Details for Tricyphona (Tricyphona) immaculata
Name:Tricyphona (Tricyphona) immaculata (Meigen, 1804)
Publication:Klass. Beschr. 1: 58 (as Limonia)
Status:Recognized taxon.
Classification:Family Pediciidae
Subfamily Pediciinae
Keys:Boardman, 2016 (craneflies Shropshire); Peeters en Oosterbroek, 2016c (craneflies Netherlands, Belgium and Luxembourg) (in Dutch); Stubbs and Kramer, 2016f; Stubbs, 2021 (British craneflies)
Tricyphona (Tricyphona) immaculata : habitus - maleTricyphona (Tricyphona) immaculata : habitus - maleTricyphona (Tricyphona) immaculata : habitus - maleTricyphona (Tricyphona) immaculata : habitus - femaleTricyphona (Tricyphona) immaculata : habitus - female
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plus 36 more images of habitus, habitus, body part(s), body part(s), body part(s), body part(s), body part(s), body part(s), body part(s), hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, ovipositor, wing, wing, wing, habitat, habitat, habitat, habitat, habitat, habitat, habitat, habitat, habitat, habitat, habitat, habitat, habitat, habitat, habitat, habitat and habitat.
Distribution:Andorra, Austria, Belarus (Berezinsky BR), Belgium, Bosnia-Herzegovina, Bulgaria, Croatia, Czech Rep., Denmark (incl. Faroe Is), Estonia, Finland, France (incl. Corsica), Germany, Great Britain, Hungary, Ireland, Italy (incl. Sicily), Latvia, Lithuania, Luxembourg, Montenegro, Netherlands, North Macedonia, Norway, Poland, Portugal, Romania, Serbia, Slovakia, Slovenia, Spain, Sweden, Switzerland, Ukraine; Russia: Novaya Zemlya, RUN, RUW, RUC, RUE, North Caucasus; Morocco; Georgia, Azerbaijan, Turkey (Asiatic part: see citations for provinces), Lebanon;; Russia: WS (Altay); Kazakhstan (east), Kyrgyzstan.
Note: see the manual for abbreviations: present-day Russia.
Region(s):Westpalaearctic;; Eastpalaearctic
General note:Added: Andorra (Geiger et al., 1994). For Italy check Stary and Oosterbroek, 1996, and citations below. For Finland check Salmela, 2012b, 2012c, and citations below. For Norway check Olsen et al., 2018, and citations below. For Portugal check Oosterbroek et al., 2020, and citations below. For Great Britain and Ireland check Stubbs, 2021, and citations below. For Morocco check Kettani and Oosterbroek, 2022b, and citations below. Removed, as found in earlier CCW versions: Armenia (excluded for Armenia by Hakobyan and Jenderedjian, 2023).
Map:Podenas et al., 2006 (Switzerland); Boardman, 2007 (Shropshire)
Figure
references:
Habitus: Papp and Schumann, 2000 (male); Quindroit, 2020a (male); Kramer and Morris, 2021a (copula)
Head: Podenas et al., 2006 (antenna)
Wing: Stary, 1968b; Slipka and Stary, 1977b; Podenas et al., 2006; Boardman, 2007
Hypopygium: Stary, 1968b; Podenas et al., 2006
Other figures: Michelsen, 1996 (neck region)

Immature stages
Larva: Podeniene, 2003a; Krivosheina and Krivosheina, 2011c
Citationson biology (mainly from 2000 onward):
Austria
Habitat. Schultz et al., 2006: 351 (Hochmoor).
Habitat. Aistleitner et al., 2024: 32 (loc(s) Vorarlberg, see habitat photo above).
Azerbaijan
Habitat. Stary and Obona, 2020: 130-131 (loc(s); with list of localities, habitats and altitudes).
Belarus
Habitat. Paramonov, 2019a: 96 (loc(s) Berezinsky Biosphere Reserve, distr, habitats, citing various authors) (in Russian).
Czech Rep.
Habitat. Stary et al., 2013b: 378 (on Limonidae from Vraz near Pisek, with general habitat description).
Habitat. Stary and Vonicka, 2018: 71 (on Limoniidae and Pediciidae of two localities in northern Bohemia, with list of localities, altitudes and habitats).
Denmark
Habitat. Nielsen and Nielsen, 2009: 123, 129 (emerging from a forest floor in a Beech [Fagus] stand) (in Danish).
Habitat. Byriel and Rojas, 2017: 118 (among the 101 species in a study on cranefly hotspots in unmanaged and managed forests) (in Danish).
Finland
Habitat. Salmela, 2001a: 143-146, 152 (a common species with one generation a year around springs in southern Finland; (in Europe a very common predator in springs and many other habitats, ranging from aquatic to terrestrial, with two generations a year in Germany and south Sweden, citing various sources)).
Habitat. Salmela, 2004: 3, 4, 9 (eurytopic, ubiquitous species inhabiting a wide spectrum of habitats).
Habitat. Salmela and Ilmonen, 2005: 89-91 (the occurrence in relation to the trophic status of mires within an open boreal mire system is investigated; identified as indicator for mesotrophic sites).
Habitat. Salmela and Vartija, 2007: 10 (very common in wetland habitats, citing Salmela, 2006).
Biology. Salmela, 2008: 12, 20-22, 26, 32, 53 (49) (ecology or habitat: eurytopic, wet, most abundant in rich fens with spring influence and headwater streams with seepages; phenology).
Habitat. Autio and Salmela, 2010: 49, 50 (habitats Aland Is).
Habitat. Salmela, 2011a: 1-28 (study on three wetland habitats: aapa mires, springs and headwater streams; among the most common species).
Habitat. Kramer, 2012f: 3-4 (found at wooded river bank and swept from Carex along river bank).
Habitat. Autio et al., 2013: 1128, 1133 (a wetland generalist, one of the species found in a study on species richness and rarity in a boreal mire).
France
Habitat. Kramer, 2007a: 33 (collected in a marsh dominated by Equisetum).
Habitat. Quindroit, 2020a: 40, 45-48 (annotated list with information on abundancy and habitat of the Tipuloidea of Pays de la Loire: Mayenne (53), Sarthe (72), Loire-Atlantique (44), Maine-et-Loire (49) and Vendee (85)) (in French).
Habitat. Quindroit, 2020b: 133 (loc(s) Nord (59) and Pas-de-Calais (62) with notes on habitats and distr) (in French).
Habitat. Quindroit and Lemoine, 2021a: 24, 25 (loc(s), resultats de deux campagnes de piegeage dans le Pas-de-Calais (62); see paper for details on habitat).
Habitat. Quindroit, 2021a: 36 (loc(s), resultats d une campagne de piegeage dans le Pas-de-Calais (62); see paper for details on habitat).
Georgia
Habitat. Stary and Obona, 2020: 130 (loc(s); with list of localities, habitats and altitudes).
Germany
Habitat. Reusch and Schrankel, 2006: 181 (Quellbereichen; Larven aquatisch, in Quellbereichen, Bachen und kleinen Flussen; Larven zoophag).
Habitat. Reusch, 2006: 134 (Kalkquellmoor [carbonate fen complex]).
Habitat. Lehmann and Reusch, 2009: 122 (from around lowland springs).
Habitat. Reusch and Hohmann, 2009: 130 (from spring brooks).
Great Britain
Habitat. Coulson and Downie, 2000: 30 (areas of blanket bog which had many pools, typical Erica-Sphagnum mire).
Biology. Godfrey, 2001a: 23, 27-28 (habitat; rearing).
Habitat. Rotheray, 2001: 15 (reared from Birch [Betula]).
Habitat. Hancock, 2002: 14 (recorded in this study emerging from water-logged wood).
Habitat. Godfrey, 2003a: 25, 45 (review of association with woody debris; review literature).
Biology. Stubbs, 2003: 18 (habitat: widespread on peaty areas including lowlands; a wet ground species; one of the abundant craneflies as a food source for nesting birds).
Habitat. Boyce, 2004: 33, 62, 78, 104 (predominantly an upland species most commonly encountered on blanket bogs but also found on lowland peatlands).
Habitat. Boardman, 2007: 38 (associated with wet places such as sluggish streams, marshes, bogs and seepages).
Habitat. Wormell, 2007: 150 (mainly along flushes) (as Pedicia).
Biology. Drake, 2011a: 13, 16 26 (among the Diptera from a wet Alder [Alnus] wood in Devon, showing two distinct flight periods in spring and autumn).
Habitat. Kramer, 2011j: 12,025 (common marshland species).
Habitat. Billker, 2017: 46 (Lundy Is, Bristol Channel: cliffs with woodland with towering oaks [Quercus] and a mixed undergrowth with a lot of bluebells [Hyacinthoides non-scripta]; a small stream cuts through the woods, making the habitat an excellent breeding place for Lower Diptera).
Habitat. Brighton, 2017b: 29, 40 (overall regional checklist Lancashire and Cheshire (VC58, 59 and 60) with notes on habitats and/or regional occurrence and/or status; see paper for details).
Phenology. Kramer and Morris, 2021a: 6 (info on the species, on the distr and map of loc(s) Leicestershire and Rutland (VC55; bivoltine, spring and autumn)
Biology. Kramer, 2022b: 12 (drawing attention to the work of Mathew Carroll [2012] from York Uni working on peat uplands; Mathew found that raising the water table by blocking drainage ditches significantly increased cranefly numbers; species identified during his study on moorland in Wales, the Pennines and the North York Moors were Tipula subnodicornis, Tricyphona immaculata, Molophilus ater, Ormosia pseudosimilis, Idioptera pulchella and Limonia dilutior).
Latvia
Habitat. Salmela and Vartija, 2007: 11 (at a small brook surrounded by deciduous trees).
Lithuania
Habitat. Podeniene, 2001a: 84 (larvae were found in leaf litter in Alnus swamp).
Habitat. Podeniene, 2003a: 1-295 (larva, descr, figs, habitat) (in Lithuanian).
Habitat. Podeniene, 2003b: 10, 20-23, 29 (larval habitat: at the bottom of small rivers, sand and silt in riparian zone of small rivulets and littoral zone of lakes, mud (in bogs of low lying woody swamps, on swampy riversides, in littoral zone of swampy lakes), fallings in swampy deciduous forests).
Luxembourg
Biology. Vogtenhuber, 2007: 356 (habitat: carnivoren Larven leben in der Laubstreu feuchter Boden, oft an Bachen; phenologie: zwei Generationen im Jahr).
Morocco
Habitat. Driauach and Belqat, 2016: 142 (loc(s) Rif Mts with list of habitats).
Norway
Habitat. Olsen and Andersen, 2021: 217-218 (loc(s) Innlandet, in former Hedmark, with habitats specified separately, for habitats see also Jonassen and Andersen, 2020).
Poland
Habitat. Chaniecka and Wiedenska, 2006: 149 (occurrence in spring-, springbrook-, and/or bogspring plant communities) (in Polish).
Habitat. Wiedenska, 2014: 15 (on the occurence of Pediciidae in six selected plant communities in the Kamienica Valley, Gorce National Park, Western Carpathians) (in Polish, habitat tables also in English).
Phenology. Wiedenska, 2015a: 47, 54 (on the flight activity of Limoniidae in the Kamienica Valley, Gorce National Park, Western Carpathians; probably two generations a year, with flight diagrams; new correct flight diagrams are given in Wiedenska, 2017b) (in Polish, tables also in English).
Habitat. Wiedenska, 2017a: 28 (faunistic and habitat data of species from the Gorce NP) (in Polish).
Portugal
Habitat. Stary, 2014b: 93 (collected at brooks, spring and marshes).
Romania
Habitat. Parvu, 2004: 187 (spring in forest).
Habitat. Ujvarosi, 2005a: 237, 248 (brook, springs, swamp, moist soil, citing various sources).
Habitat. Ujvarosi et al., 2011b: 110 (loc(s) Dupa Lanca marshy area near Voslobeni) (in Hungarian, species list with habitat information in English).
Habitat. Kolcsar et al., 2013: 73 (loc(s) Cluj-Napoca and surroundings; list of habitats with altitudes).
Russia
Habitat. Przhiboro, 2003: 363 (larvae were common in the water margin zone of an oligotrophic lake; rearing).
Habitat. Przhiboro, 2004: 103, 115 (in Russian).
Habitat. Lantsov in litt., 2014: (found in beech-hornbeam broad-leaved forest with young growth of maple).
Spain
Habitat. Carles-Tolra et al., 2006: 97 (in Spanish).
Habitat. Hancock and Hewitt, 2020: 45-49 (in the Cazorla Parque Natural, Jaen Province, Spain, in June 2019, an area of cliff face was seen covered with a butterwort, Pinguicula vallisneriifolia (Webb), the leaves of which had trapped large numbers of insects, including 19 species of craneflies; the effectiveness of the leaf glandular secretions in relation to immobilising different sizes of flies is discussed; see paper for details, habitat figs).
Habitat. Hancock, 2020: 98 (found attached to leaves of butterwort (Pinguicula vallisneriifolia Webb)) (see also Hancock and Hewitt, 2020).
Sweden
Habitat. Lindstrom and Fritz, 2015: 44, 48, 49 (distributin and ecology in five fens in Hallands County) (in Schwedish).
Ukraine
Habitat. Kolcsar and Torok, 2017: 26 (springs in Spruce [Picea] forest).
on characters and taxonomy (mainly from 2000 onward):
Country not relevant
Phylogeny. Michelsen, 1996: 79, 101 (ingroup taxon in a study titled: Neodiptera: New insights into the adult morphology and higher level phylogeny of Diptera (Insecta); figs).
Phylogeny. Ahonen, 2007: 1-28 (included in phylogenetic analysis of Tipuloidea subfamilies relationships based on molecular data).
Characters. Conrow and Gelhaus, 2022: 138 (among the 45 species examined for wing interference patterns, 13 species were examined in more detail; see paper for details).
Great Britain
Characters. Boardman, 2007: 38 (characters, figs).
Immatures. Reusch, 2007: 872 (larvae in coll. Beling).
Characters. Kramer and Morris, 2021a: 6 (diagnosis, fig(s)).
Lithuania
Immatures. Podeniene, 2003a: 1-295 (larva, descr, figs, habitat) (in Lithuanian).
Portugal
Characters. Ferreira et al., 2021: 11 (barcoding based on material mentioned in Oosterbroek et al. 2020).
on distribution (mainly from 2000 onward):
Andorra
Distribution. Eiroa, 2002b: 65 (checklist).
Armenia
Misinterpretation. Obona et al., 2016: 140 (checklist but excluded for Armenia by Hakobyan and Jenderedjian, 2023).
Austria
Distribution. Schultz et al., 2006: 351 (loc(s)).
Distribution. Reusch and Heiss, 2012: 343-351 (loc(s) Nat. Park Gesause, see PDF for months and altitudes).
Distribution. Vogtenhuber and Kofler, 2017: 142 (loc(s) Osttirol).
Distribution. Aistleitner et al., 2024: 32 (loc(s) Vorarlberg, see habitat photo above).
Azerbaijan
Distribution. Stary and Obona, 2020: 130-131 (loc(s); with list of localities, habitats and altitudes).
Distribution. Snegovaya, 2021: 1154 (checklist with details on distr in Azerbaijan).
Belarus
First record. Paramonov, 2019a: 96 (loc(s) Berezinsky Biosphere Reserve, distr, habitats, citing various authors) (in Russian).
Belgium
Distribution. Martens et al., 2014: 114, 124 (loc(s) Oost-Vlaanderen).
Distribution. Peeters in litt., 2016: (among Malaisetrap material from Jardin Massart, Brussels).
Bosnia-Herzegowina
Distribution. Oosterbroek and Simova-Tosic, 2004: 443 (review literature).
Bulgaria
Distribution. Hubenov, 2015: 222 (annotated list Diptera Pirin Mt, loc(s), faunistics and zoogeography).
Distribution. Hubenov, 2017: 71 (on vertical distribution in the Pirin and Rila Mts; see paper for details on forests and subalpine-alpine zones, distr).
Distribution. Hubenov, 2018a: 17 (on the Diptera of the Vitosha Mt; see paper for details on vegetation belts and local distr).
Distribution. Hubenov, 2021a: 39 (checklist with summary of distr in Bulgaria, references).
Channel Is
Distribution. Stubbs, 2022b: 5 (on a collection of craneflies from Jersey, collected between 1985 and 2002 by Tony Warne).
Croatia
First record. Kolcsar et al., 2015a: 28 (loc(s), distr).
Distribution. Kolcsar et al., 2023b: 33 (loc(s)).
Czech Rep.
Distribution. Stary et al., 2005b: 32-33 (loc(s), the most frequent pediciid species in the studied area).
Distribution. Stary, 2009j: webpage (checklist).
Distribution. Stary et al., 2013b: 378 (on Limonidae from Vraz near Pisek, with general habitat description).
Distribution. Mocek, 2014: 156 (loc(s) NE Bohemia).
Distribution. Stary and Vonicka, 2018: 71 (on Limoniidae and Pediciidae of two localities in northern Bohemia, with list of localities, altitudes and habitats).
Denmark
Distribution. Petersen et al., 2001: 157 (checklist).
Distribution. Nielsen and Nielsen, 2009: 123 (loc(s)).
Estonia
First record. Salmela in litt., 2003: (new for Estonia).
Finland
Distribution. Polevoi, 2001: 191 (loc(s)).
Distribution. Salmela, 2001a: 143-146, 152 (loc(s)).
Distribution. Salmela, 2002: 30, 31 (loc(s)).
Distribution. Salmela, 2004: 3, 4, 9 (loc(s)).
Distribution. Salmela and Ilmonen, 2005: 89-91 (loc(s)).
Distribution. Salmela and Autio, 2007b: 54 (loc(s)).
Distribution. Salmela et al., 2007b: 37, 46 (abundance, loc(s)).
Distribution. Salmela, 2008: 12, 32, 53 (49) (loc(s), known from all Finnish ecoregions).
Distribution. Autio and Salmela, 2010: 49 (loc(s) Aland Is).
Distribution. Salmela, 2010a: 8, 13 (survey fauna Tahmela spring in city of Tampere) (in Finnish).
Distribution. Salmela, 2011a: 24 (loc(s) subalpine northern boreal Finland).
Distribution. Kramer, 2012f: 3-4 (loc(s) Obb: Syote National Park).
Distribution. Salmela, 2012b: 242 (annotated list of Finnish crane flies).
Distribution. Salmela, 2013: 18 (belongs to the most abundant and frequent cranefly species of Finland, see table above under Habitat).
Distribution. Salmela and Petrasiunas, 2014: 23 (checklist Finnish Tipulomorpha).
Distribution. Salmela et al., 2015: 109 (list of localities in extensive report on malaise-trapping of insects in conservation areas in Lapland in 2012–2014) (in Finnish).
France
Distribution. Kramer, 2007a: 33 (loc(s) Isere (38)).
Distribution. Kramer and Langlois, 2019a: 76 (record(s) Ravin de Valbois, Doubs (25)).
Distribution. Quindroit, 2020a: 40, 45-48 (annotated list with information on abundancy and habitat of the Tipuloidea of Pays de la Loire: Mayenne (53), Sarthe (72), Loire-Atlantique (44), Maine-et-Loire (49) and Vendee (85)) (in French).
Distribution. Quindroit, 2020b: 133 (loc(s) Nord (59) and Pas-de-Calais (62) with notes on habitats and distr) (in French).
Distribution. Quindroit and Lemoine, 2021a: 24, 25 (loc(s), resultats de deux campagnes de piegeage dans le Pas-de-Calais (62); see paper for details on habitat).
Distribution. Quindroit, 2021a: 36 (loc(s), resultats d une campagne de piegeage dans le Pas-de-Calais (62); see paper for details on habitat).
Distribution. Tissot et al., 2021: 210 (the Diptera of the Reserves Naturelles Nationales du Doubs (25), results of Malaise trapping since 2009) (in French).
Distribution. Quindroit, 2022a: 25-28 (additions to the annotated list of the Tipuloidea of Pays de la Loire as given in Quindroit, 2020a).
Georgia
Distribution. Stary and Obona, 2020: 130 (loc(s); with list of localities, habitats and altitudes).
Germany
Distribution. Schacht, 1999: 134 (checklist Bayern).
Distribution. Reusch and Oosterbroek, 2000: 164 (checklist German Bundeslander).
Distribution. Reusch and Bellstedt, 2001b: 76 (checklist Thuringen).
Distribution. Schacht, 2005: 4 (checklist Bayern).
Distribution. Reusch and Schrankel, 2006: 181 (loc(s) Bayern, Nationalpark Berchtesgaden, distr).
Distribution. Reusch, 2006: 134 (loc(s)).
Distribution. Lehmann and Reusch, 2009: 121-122 (loc(s) Niedersachsen).
Distribution. Reusch and Hohmann, 2009: 130 (loc(s) Sachsen-Anhalt).
Distribution. Hable et al., 2010b: 90 (loc(s) Bayern).
Distribution. Schacht, 2010: 18 (checklist Diptera Bayern).
Distribution. Dunk and Kraus, 2014: 169 (loc(s) Bayern).
Distribution. Stuke, 2019: 163 (checklist Niedersachsen and Bremen with reference to original sources).
Great Britain
Distribution. Coulson and Downie, 2000: 30 (loc(s)).
Distribution. Godfrey, 2001a: 23, 27-28 (loc(s)).
Distribution. Rotheray, 2001: 1, 14 (loc(s) Scotland).
Distribution. Hancock, 2002: 13 (loc(s) Scotland).
Distribution. Godfrey, 2003a: 25, 45 (loc(s), distr, review literature).
Distribution. Boyce, 2004: 33, 62, 78, 104 (distr).
Distribution. Corbet, 2004: 128 (loc(s)).
Distribution. Hewitt et al., 2005: 37 (loc(s) Cumbria) (as Pedicia).
Distribution. Boardman, 2007: 38 (distr, map).
Distribution. Wormell, 2007: 150 (loc(s) Isle of Rum) (as Pedicia).
Distribution. Skidmore, 2009: 125 (review of occurrence on the Western Isles of Scotland, distr).
Distribution. Drake, 2011a: 26 (loc(s) Devon).
Distribution. Kramer, 2011g: 2 (loc(s) Devon).
Distribution. Kramer, 2011j: 12 (loc(s) Leicestershire and Rutland (VC55)).
Distribution. Kramer, 2013i: 8 (checklist of species from Leigh Woods NNR, Bristol district).
Distribution. Hewitt, 2014: (provisional checklist Cumbrian Diptera).
Distribution. Kramer, 2014c: 4 (in table of cranefly species recorded at Shapwick Heath NNR, as given in H. Audcent, 1949 and/or collected by J. Kramer in 2013).
Distribution. Wolton et al., 2014: 26 (among Diptera associated with a hedge in Devon, caught in emergence trap).
Distribution. Billker, 2017: 46 (loc(s) Lundy Is, Bristol Channel).
Distribution. Brighton, 2017b: 29, 40 (overall regional checklist Lancashire and Cheshire (VC58, 59 and 60) with notes on habitats and/or regional occurrence and/or status; see paper for details).
Distribution. Showers, 2017a: 4 (loc(s) Northamptonshire).
Distribution. Kramer and Morris, 2021a: 6 (info on the species, on the distr and map of loc(s) Leicestershire and Rutland (VC55; bivoltine, spring and autumn)
Distribution. Kramer, 2022b: 12 (drawing attention to the work of Mathew Carroll [2012] from York Uni working on peat uplands; Mathew found that raising the water table by blocking drainage ditches significantly increased cranefly numbers; species identified during his study on moorland in Wales, the Pennines and the North York Moors were Tipula subnodicornis, Tricyphona immaculata, Molophilus ater, Ormosia pseudosimilis, Idioptera pulchella and Limonia dilutior).
Hungary
Distribution. Kolcsar and Keresztes, 2016: 123 (loc(s), distr).
Ireland
Distribution. Ashe et al., 2008: 15 (additional Irish record(s), distr).
Distribution. Chandler et al., 2008: 8 (checklist).
Italy
Distribution. Podenas and Podeniene, 2008: 350, 353 (loc(s), annotated checklist South Tyrol [Prov. Bolzano]).
Kazakhstan
Distribution. Devyatkov, 2021d: 337 (loc(s) East Kazakhstan, distr) (in Russian).
Latvia
First record. Stary, 2004g: (Fauna Europaea: added Latvia).
Distribution. Salmela and Vartija, 2007: 11 (loc(s)).
Lithuania
Distribution. Podeniene, 2001a: 84 (loc(s)).
Distribution. Pakalniskis et al., 2006: 25 (checklist).
Luxembourg
First record. Vogtenhuber, 2007: 356 (loc(s)).
Montenegro
Distribution. Oosterbroek and Simova-Tosic, 2004: 443 (review literature).
Morocco
First record. Driauach and Belqat, 2016: 142 (loc(s) Rif Mts with list of habitats).
Distribution. Kettani and Oosterbroek, 2022b: 28 (annotated checklist, with review loc(s) and/or references).
Netherlands
Distribution. de Jong and Oosterbroek, 2002c: 37 (checklist).
Distribution. Oosterbroek and de Jong, 2017b: 238 (loc(s) Zeeland) (in Dutch).
North Macedonia
Distribution. Oosterbroek and Simova-Tosic, 2004: 443 (review literature).
Norway
Distribution. Olsen et al., 2018: 152-153 (annotated checklist Nordic countries and for Norway according to the Strand regions).
Distribution. Olsen and Andersen, 2021: 217-218 (loc(s) Innlandet, in former Hedmark, with habitats specified separately, for habitats see also Jonassen and Andersen, 2020).
Poland
Distribution. Palaczyk, 2001: 269 (loc(s)).
Distribution. Chaniecka and Wiedenska, 2006: 149 (loc(s)).
Distribution. Wiedenska, 2007b: 77 (checklist).
Distribution. Wiedenska, 2008: 47 (loc(s) Parowy Janinowski nature reserve near Lodz) (in Polish).
Distribution. Wiedenska, 2010: 136 (loc(s) Parku Krajobrazowego Wzniesien Lodzkich, Lodz).
Distribution. Wiedenska, 2014: 15 (on the occurence of Pediciidae in six selected plant communities in the Kamienica Valley, Gorce National Park, Western Carpathians) (in Polish, habitat tables also in English).
Distribution. Wiedenska, 2017a: 28 (faunistic and habitat data of species from the Gorce NP) (in Polish).
Distribution. Obona et al., 2019a: 125 (records of Diptera from the Babia Gora Massif).
Distribution. Wiedenska and Syratt, 2023: 153 (loc(s) Bieszczady Mountains, distr and remarks).
Portugal
First record. Stary, 2014b: 93 (loc(s) Faro).
Distribution. Oosterbroek et al., 2020: 336 (loc(s) Guarda, also known from Faro, distr).
Romania
Distribution. Ujvarosi and Stary, 2003: 49 (loc(s), checklist).
Distribution. Parvu, 2004: 187 (loc(s)).
Distribution. Parvu, 2007: 234 (loc(s) Maramures Depression).
Distribution. Ujvarosi, 2007: 231-232 (checklist Pediciidae).
Distribution. Ujvarosi et al., 2011b: 110 (loc(s) Dupa Lanca marshy area near Voslobeni) (in Hungarian, species list with habitat information in English).
Distribution. Kolcsar et al., 2012: 25 (checklist Romania).
Distribution. Kolcsar et al., 2013: 73 (loc(s) Cluj-Napoca and surroundings; list of habitats with altitudes).
Russia
Distribution. Przhiboro, 2003: 363 (loc(s) NW Russia).
Distribution. Przhiboro, 2004: 103, 115 (loc(s) RUW: Leningradskaya oblast) (in Russian).
Distribution. Humala and Polevoi, 2009: 66 (loc(s) RUN: Kareliya) (in Russian).
Distribution. Gavryushin in litt., 2011e: (loc(s) RUN: Murmanskaya oblast).
Distribution. Lantsov in litt., 2014: (North Caucasus, no further locality details presented).
Distribution. Gavryushin in litt., 2015a: (loc(s) RUE: Bashkortostan Rep.).
Distribution. Lantsov, 2015a: 366, 367 (North Caucasus: loc(s) Karachay-Cherkess Rep.).
Distribution. Lantsov, 2016: 165-170 (on craneflies from Arkhyz, Karachay-Cherkessia Rep., North Caucasus) (in Russian).
Distribution. Paramonov and Pilipenko, 2016: 100 (loc(s) RUC: Tverskaya oblast) (in Russian).
Distribution. Lantsov, 2020: 293 (North Caucasus: found in the plains of Dagestan).
Distribution. Pilipenko et al., 2020: 362 (loc(s) RUC: Mordoviya Rep., distr).
Serbia
Distribution. Oosterbroek and Simova-Tosic, 2004: 443 (review literature).
Distribution. Gavryushin in litt., 2015b: (loc(s) Serbia).
Slovakia
Distribution. Stary, 2009e: 41 (loc(s) Polana area).
Distribution. Stary, 2009j: webpage (checklist).
Slovenia
Distribution. Oosterbroek and Simova-Tosic, 2004: 443 (review literature).
Spain
Distribution. Eiroa, 2002b: 65 (checklist).
Distribution. Carles-Tolra et al., 2006: 97 (loc(s)).
Distribution. Mederos and Eiroa, 2015: 267 (loc(s) Lerida).
Distribution. Hancock and Hewitt, 2020: 45-49 (in the Cazorla Parque Natural, Jaen Province, Spain, in June 2019, an area of cliff face was seen covered with a butterwort, Pinguicula vallisneriifolia (Webb), the leaves of which had trapped large numbers of insects, including 19 species of craneflies; the effectiveness of the leaf glandular secretions in relation to immobilising different sizes of flies is discussed; see paper for details, habitat figs).
Distribution. Hancock, 2020: 98 (loc(s) Jaen, Cazorla National Park).
Distribution. Keresztes et al., 2022: 196 (loc(s) Asturias and review distribution in Spain) (in Spanish).
Sweden
Distribution. Salmela, 2010b: 129, 134 (loc(s) Malmo, distr).
Distribution. Fritz and Lindstrom, 2013: 63 (loc(s) Hallands County) (in Schwedish).
Distribution. Lindstrom and Fritz, 2015: 44, 48, 49 (distributin and ecology in five fens in Hallands County) (in Schwedish).
Switzerland
Distribution. Podenas et al., 2006: 189, 358 (map, vertical distr per thermic level).
Distribution. Dufour and Merz, 2012: 355 (annotated checklist canton Geneva).
Turkey
First record. Stary and Oosterbroek, 2008: 18 (loc(s), distr).
Distribution. Ozgul and Koc, 2016: 781-782 (loc(s) various provinces in NW Turkey, checklist Turkish Pedicidae with provinces).
Distribution. Provinces Asiatic part, 2024: (known from Afyonkarahisar, Bursa, Canakkale, Erzurum, Eskisehir).
Ukraine
Distribution. Kolcsar and Torok, 2017: 26 (loc(s) Hoverla).
on flight period (mainly from 2000 onward):
Austria
Vogtenhuber and Kofler, 2017: month(s): 7.
Aistleitner et al., 2024: month(s): 7.
Azerbaijan
Stary and Obona, 2020: month(s): 5, 9-10.
Belarus
Paramonov, 2019a: month(s): 9.
Belgium
Martens et al., 2014: month(s): 4-6.
Bulgaria
Hubenov, 2015: month(s): 5-6, 8.
Channel Is
Stubbs, 2022b: month(s): 6.
Croatia
Kolcsar et al., 2015a: month(s): 4-5, 10.
Kolcsar et al., 2023b: month(s): 1, 9-10.
Czech Rep.
Stary et al., 2005b: month(s): 3-6, 9.
Stary and Vonicka, 2018: month(s): 5-9(10).
Finland
Salmela, 2001a: month(s): 5-6.
Salmela, 2008: month(s): 6-8.
Kramer, 2012f: month(s): 6.
France
Kramer, 2007a: month(s): 6.
Quindroit, 2020a: month(s): 4-5, 9.
Quindroit, 2020b: month(s): 4-5, 9-10.
Georgia
Stary and Obona, 2020: month(s): 4-5, 7, 9.
Germany
Reusch and Schrankel, 2006: month(s): 5-6.
Great Britain
Godfrey, 2001a: month(s): 5.
Hancock, 2002: month(s): 5.
Corbet, 2004: month(s): 9.
Wormell, 2007: month(s): 8.
Kramer, 2011j: month(s): 5-6.
Kramer, 2014c: month(s): 5.
Kramer and Morris, 2021a: month(s): 4-6, 9-10(11).
Hungary
Kolcsar and Keresztes, 2016: month(s): 5-6, 8-10.
Ireland
Ashe et al., 2008: month(s): 5-6, 8-9.
Italy
Podenas and Podeniene, 2008: month(s): 5-6.
Kazakhstan
Devyatkov, 2021d: month(s): 5-8.
Latvia
Salmela and Vartija, 2007: month(s): 5.
Morocco
Driauach and Belqat, 2016: month(s): 4-5.
Norway
Olsen and Andersen, 2021: month(s): (4-)6-9.
Poland
Wiedenska, 2008: month(s): 5.
Wiedenska, 2014: month(s): 5-10.
Wiedenska, 2015a: month(s): 5, 7-9.
Wiedenska, 2017a: month(s): 5-6, 8.
Obona et al., 2019a: month(s): 6.
Wiedenska and Syratt, 2023: month(s): 5-7, 10.
Portugal
Stary, 2014b: month(s): 4-5.
Oosterbroek et al., 2020: month(s): 5.
Romania
Parvu, 2004: month(s): 6.
Parvu, 2007: month(s): 5.
Ujvarosi et al., 2011b: month(s): 5-6.
Kolcsar et al., 2013: month(s): 5-6, 9.
Russia
Przhiboro, 2003: month(s): 6-7.
Gavryushin in litt., 2011e: month(s): 7.
Gavryushin in litt., 2015a: month(s): 7.
Lantsov, 2015a: month(s): 6-7.
Paramonov and Pilipenko, 2016: month(s): 5-6, 8-9.
Pilipenko et al., 2020: month(s): 5, 9.
Serbia
Gavryushin in litt., 2015b: month(s): 5, 9.
Slovakia
Stary, 2009e: month(s): 5-7, 10.
Spain
Carles-Tolra et al., 2006: month(s): 4.
Hancock and Hewitt, 2020: month(s): 6.
Hancock, 2020: month(s): 6.
Keresztes et al., 2022: month(s): 6.
Switzerland
Podenas et al., 2006: month(s): 4-11.
Turkey
Stary and Oosterbroek, 2008: month(s): 6.
Ozgul and Koc, 2016: month(s): 4-6.
Ukraine
Kolcsar and Torok, 2017: month(s): 7.
on altitude (mainly from 2000 onward):
Austria
Vogtenhuber and Kofler, 2017: altitude: 930 m.
Aistleitner et al., 2024: altitude: 2020 m.
Bulgaria
Hubenov, 2015: altitude: 1900-2200 m.
Hubenov, 2017: altitude: 1900-2200 m.
Hubenov, 2021a: altitude: 1200-2200 m.
Croatia
Kolcsar et al., 2015a: altitude: 630-720 m.
Kolcsar et al., 2023b: altitude: 630-720 m.
France
Kramer, 2007a: altitude: 1025 m.
Germany
Reusch and Schrankel, 2006: altitude: 1100-1250 m.
Reusch and Hohmann, 2009: altitude: 375-842 m.
Italy
Podenas and Podeniene, 2008: altitude: 1220 m.
Kazakhstan
Devyatkov, 2021d: altitude: 386-1468 m.
Morocco
Driauach and Belqat, 2016: altitude: 966-1674 m.
Poland
Wiedenska, 2017a: altitude: 680-1230 m.
Obona et al., 2019a: altitude: 1001 m.
Wiedenska and Syratt, 2023: altitude: 600-1000 m.
Portugal
Stary, 2014b: altitude: 450-870 m.
Romania
Ujvarosi and Stary, 2003: altitude: 450-1250 m.
Russia
Lantsov in litt., 2014: altitude: 1048 m.
Gavryushin in litt., 2015a: altitude: 607 m.
Lantsov, 2015a: altitude: 1048-1342 m.
Serbia
Gavryushin in litt., 2015b: altitude: 1030-1500 m.
Slovakia
Stary, 2009e: altitude: 560-870 m.
Spain
Carles-Tolra et al., 2006: altitude: 100 m.
Mederos and Eiroa, 2015: altitude: 130 m.
Keresztes et al., 2022: altitude: 1485-1690 m.
Turkey
Stary and Oosterbroek, 2008: altitude: 2000 m.
Ozgul and Koc, 2016: altitude: 430-1701 m.
Ukraine
Kolcsar and Torok, 2017: altitude: 1180-1800 m.

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