Details for Molophilus (Molophilus) appendiculatus

Name: Molophilus (Molophilus) appendiculatus (Staeger, 1840) Publication: Naturh. Tidsskr. 3: 55 (as Erioptera) Status: Recognized taxon, Synonym(s): armatus Meijere, 1918. Classification: Family Limoniidae Subfamily Chioneinae Descriptive note: Aberrant Scottish material is discussed and figured in Hancock, 2005. Keys: Boardman, 2016 (craneflies Shropshire); Stubbs and Kramer, 2016k; Stubbs, 2021 (British craneflies) Variant spelling(s): appendiculata habitus male habitus male habitus male habitus male hypopygium plus 18 more images of hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, wing, habitat, habitat, habitat, habitat, habitat, habitat, habitat, habitat, habitat, habitat and habitat. Distribution: Austria, Belgium, Bulgaria, Croatia, Czech Rep., Denmark, Estonia, Finland, France, Germany, Great Britain, Hungary, Ireland, Italy, Latvia, Lithuania, Luxembourg, Montenegro, Netherlands, Norway, Poland, Romania, Serbia, Slovakia, Slovenia, Spain, Sweden, Switzerland, Turkey (European part: Kirklareli), Ukraine; Russia: RUN, RUW, RUC, RUE; Turkey (Asiatic part: Canakkale, Eskisehir, Kutahya);; Russia: WS (Altay); Kazakhstan (east). Note: see the manual for abbreviations: present-day Russia. Region(s): Westpalaearctic;; Eastpalaearctic General note: For Italy check Stary and Oosterbroek, 1996, and citations below. For Finland check Salmela, 2012b, 2012c, and citations below. For Norway check Olsen et al., 2018, and citations below. For Great Britain and Ireland check Stubbs, 2021, and citations below. Map: Podenas et al., 2006 (Switzerland); Ashe et al., 2007b (Ireland); Boardman, 2007 (Shropshire) Figure references: Hypopygium: Hancock, 2005 (aberrant male); Podenas et al., 2006; Kramer and Morris, 2022a Citations on biology (mainly from 2000 onward): Bulgaria Habitat. Hubenov, 2025b: suppl: 4 (annotated list Diptera high Bulgarian mts, local distr, habitats and zoogeography). Czech Rep. Habitat. Stary and Vonicka, 2018: 56 (on Limoniidae and Pediciidae of two localities in northern Bohemia, with list of localities, altitudes and habitats). Denmark Habitat. Nielsen and Nielsen, 2009: 121, 129, 131 (emerging from a forest floor in a Beech [Fagus] stand) (in Danish). Habitat. Byriel and Rojas, 2017: 118 (among the 101 species in a study on cranefly hotspots in unmanaged and managed forests) (in Danish). Finland Habitat. Salmela, 2001a: 144, 146, 150 (terrestrial species of more humid locations, potential semiaquatic, citing Brinkmann, 1991). Habitat. Rasimus, 2018: 17 (on saproxylic craneflies in the Tampere region) (in Finnish). France Biology. Quindroit, 2020a: 27-28, 45-48 (annotated list with information on abundancy and habitat of the Tipuloidea of the Pays de la Loire regoin, distr) (in French). Habitat. Quindroit, 2020b: 139 (loc(s) Nord (59) and Pas-de-Calais (62) with notes on habitats and distr) (in French). Habitat. Quindroit and Lemoine, 2021a: 25 (loc(s), resultats de deux campagnes de piegeage dans le Pas-de-Calais (62); see paper for details on habitat). Habitat. Quindroit, 2021a: 36 (loc(s), resultats d une campagne de piegeage dans le Pas-de-Calais (62); see paper for details on habitat). Habitat. Quindroit and Lemoine, 2022a: 27 (loc(s) Aisne (02) and Oise (60) with notes on habitats and distr) (in French). Germany Habitat. Hovemeyer and Schauermann, 2003: 72 (found in a study on succession of Diptera on dead Beach [Fagus] wood near Gottingen; see paper for details). Habitat. Reusch, 2006: 134 (Kalkquellmoor [carbonate fen complex]). Habitat. Lehmann and Reusch, 2009: 122 (from around lowland springs). Great Britain Habitat. Godfrey, 2000a: 19, 21. Habitat. Godfrey, 2001a: 11, 21-23. Habitat. Godfrey, 2003a: 24, 44 (adults of this almost certainly non-xylophagous species were taken in an emergence trap filled with semi-submerged logs, probably from the mud adhering to the woody debris in the trap). Habitat. Boardman, 2007: 49-50 (found in most types of wet woodland with streams in the lowlands). Habitat. Drake, 2011a: 13, 24 (among the Diptera from a wet Alder [Alnus] wood in Devon). Habitat. Kramer, 2011j: 15 (a common species of marshland). Biology. Hancock, 2015: 30-31 (Scottish data, from a Malaise trap set in Oak [Quercus]-woodland are from june-september, showing a distinct summer peak). Habitat. Showers, 2015: 2 (on craneflies in moth traps at Pitsford Water Nature Reserve in 2013 and 2014, including information for the two traps and photos of their habitat). Biology. Wolton, 2022: 6-8 (on observations on the phenology and sex ratios of craneflies (Limoniidae) found in emergencetraps in a wet woodland at Locks Park Farm, Devon; see paper for details on this species). Norway Habitat. Olsen and Andersen, 2021: 209 (loc(s) Innlandet, in former Hedmark, with habitats specified separately, for habitats see also Jonassen and Andersen, 2020). Poland Habitat. Chaniecka and Wiedenska, 2006: 148 (occurrence in spring-, springbrook-, and/or bogspring plant communities) (in Polish). Habitat. Wiedenska, 2014: 13 (on the occurence of Limoniidae in six selected plant communities in the Kamienica Valley, Gorce National Park, Western Carpathians) (in Polish, habitat tables also in English). Phenology. Wiedenska, 2015a: 53 (on the flight activity of Limoniidae in the Kamienica Valley, Gorce National Park, Western Carpathians) (in Polish, tables also in English). Habitat. Wiedenska, 2017a: 16 (faunistic and habitat data of species from the Gorce NP) (in Polish). Romania Habitat. Ujvarosi, 2005a: 246 (brooks, small river, swamp, moist soil, citing various sources). Habitat. Ujvarosi et al., 2011b: 107 (loc(s) Dupa Lanca marshy area near Voslobeni) (in Hungarian, species list with habitat information in English). Habitat. Kolcsar et al., 2013: 72 (loc(s) Cluj-Napoca and surroundings; list of habitats with altitudes). Russia Habitat. Paramonov and Pilipenko, 2016: 97 (habitat, citing Savchenko, 1962) (in Russian). Habitat. Kolcsar et al., 2021b: 159 (Tatarstan Rep.: ravine, wetland). Spain Habitat. Mederos et al., 2025b: 159 (loc(s) Lerida, distr; see habitat photo). Sweden Phenology. Salmela, 2010b: 127-28 (phenology diagram, most numerous in july to mid august). Habitat. Fritz and Lindstrom, 2013: 23, 62 (on distribution, habitat preference and habitat conservation in nature reserved forest and tree habitats in Hallands County) (in Schwedish). Habitat. Lindstrom and Fritz, 2015: 44, 48 (distributin and ecology in five fens in Hallands County) (in Schwedish). on characters and taxonomy (mainly from 2000 onward): Country not relevant Phylogeny. Ahonen, 2007: 1-28 (included in phylogenetic analysis of Tipuloidea subfamilies relationships based on molecular data). Characters. Boardman and Stary, 2020: 155 (comparison). Great Britain Taxonomy. Hancock, 2005: 62-63 (aberrant Scotisch male: characters, comparison, figs) Characters. Boardman, 2007: 49-50 (characters). Characters. Kramer and Morris, 2022a: 12 (diagnosis, figs). on distribution (mainly from 2000 onward): Austria Distribution. Reusch and Heiss, 2012: 323-332 (loc(s) Nat. Park Gesause, see PDF for months and altitudes). Bulgaria Distribution. Hubenov, 2015: 220 (annotated list Diptera Pirin Mt, loc(s), faunistics and zoogeography). Distribution. Hubenov, 2016: 48 (annotated checklist Diptera of the Rila Mountains based on data from the literature). Distribution. Hubenov, 2017: 68 (on vertical distribution in the Pirin and Rila Mts; see paper for details on forests and subalpine-alpine zones, distr). Distribution. Hubenov, 2021a: 32 (checklist with summary of distr in Bulgaria, references). Distribution. Hubenov, 2021a: 32 (checklist with summary of distr in Bulgaria, references). Distribution. Hubenov, 2025b: suppl: 4 (annotated list Diptera high Bulgarian mts, local distr, habitats and zoogeography). Channel Is Distribution. Stubbs, 2022b: 5 (on a collection of craneflies from Jersey, collected between 1985 and 2002 by Tony Warne). Croatia Distribution. Oosterbroek and Simova-Tosic, 2004: 444 (review literature). Distribution. Kolcsar et al., 2023b: 25 (loc(s)). Czech Rep. Distribution. Stary et al., 2005a: 28 (loc(s)). Distribution. Stary, 2009i: webpage (checklist). Distribution. Stary and Vonicka, 2018: 56 (on Limoniidae and Pediciidae of two localities in northern Bohemia, with list of localities, altitudes and habitats). Denmark Distribution. Petersen and Jong, 2001b: 150 (checklist). Distribution. Nielsen and Nielsen, 2009: 121 (loc(s)). Estonia Distribution. Stary, 2004f: (Fauna Europaea: added Estonia). Finland Distribution. Salmela, 2001a: 144, 146, 150 (loc(s)). Distribution. Salmela, 2002: 30-32 (loc(s) lake Iidesjarvi) (in Finnish). Distribution. Salmela et al., 2007b: 45 (loc(s)). Distribution. Salmela, 2010a: 12 (survey fauna Tahmela spring in city of Tampere) (in Finnish). Distribution. Salmela, 2012b: 238 (annotated list of Finnish crane flies). Distribution. Salmela and Petrasiunas, 2014: 25 (checklist Finnish Tipulomorpha). Distribution. Salmela et al., 2015: 103 (list of localities in extensive report on malaise-trapping of insects in conservation areas in Lapland in 2012–2014) (in Finnish). Distribution. Rasimus, 2018: 17 (on saproxylic craneflies in the Tampere region) (in Finnish). France Distribution. Kramer and Langlois, 2019a: 76 (record(s) Ravin de Valbois, Doubs (25)). Distribution. Kramer and Langlois, 2019b: 227 (list of craneflies captured 25-27 june 2019 in Ravin de Valbois, Doubs (25)). Distribution. Quindroit, 2020a: 27-28, 45-48 (annotated list with information on abundancy and habitat of the Tipuloidea of the Pays de la Loire regoin, distr) (in French). Distribution. Quindroit, 2020b: 139 (loc(s) Nord (59) and Pas-de-Calais (62) with notes on habitats and distr) (in French). Distribution. Labat, 2021: 92 (on Diptera taxa recorded along the over-all length of the Dordogne River and some of its tributaries; see paper for the locality details) (in French). Distribution. Quindroit and Lemoine, 2021a: 25 (loc(s), resultats de deux campagnes de piegeage dans le Pas-de-Calais (62); see paper for details on habitat). Distribution. Quindroit, 2021a: 36 (loc(s), resultats d une campagne de piegeage dans le Pas-de-Calais (62); see paper for details on habitat). Distribution. Tissot et al., 2021: 209 (the Diptera of the Reserves Naturelles Nationales du Doubs (25), results of Malaise trapping since 2009) (in French). Distribution. Quindroit and Lemoine, 2022a: 27 (loc(s) Aisne (02) and Oise (60) with notes on habitats and distr) (in French). Distribution. Quindroit and Lemoine, 2022a: 36 (table specifying the presence in the five departments of the Hauts-de-France region). Distribution. Lemoine and Quindroit, 2023: 9 (loc(s) Hautes-Alpes (05)). Distribution. Quindroit and Racine, 2025: 358 (table specifying the presence in the five departments of the Pays de la Loire region). Germany Distribution. Schacht, 1999: 135 (checklist Bayern). Distribution. Reusch and Oosterbroek, 2000: 158 (checklist German Bundeslander). Distribution. Reusch and Bellstedt, 2001b: 73 (checklist Thuringen). Distribution. Hovemeyer and Schauermann, 2003: 72 (found in a study on succession of Diptera on dead Beach [Fagus] wood near Gottingen; see paper for details). Distribution. Schacht, 2005: 5 (checklist Bayern). Distribution. Reusch, 2006: 134 (loc(s) Bayern). Distribution. Lehmann and Reusch, 2009: 122 (loc(s) Niedersachsen). Distribution. Hable et al., 2010b: 91 (loc(s) Bayern). Distribution. Schacht, 2010: 17 (checklist Diptera Bayern). Distribution. Stuke, 2019: 144 (checklist Niedersachsen and Bremen with reference to original sources). Great Britain Distribution. Godfrey, 2000a: 19, 21 (see paper for details of sites surveyed in Shropshire and into Wales). Distribution. Godfrey, 2001a: 11, 21-23 (recorded from sites in Shropshire and just over the country border in Montgmeryshire; see paper for details). Distribution. Godfrey, 2003a: 24, 44 (loc(s), distr). Distribution. Hancock, 2005: 62-63 (loc(s) aberrant Scottisch male) Distribution. Hewitt et al., 2005: 38 (loc(s) Cumbria). Distribution. Boardman, 2007: 49-50 (distr, map). Distribution. Skidmore, 2008a: 59 (loc(s) Tiree, Western Isles, Scotland). Distribution. Skidmore, 2008b: 68 (loc(s) Kerrera, Western Isles, Scotland). Distribution. Skidmore, 2009: 126 (review of occurrence on the Western Isles of Scotland, distr). Distribution. Drake, 2011a: 24 (loc(s) Devon). Distribution. Kramer, 2011j: 15 (loc(s) Leicestershire and Rutland (VC55)). Distribution. Kramer, 2013i: 8 (checklist of species from Leigh Woods NNR, Bristol district). Distribution. Hewitt, 2014: (provisional checklist Cumbrian Diptera). Distribution. Kramer, 2014c: 4 (in table of cranefly species recorded at Shapwick Heath NNR, as given in H. Audcent, 1949 and/or collected by J. Kramer in 2013). Distribution. Wolton et al., 2014: 27 (among Diptera associated with a hedge in Devon, caught in emergence trap). Distribution. Chandler, 2015: 89 (loc(s) Bushy Park, Middlesex). Distribution. Hancock, 2015: 31 (loc(s) Scotland). Distribution. Showers, 2015: 2 (on craneflies in moth traps at Pitsford Water Nature Reserve in 2013 and 2014, including information for the two traps and photos of their habitat). Distribution. Brighton, 2017c: 26 (overall regional checklist Lancashire and Cheshire (VC58, 59 and 60) with notes on habitats and/or regional occurrence and/or status; see paper for details). Distribution. Kramer and Morris, 2022a: 12 (info on the species, on the distr and map of loc(s) Leicestershire and Rutland (VC55)). Distribution. Wolton, 2022: 6-8 (on observations on the phenology and sex ratios of craneflies (Limoniidae) found in emergencetraps in a wet woodland at Locks Park Farm, Devon; see paper for details on this species). Ireland Distribution. Ashe et al., 2007b: 362, 377, 404 (review Irish records, map). Distribution. Chandler et al., 2008: 10 (checklist). Italy Distribution. Podenas and Podeniene, 2008: 348, 351 (loc(s), annotated checklist South Tyrol [Prov. Bolzano]). Kazakhstan First record. Devyatkov, 2020b: 221 (loc(s) East Kazakhstan, distr) (in Russian). Lithuania Distribution. Pakalniskis et al., 2006: 22 (checklist). Luxembourg Distribution. Vogtenhuber, 2007: 354 (loc(s)). Montenegro Distribution. Oosterbroek and Simova-Tosic, 2004: 444 (review literature). Netherlands Distribution. Jong and Oosterbroek, 2002b: 29 (checklist). Distribution. Oosterbroek and Dek, 2020: 16 (Limoniidae and Tipulidae Nationaal Park Hollandse Duinen in 2018, Zuid-Holland). Distribution. Stary, 2020a: 45 (Limoniidae Kaaistoep, Noord-Brabant). Norway Distribution. Olsen et al., 2018: 154-155 (annotated checklist Nordic countries and for Norway according to the Strand regions). Distribution. Olsen and Andersen, 2021: 209 (loc(s) Innlandet, in former Hedmark, with habitats specified separately, for habitats see also Jonassen and Andersen, 2020). Poland Distribution. Chaniecka and Wiedenska, 2006: 148 (loc(s)). Distribution. Wiedenska, 2007a: 76 (checklist). Distribution. Wiedenska, 2008: 45 (loc(s) Parowy Janinowski nature reserve near Lodz) (in Polish). Distribution. Wiedenska, 2010: 134 (loc(s) Parku Krajobrazowego Wzniesien Lodzkich, Lodz). Distribution. Wiedenska, 2014: 13 (on the occurence of Limoniidae in six selected plant communities in the Kamienica Valley, Gorce National Park, Western Carpathians) (in Polish, habitat tables also in English). Distribution. Wiedenska, 2017a: 16 (faunistic and habitat data of species from the Gorce NP) (in Polish). Distribution. Wiedenska and Syratt, 2023: 137 (loc(s) Bieszczady Mountains, distr and remarks). Romania Distribution. Parvu, 2004: 185 (loc(s)). Distribution. Ujvarosi, 2007: 228-231 (checklist Limoniidae). Distribution. Ujvarosi et al., 2011b: 107 (loc(s) Dupa Lanca marshy area near Voslobeni) (in Hungarian, species list with habitat information in English). Distribution. Kolcsar et al., 2013: 72 (loc(s) Cluj-Napoca and surroundings; list of habitats with altitudes). Russia Distribution. Polevoi, 2006: 96 (loc(s) RUN: Kareliya) (in Russian). Distribution. Paramonov and Pilipenko, 2016: 97 (loc(s) RUC: Tverskaya oblast, distr) (in Russian). Distribution. Kolcsar et al., 2021b: 157-159 (loc(s) RUE: Bashkortostan Rep. and Tatarstan Rep.). Serbia Distribution. Oosterbroek and Simova-Tosic, 2004: 444 (review literature). Slovakia Distribution. Stary, 2009d: 29 (loc(s) Polana area). Distribution. Stary, 2009i: webpage (checklist). Slovenia Distribution. Oosterbroek and Simova-Tosic, 2004: 444 (review literature). Distribution. Kolcsar et al., 2023b: 25 (loc(s)). Spain Distribution. Eiroa and Baez, 2002a: 54 (checklist). Distribution. Mederos et al., 2025b: 159 (loc(s) Lerida, distr; see habitat photo). Sweden Distribution. Salmela, 2010b: 126, 129, 133 (loc(s) Malmo, abundance, distr). Distribution. Fritz and Lindstrom, 2013: 23, 62 (on distribution, habitat preference and habitat conservation in nature reserved forest and tree habitats in Hallands County) (in Schwedish). Distribution. Lindstrom and Fritz, 2015: 44, 48 (distributin and ecology in five fens in Hallands County) (in Schwedish). Switzerland Distribution. Podenas et al., 2006: 106, 286 (map, vertical distr per thermic level). Distribution. Dufour and Merz, 2012: 354 (annotated checklist canton Geneva). Turkey Distribution. Bilgin et al., 2015: 5, 8 (loc(s) Eskisehir, distr) (in Turkish). Distribution. Koc et al., 2016: 10 (provinces Marmara region). on flight period (mainly from 2000 onward): Bulgaria Hubenov, 2015: month(s): 8-9. Hubenov, 2016: month(s): 8-9. Channel Is Stubbs, 2022b: month(s): 6-7. Croatia Kolcsar et al., 2023b: month(s): 7. Czech Rep. Stary et al., 2005a: month(s): 5-6. Stary and Vonicka, 2018: month(s): (5)6-7, 9. Finland Salmela, 2001a: month(s): 6-8. France Kramer and Langlois, 2019b: month(s): 6. Quindroit, 2020a: month(s): 4-6. Quindroit, 2020b: month(s): 5, 7. Quindroit and Lemoine, 2022a: month(s): 6-8. Lemoine and Quindroit, 2023: month(s): 6. Great Britain Godfrey, 2000a: month(s): 6. Godfrey, 2001a: month(s): 5. Hancock, 2005: month(s): 6. Boardman, 2007: month(s): 5-9. Skidmore, 2008a: month(s): 6. Skidmore, 2008b: month(s): 6. Kramer, 2011j: month(s): 7-9. Kramer, 2014c: month(s): 6. Chandler, 2015: month(s): 5-8. Hancock, 2015: month(s): 6-9. Ireland Ashe et al., 2007b: month(s): 5-10. Italy Podenas and Podeniene, 2008: month(s): 8. Kazakhstan Devyatkov, 2020b: month(s): 6-8. Netherlands Oosterbroek and Dek, 2020: month(s): 4. Norway Olsen and Andersen, 2021: month(s): 6-7(-9). Poland Wiedenska, 2014: month(s): 7-9. Wiedenska, 2015a: month(s): 6-8. Wiedenska, 2017a: month(s): 6-8. Wiedenska and Syratt, 2023: month(s): 6-8. Romania Parvu, 2004: month(s): 7. Ujvarosi et al., 2011b: month(s): 5-6. Kolcsar et al., 2013: month(s): 6. Russia Paramonov and Pilipenko, 2016: month(s): 7. Kolcsar et al., 2021b: month(s): 6-8. Slovakia Stary, 2009d: month(s): 6-7. Slovenia Kolcsar et al., 2023b: month(s): 7. Spain Mederos et al., 2025b: month(s): 8. Switzerland Podenas et al., 2006: month(s): 5-9. on altitude (mainly from 2000 onward): Bulgaria Hubenov, 2015: altitude: 1950 m. Hubenov, 2016: altitude: 1230-1390 m. Hubenov, 2017: altitude: 1230-1390-1950 m. Hubenov, 2021a: altitude: 1230-1950 m. Hubenov, 2025b: altitude: 1230-1950 m. Croatia Kolcsar et al., 2023b: altitude: 346 m. France Labat, 2021: altitude: 1241 m. Italy Podenas and Podeniene, 2008: altitude: 940 m. Kazakhstan Devyatkov, 2020b: altitude: 300-1170 m. Norway Olsen and Andersen, 2021: altitude: 290-515 m. Poland Wiedenska, 2017a: altitude: 760-1140 m. Wiedenska and Syratt, 2023: altitude: 500-1000 m. Russia Kolcsar et al., 2021b: altitude: 80-777 m. Slovakia Stary, 2009d: altitude: 560-780 m. Slovenia Kolcsar et al., 2023b: altitude: 600 m. Spain Mederos et al., 2025b: altitude: 1200 m.