
Map:Koc, 2004 (Mugla); Podenas et al., 2006 (Switzerland); Ashe et al., 2007b (Ireland); Boardman, 2007 (Shropshire)
Biology:Saltmarsh specialist (Gibbs, 1992).
Figure
references:
Habitus: Quindroit, 2020a (female); Ferreira et al., 2021 (female); Kramer and Morris, 2022a (female)
Wing: Podenas et al., 2006; Bratton, 2013
Hypopygium: Podenas et al., 2006; Kramer and Morris, 2022a

Immature stages
Larva: Podeniene, 2003a; Podeniene, 2009
Citationson biology (mainly from 2000 onward):
Azerbaijan
Habitat. Stary and Obona, 2020: 134-135 (loc(s); with list of localities, habitats and altitudes).
Bulgaria
Habitat. Hubenov, 2025b: suppl: 4 (annotated list Diptera high Bulgarian mts, local distr, habitats and zoogeography).
Country not relevant
Habitat. Krivosheina and Krivosheina, 2011c: 1-294 (larva: descr, figs, biology, key) (in Russian, key also in English).
Czech Rep.
Habitat. Stary and Vonicka, 2018: 62 (on Limoniidae and Pediciidae of two localities in northern Bohemia, with list of localities, altitudes and habitats).
Denmark
Habitat. Byriel and Rojas, 2017: 118 (among the 101 species in a study on cranefly hotspots in unmanaged and managed forests) (in Danish).
Finland
Habitat. Autio and Salmela, 2010: 48, 51 (habitats Aland Is, inhabitant of Baltic shore meadows and marshlands).
Habitat. Rasimus, 2022: 6 (on the craneflies fauna of Tavastia australis biogeographical province) (in Finnish).
France
Biology. Quindroit, 2020a: 29, 45-48 (annotated list with information on abundancy and habitat of the Tipuloidea of the Pays de la Loire regoin, distr) (in French).
Habitat. Quindroit, 2020b: 142 (loc(s) Nord (59) and Pas-de-Calais (62) with notes on habitats and distr) (in French).
Habitat. Herbrecht et al., 2021: 57 (loc(s) Loire-Atlantique (44) based on malaise-trapping with notes on habitats) (in French).
Habitat. Quindroit and Lemoine, 2022a: 28 (loc(s) Aisne (02) and Somme (80) with notes on habitats and distr) (in French).
Georgia
Habitat. Stary and Obona, 2020: 135 (loc(s); with list of localities, habitats and altitudes).
Germany
Habitat. Dau, 2013: 42, Anhang 1: 33 (among species found in Salzquellen in Schleswig-Holstein) (also as similis).
Great Britain
Habitat. Godfrey, 2000a: 20.
Habitat. Godfrey, 2001a: 21, 23, 32 (reared from woody material).
Habitat. Alexander, 2002: 89 (develops in wet soil, has been reared from rot-hole material).
Habitat. Godfrey, 2003a: 25, 44 (emerged from woody debris taken from a log, possibly a facultative xylophage; review literature).
Habitat. Stubbs, 2003: 31 (of ecological interest in being stunted and with short wings in lower and middle saltmarsh but more normal fully developed individuals are frequent in higher saltmarsh).
Habitat. Stubbs, 2003: 31, 95 (often plentiful in all zones of coastal saltmarshes; of ecological interest in being stunted and with short wings in lower and middle saltmarsh but more normal fully developed individuals are frequent in higher saltmarsh; also occurring inland in brackish and freshwater muddy marginal situations; populations are suppressed by close grazing but survive).
Habitat. Boardman, 2007: 58 (wet places).
Habitat. Kramer, 2009g: 2 (recorded from dune scrub and/or dune grassland).
Habitat. Irwin, 2010: 169 (a marine, estuarine or brackish species that can develop in mud).
Habitat. Bratton, 2013: 8 (reared from mud collected from among reed litter to the waterline of a brackish pond (6.8% seawater)).
Habitat. Showers, 2015: 2 (on craneflies in moth traps at Pitsford Water Nature Reserve in 2013 and 2014, including information for the two traps and photos of their habitat).
Habitat. Hancock and Horsfield, 2017: 8 (reduced wings can occur with S. stictica (Meigen) in Britain on exposed saltmarshes, citing Stubbs, 2003).
Habitat. Drake, 2019: 169 (flies from inland saltmarshes, loc(s) Cheshire).
Habitat. Kramer and Morris, 2022a: 23 (info on the species, on the distr and map of loc(s) Leicestershire and Rutland (VC55); a fly of marshland and often abundant in salt-marshes).
Iceland
Habitat. Kolcsar et al., 2021b: 234-235 (loc(s) [Olsen in litt., 2018: collected at a salt marsh (see habitat photo(s) above]).
Ireland
Habitat. Ashe et al., 2007b: 392 (bred from peaty soil, citing Blackith et al., 1991).
Italy
Habitat. Stary, 2002a: 106 (semi-dry Quercus forest and neighbouring open habitats).
Habitat. Stary, 2003a: 125.
Lithuania
Habitat. Podeniene, 2003a: 1-295 (larva, descr, figs, habitat) (in Lithuanian).
Habitat. Podeniene, 2009: 137 (can be found in very different wet habitats).
Netherlands
Habitat. Dek and Oosterbroek, 2013: 87 (collected in a damp deciduous forest with undergrowth largely of Aegopodium podagraria and forest floor with much dead wood).
Habitat. Drake et al., 2019: 75-77 (on Diptera of inland saltmarshes in Cheshire and Staffordshire).
Portugal
Habitat. Stary, 2014b: 86-87 (collected at saltmarshes).
Romania
Habitat. Parvu, 2003: 233.
Russia
Habitat. Lantsov, 2009e: 119 (a typically halophilous species, part of the coastal community of Lake Sukhoye).
Habitat. Lantsov, 2011b: 70 (on ecological groups of tipuloid larvae in the Caucasus) (in Russian).
Habitat. Lantsov, 2017b: 107 (on dominant species of the crane flies (Diptera, Tipuloidea) of the North Caucasus as biological indicators of habitats) (in Russian).
Habitat. Lantsov, 2017c: 286 (loc(s) North Caucasus: Dagestan with information on habitats and ecology) (in Russian).
Slovakia
Habitat. Beuk et al., 2023: 46, 54 (wetland at the bottom of the hill).
Spain
Habitat. Stary, 2014b: 87 (collected at saltmarsh and canal).
Sweden
Biology. Salmela, 2010b: 127-129 (a species of brackish or saline habitats, citing various authors; phenology diagram, peak abundance in april-may).
Turkey
Biology. Koc, 2004: 91-92, 116-118, 151 (habitat; phenology) (in Turkish).
on characters and taxonomy (mainly from 2000 onward):
Country not relevant
Characters. Chandler and Crossley, 2003: 50 (comparison).
Immatures. Krivosheina and Krivosheina, 2011c: 1-294 (larva: descr, figs, biology, key) (in Russian, key also in English).
Great Britain
Characters. Stubbs, 2003: 31 (of ecological interest in being stunted and with short wings in lower and middle saltmarsh but more normal fully developed individuals are frequent in higher saltmarsh).
Characters. Boardman, 2007: 58 (characters).
Characters. Bratton, 2013: 8 (figs males with and without discal cell).
Characters. Hancock and Horsfield, 2017: 8 (reduced wings can occur with S. stictica (Meigen) in Britain on exposed saltmarshes, citing Stubbs, 2003).
Characters. Kramer and Morris, 2022a: 23 (diagnosis, figs).
Israel
Key. Stary and Freidberg, 2007: 302-307, 327-328 (only known from older literature, confirmation required, key).
Lithuania
Immatures. Podeniene, 2003a: 1-295 (larva, descr, figs, habitat) (in Lithuanian).
Portugal
Characters. Ferreira et al., 2021: 6 (barcoding based on material mentioned in Oosterbroek et al. 2020).
on distribution (mainly from 2000 onward):
Albania
Distribution. Kolcsar et al., 2023b: 29-30 (loc(s)).
Armenia
Misinterpretation. Obona et al., 2016: 139 (checklist but excluded for Armenia by Hakobyan and Jenderedjian, 2023).
Azerbaijan
Distribution. Stary and Obona, 2020: 134-135 (loc(s); with list of localities, habitats and altitudes).
Distribution. Snegovaya, 2021: 1145 (checklist with details on distr in Azerbaijan).
Belgium
Distribution. Martens et al., 2014: 114, 124 (loc(s) Oost-Vlaanderen).
Distribution. Dek and Peeters, 2023a: 140 (loc(s) Botanic Garden Jean Massart, Brussels-Capital Region, common in Belgium).
Bosnia-Herzegowina
Distribution. Oosterbroek and Simova-Tosic, 2004: 445 (review literature).
Bulgaria
Distribution. Hubenov, 2021a: 32 (checklist with summary of distr in Bulgaria, references).
Distribution. Hubenov, 2025b: suppl: 4 (annotated list Diptera high Bulgarian mts, local distr, habitats and zoogeography).
Channel Is
Distribution. Stubbs, 2022b: 5 (on a collection of craneflies from Jersey, collected between 1985 and 2002 by Tony Warne and present in the Societe Jersiaise collection, identifier unknown).
Croatia
Distribution. Oosterbroek and Simova-Tosic, 2004: 445 (review literature).
Czech Rep.
Distribution. Stary et al., 2005a: 30 (loc(s)).
Distribution. Stary, 2009i: webpage (checklist).
Distribution. Stary and Vonicka, 2018: 62 (on Limoniidae and Pediciidae of two localities in northern Bohemia, with list of localities, altitudes and habitats).
Denmark
Distribution. Petersen and Jong, 2001b: 151 (checklist).
Estonia
Distribution. Stary, 2004f: (Fauna Europaea: added Estonia).
Finland
Distribution. Autio and Salmela, 2010: 48 (loc(s) Aland Is, recorded from the whole finnish coastal area southwards from the gulf of Bothnia).
Distribution. Salmela, 2012b: 239 (annotated list of Finnish crane flies).
Distribution. Salmela and Petrasiunas, 2014: 26 (checklist Finnish Tipulomorpha).
Distribution. Rasimus, 2022: 6 (on the craneflies fauna of Tavastia australis biogeographical province) (in Finnish).
France
Distribution. Garrin and Herbrecht, 2016: Annexe 11 (loc(s) Notre-Dame-des-Landes, Loire-Atlantique (44)).
Distribution. Quindroit, 2020a: 29, 45-48 (annotated list with information on abundancy and habitat of the Tipuloidea of the Pays de la Loire regoin, distr) (in French).
Distribution. Quindroit, 2020b: 142 (loc(s) Nord (59) and Pas-de-Calais (62) with notes on habitats and distr) (in French).
Distribution. Herbrecht et al., 2021: 57 (loc(s) Loire-Atlantique (44) based on malaise-trapping with notes on habitats) (in French).
Distribution. Quindroit and Lemoine, 2022a: 28 (loc(s) Aisne (02) and Somme (80) with notes on habitats and distr) (in French).
Distribution. Quindroit and Lemoine, 2022a: 36 (table specifying the presence in the five departments of the Hauts-de-France region).
Distribution. Quindroit and Racine, 2025: 358 (table specifying the presence in the five departments of the Pays de la Loire region).
Georgia
Distribution. Stary and Obona, 2020: 135 (loc(s); with list of localities, habitats and altitudes).
Germany
Distribution. Schacht, 1999: 134 (checklist Bayern).
Distribution. Reusch and Oosterbroek, 2000: 160 (checklist German Bundeslander).
Distribution. Reusch and Bellstedt, 2001b: 74 (checklist Thuringen).
Distribution. Schacht, 2005: 6 (checklist Bayern).
Distribution. Schacht, 2010: 18 (checklist Diptera Bayern).
Distribution. Dau, 2013: 42, Anhang 1: 33 (among species found in Salzquellen in Schleswig-Holstein) (also as similis).
Distribution. Dunk and Kraus, 2014: 169 (loc(s) Bayern).
Distribution. Stuke, 2019: 146 (checklist Niedersachsen and Bremen with reference to original sources).
Great Britain
Distribution. Godfrey, 2000a: 20 (see paper for details of sites surveyed in Shropshire and into Wales).
Distribution. Godfrey, 2001a: 21, 23, 32 (recorded from sites in Shropshire and just over the country border in Montgmeryshire; see paper for details).
Distribution. Godfrey, 2003a: 25, 44 (loc(s), distr, review literature).
Distribution. Hewitt et al., 2005: 38 (loc(s) Cumbria) (as Erioptera).
Distribution. Boardman, 2007: 58 (loc(s), map).
Distribution. Skidmore, 2008a: 59 (loc(s) Tiree, Western Isles, Scotland).
Distribution. Kramer, 2009g: 2 (loc(s) Swansea).
Distribution. Skidmore, 2009: 127 (review of occurrence on the Western Isles of Scotland, distr).
Distribution. Hancock, 2011a: 2 (loc(s) Tiree and Coll, Southern Inner Hebrides).
Distribution. Kramer, 2011g: 2 (loc(s) Devon).
Distribution. Kramer, 2011j: 17 (loc(s) Leicestershire and Rutland (VC55)).
Distribution. Bratton, 2013: 8 (loc(s) North Wales).
Distribution. Hewitt, 2014: (provisional checklist Cumbrian Diptera).
Distribution. Kramer, 2014c: 4 (in table of cranefly species recorded at Shapwick Heath NNR, as given in H. Audcent, 1949 and/or collected by J. Kramer in 2013).
Distribution. Wolton et al., 2014: 27 (among Diptera associated with a hedge in Devon).
Distribution. Chandler, 2015: 89 (loc(s) Bushy Park, Middlesex).
Distribution. Showers, 2015: 2 (on craneflies in moth traps at Pitsford Water Nature Reserve in 2013 and 2014, including information for the two traps and photos of their habitat).
Distribution. Drake, 2016: 20 (study on the relative importance to Diptera of pasture and ditch margins on Catcott Reserve, Somerset).
Distribution. Robinson et al., 2016: 79 (list of terrestrial invertebrates of Mingulay Is, Outer Hebrides).
Distribution. Brighton, 2017c: 27 (overall regional checklist Lancashire and Cheshire (VC58, 59 and 60) with notes on habitats and/or regional occurrence and/or status; see paper for details).
Distribution. Drake, 2019: 169 (flies from inland saltmarshes, loc(s) Cheshire).
Distribution. Kramer and Morris, 2022a: 23 (info on the species, on the distr and map of loc(s) Leicestershire and Rutland (VC55); a fly of marshland and often abundant in salt-marshes).
Greece
Distribution. Oosterbroek, 2009c: 181 (new for Lesvos and Mykonos).
Hungary
Distribution. Toth, 2005: 169 (loc(s)).
Iceland
First record. Kolcsar et al., 2021b: 234-235 (loc(s) [Olsen in litt., 2018: collected at a salt marsh (see habitat photo(s) above]).
Ireland
Distribution. Ashe et al., 2007b: 363, 391-392, 406 (review Irish records, map).
Distribution. Ashe et al., 2008: 8 (refers to record in Speight, 1990).
Distribution. Chandler et al., 2008: 11 (checklist).
Israel
Distribution. Stary and Freidberg, 2007: 302-307, 327-328 (only known from older literature, confirmation required, distr).
Italy
Distribution. Stary, 2002a: 106 (loc(s)).
Distribution. Stary, 2003a: 125 (loc(s)).
Kazakhstan
First record. Devyatkov, 2019b: 245 (loc(s) Pavlodarskaya oblast, distr) (in Russian).
Latvia
Distribution. Stary, 2004f: (Fauna Europaea: added Latvia).
Lithuania
Distribution. Pakalniskis et al., 2006: 24 (checklist).
Malta
First record. Ebejer, 2015: 53 (loc(s) Malta and Gozo).
Distribution. Ebejer and Gatt, 2021: 17 (checklist of the Diptera of Malta).
Netherlands
Distribution. Jong and Oosterbroek, 2002b: 30 (checklist).
Distribution. Cuppen, 2009: 7 (loc(s) larval records).
Distribution. Dek and Oosterbroek, 2013: 87 (loc(s) Zeeland).
Distribution. Hoven, 2018: 16 (loc(s) Zeeland).
Distribution. Drake et al., 2019: 75-77 (on Diptera of inland saltmarshes in Cheshire and Staffordshire).
Distribution. Steenis, 2019: 15 (loc(s) Noord-Brabant, Biesbosch).
Distribution. Oosterbroek and Dek, 2020: 17 (Limoniidae and Tipulidae Nationaal Park Hollandse Duinen in 2018, Zuid-Holland).
Distribution. Stary, 2020a: 45 (Limoniidae Kaaistoep, Noord-Brabant).
North Macedonia
Distribution. Oosterbroek and Simova-Tosic, 2004: 445 (review literature).
Norway
Distribution. Olsen et al., 2018: 156-157 (annotated checklist Nordic countries and for Norway according to the Strand regions).
Poland
Distribution. Wiedenska, 2007a: 76 (checklist).
Distribution. Wiedenska and Syratt, 2023: 135 (loc(s) Bieszczady Mountains, distr and remarks).
Portugal
First record. Stary, 2014b: 86-87 (loc(s) Faro).
Distribution. Oosterbroek et al., 2020: 324 (loc(s) Aveiro and Coimbra, also known from Faro, distr).
Romania
Distribution. Parvu, 2003: 233 (loc(s)).
Distribution. Parvu, 2004: 185 (loc(s)).
Distribution. Ujvarosi, 2007: 228-231 (checklist Limoniidae).
Russia
Distribution. Lantsov, 2003e: 81 (loc(s) Caucasus region, distr).
Distribution. Lantsov, 2004: 63, 67 (loc(s) Caucasus region, distr).
Distribution. Humala and Polevoi, 2008: 133 (loc(s) RUN: Kareliya).
Distribution. Lantsov, 2009e: 119 (loc(s) North Caucasus: Lake Sukhoye).
Distribution. Lantsov, 2011c: 86-87 (loc(s) RUN: Arkhangelskaya oblast, Nenetskiy AO, distr, first record for Subarctic Russia) (in Russian).
Distribution. Paramonov and Pilipenko, 2016: 97 (loc(s) RUC: Tverskaya oblast, distr) (in Russian).
Distribution. Lantsov, 2017b: 107 (on dominant species of the crane flies (Diptera, Tipuloidea) of the North Caucasus as biological indicators of habitats) (in Russian).
Distribution. Lantsov, 2017c: 286 (loc(s) North Caucasus: Dagestan with information on habitats and ecology) (in Russian).
Serbia
Distribution. Oosterbroek and Simova-Tosic, 2004: 445 (review literature).
Distribution. Gavryushin in litt., 2015b: (loc(s) Serbia).
Slovakia
Distribution. Stary, 2009d: 33 (loc(s) Polana area).
Distribution. Stary, 2009i: webpage (checklist).
Distribution. Beuk et al., 2023: 46, 54 (loc(s)).
Slovenia
Distribution. Oosterbroek and Simova-Tosic, 2004: 445 (review literature).
Spain
Distribution. Eiroa and Baez, 2002a: 54, 56 (checklist, loc(s)).
Distribution. Carles-Tolra and Ventura, 2009: 162 (loc(s) Menorca).
Distribution. Stary, 2014b: 87 (loc(s) Mallorca).
Distribution. Eiroa and Carles-Tolra, 2019: 131 (loc(s) Barcelona, list of Spanish provinces, distr) (in Spanish).
Distribution. Keresztes et al., 2022: 193 (loc(s) Asturias and review distribution in Spain) (in Spanish).
Distribution. Mederos et al., 2025b: 159 (loc(s) Gerona, distr).
Sweden
Distribution. Salmela, 2010b: 133 (loc(s) Malmo).
Switzerland
Distribution. Podenas et al., 2006: 130, 341 (map, vertical distr per thermic level).
Distribution. Dufour and Merz, 2012: 354 (annotated checklist canton Geneva).
Turkey
Distribution. Koc, 2004: 91-92, 116-118, 151 (loc(s), distr, map) (in Turkish).
Distribution. Ozgul et al., 2009: 64 (survey provinces SW Turkey).
Distribution. Bilgin et al., 2015: 5 (loc(s) Eskisehir, distr) (in Turkish).
Distribution. Koc et al., 2016: 11 (provinces Marmara region).
Distribution. Akpinar and Ozgul, 2023: 55 (loc(s) Aksaray).
Distribution. Provinces Asiatic part, 2025: (known from Ankara, Aydin, Burdur, Denizli, Eskisehir, Mugla, Marmara region).
on flight period (mainly from 2000 onward):
Albania
Kolcsar et al., 2023b: month(s): 8.
Azerbaijan
Stary and Obona, 2020: month(s): 5.
Belgium
Martens et al., 2014: month(s): 5, 8-9.
Dek and Peeters, 2023a: month(s): 4, 6, 8.
Channel Is
Stubbs, 2022b: month(s): 6.
Czech Rep.
Stary et al., 2005a: month(s): 4-5, 7-9.
Stary and Vonicka, 2018: month(s): 5-7, 9.
Finland
Rasimus, 2022: month(s): 9.
France
Quindroit, 2020a: month(s): 4-7.
Quindroit, 2020b: month(s): 3-4.
Quindroit and Lemoine, 2022a: month(s): 5, 7.
Georgia
Stary and Obona, 2020: month(s): 5.
Great Britain
Godfrey, 2000a: month(s): 6.
Godfrey, 2001a: month(s): 5.
Skidmore, 2008a: month(s): 6.
Kramer, 2009g: month(s): 7.
Hancock, 2011a: month(s): 6.
Kramer, 2011j: month(s): 4, 6.
Bratton, 2013: month(s): 11.
Kramer, 2014c: month(s): 6.
Chandler, 2015: month(s): 5-8.
Drake, 2019: month(s): 6.
Iceland
Kolcsar et al., 2021b: month(s): 7.
Ireland
Ashe et al., 2007b: month(s): 3-10.
Kazakhstan
Devyatkov, 2019b: month(s): 8.
Malta
Ebejer, 2015: month(s): 3-4.
Netherlands
Dek and Oosterbroek, 2013: month(s): 10.
Hoven, 2018: month(s): 5.
Oosterbroek and Dek, 2020: month(s): 3-5, 7.
Poland
Wiedenska and Syratt, 2023: month(s): 7.
Portugal
Stary, 2014b: month(s): 3-5.
Oosterbroek et al., 2020: month(s): 2, 5.
Romania
Parvu, 2003: month(s): 7.
Parvu, 2004: month(s): 6.
Russia
Lantsov, 2003e: month(s): 6.
Lantsov, 2004: month(s): 4-6.
Lantsov, 2011c: month(s): 7.
Lantsov, 2017c: month(s): 5.
Serbia
Gavryushin in litt., 2015b: month(s): 7.
Slovakia
Stary, 2009d: month(s): 6-7.
Beuk et al., 2023: month(s): 5.
Spain
Eiroa and Baez, 2002a: month(s): 5.
Carles-Tolra and Ventura, 2009: month(s): 4.
Stary, 2014b: month(s): 5.
Eiroa and Carles-Tolra, 2019: month(s): 4-5, 11.
Keresztes et al., 2022: month(s): 8.
Mederos et al., 2025b: month(s): 3.
Sweden
Salmela, 2010b: month(s): 4-8.
Switzerland
Podenas et al., 2006: month(s): 4-11.
Turkey
Koc, 2004: month(s): 3, 5-6.
on altitude (mainly from 2000 onward):
Albania
Kolcsar et al., 2023b: altitude: 1290 m.
Azerbaijan
Stary and Obona, 2020: altitude: 200-105 m.
Bulgaria
Hubenov, 2021a: altitude: 0-1500 m.
Hubenov, 2025b: altitude: 0-1500 m.
Georgia
Stary and Obona, 2020: altitude: 465 m.
Iceland
Kolcsar et al., 2021b: altitude: 5 m.
Italy
Stary, 2003a: altitude: 1500-1600 m.
Kazakhstan
Devyatkov, 2019b: altitude: 134 m.
Poland
Wiedenska and Syratt, 2023: altitude: 650 m.
Portugal
Stary, 2014b: altitude: 11-24 m.
Russia
Lantsov, 2003e: altitude: 650 m.
Slovakia
Beuk et al., 2023: altitude: 460 m.
Spain
Stary, 2014b: altitude: 0-1 m.
Keresztes et al., 2022: altitude: 574 m.
Turkey
Koc, 2004: altitude: 5-700 m.

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