Details for Metalimnobia (Metalimnobia) quadrimaculata

Name: Metalimnobia (Metalimnobia) quadrimaculata (Linnaeus, 1760) Publication: Fauna Sveciae, Ed. 2: 430 (as Tipula) Status: Recognized taxon, Synonym(s): calmariensis (Linnaeus, 1760); quatuormaculata (Meigen, 1804); annulus (Meigen, 1818); variegata (Macquart, 1826); ?truncata (Alexander, 1924). Classification: Family Limoniidae Subfamily Limoniinae Descriptive note: Synonymy: Linnaeus, 1767. Synonymy of variagata after Quindroit, 2025a. Usually cited as 1761. Keys: Podenas and Gelhaus, 2007 (Limoniinae Mongolia and nearby); Krivosheina and Krivosheina, 2010 (Metalimnobia larvae (5 species) and pupae (3 species); Mao and Yang, 2010a (Metalimnobia China); Peeters en Oosterbroek, 2014a (craneflies Netherlands, Belgium and Luxembourg) (in Dutch); Podenas and Byun, 2016 (Metalimnobia Korea); Stubbs and Kramer, 2016h; Stubbs, 2021 (British craneflies); Jiang and Zhang, 2020 (Metalimnobia China) habitus male habitus male habitus male habitus male habitus male plus 41 more images of habitus, habitus, habitus, habitus, habitus, habitus, habitus, habitus, body part(s), hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, ovipositor, ovipositor, ovipositor, ovipositor, ovipositor, wing, wing, wing, wing, wing, wing, wing, habitat, habitat, habitat, habitat, habitat, habitat, habitat, habitat and habitat. Distribution: USA (Pa);; Albania, Austria, Belarus, Belgium, Bosnia-Herzegovina, Czech Rep., Denmark, Finland, France, Germany, Great Britain, Hungary, Italy, Latvia, Lithuania, Luxembourg, Montenegro, Netherlands, Norway, Poland, Romania, Slovakia, Spain, Sweden, Switzerland, Turkey (European part: Istanbul, Kirklareli), Ukraine; Russia: RUN, RUW, RUC, RUE, RUS, North Caucasus; Georgia, Azerbaijan, Turkey (Asiatic part: Bolu, Kocaeli, Zonguldak);; Russia: WS (south), ES (south), FE (Kamchatka, Amurskaya oblast, Khabarovskiy kray, Primorskiy kray, Sakhalin (incl. Kuril Is)); Kazakhstan (east); Mongolia, North Korea, South Korea, Japan (Hokkaido, Honshu), China (Hebei, Henan). Note: see the manual for abbreviations: Canada and USA; present-day Russia. Region(s): Nearctic;; Westpalaearctic;; Eastpalaearctic General note: Added: USA (Pa) (Young and Gelhaus, 2000). For Italy check Stary and Oosterbroek, 1996, and citations below. For Finland check Salmela, 2012b, 2012c, and citations below. For Norway check Olsen et al., 2018, and citations below. For Spain check Eiroa and Carles-Tolra, 2019, and citations below. For Great Britain check Stubbs, 2021, and citations below. Removed, as found in earlier CCW versions: Armenia (excluded for Armenia by Hakobyan and Jenderedjian, 2023). Map: Koc et al., 2005b (NW Turkey); Podenas et al., 2006 (Switzerland); Podenas and Byun, 2016 (Korea) Biology: Alexander, 1919e (habitat). Yakovlev and Polevoi, 1997 (list of Fungi). Young and Gelhaus, 2000 (habitat, phenology) (all as annulus Meigen). Figure references: Habitus: Savchenko, 1985b; Kinota, 2006 (male, as subspecies truncata); Nakamura, 2006b (male); Podenas and Gelhaus, 2007 Head: Jo, 2012; Jo and Kong, 2014 Wing: Alexander, 1972g; Podenas et al., 2006; Podenas and Gelhaus, 2007; Salmela and Stary, 2009; Mao and Yang, 2010a; Jo, 2012; Jo and Kong, 2014; Podenas and Byun, 2016 Hypopygium: Podenas et al., 2006; Podenas and Gelhaus, 2007; Salmela and Stary, 2009; Mao and Yang, 2010a; Podenas and Byun, 2016; Jiang and Zhang, 2020 Ovipositor: Savchenko, 1985b; Podenas and Gelhaus, 2007; Jo, 2012; Jo and Kong, 2014; Podenas and Byun, 2016 Immature stages Larva: Podeniene, 2003a; Krivosheina and Krivosheina, 2010; Krivosheina and Krivosheina, 2011c Pupa: Skidmore, 2003; Krivosheina and Krivosheina, 2010 Citations on biology (mainly from 2000 onward): Albania Habitat. Kolcsar et al., 2021b: 152 (Beech [Fagus] forest, next to stream). Belarus Habitat. Paramonov, 2019a: 95 (loc(s) Berezinsky Biosphere Reserve, distr, habitats, citing various authors) (in Russian) (first published record for Belarus). Belgium Habitat. Tack et al., 2021: 149 (loc(s) Antwerpen, alle taxa biodiversiteit inventarisatie Bos t Ename, op paddestoelen op/in dood hout in nat bos (in Dutch), see paper for description). Country not relevant Habitat. Sevcik, 2003: 161 (associated with the wood-decaying fungus Abortiporus biennis, Polyporaceae, citing Sevcik, 2001). Habitat. Krivosheina and Krivosheina, 2011c: 1-294 (larva: loc(s), descr, figs, biology, key) (in Russian, key also in English). Biology. Gudin et al., 2022: 399 (paper includes an updated catalogue of dipteran hosts of Tachinidae with list of the tachinid species involved; of the host records from Diptera, most are from larvae of Tabanidae and Tipulidae, with a few occasional records in other families). Czech Rep. Habitat. Sevcik, 2006: 8 (rearing records; possibly associated with Bjerkandera adusta, but it was reared also from other species of fungi, as well as from decaying wood (Jakovlev 1994), probably due to the pupation of larvae there when the fungus has decomposed). Biology. Sevcik, 2010: 10 (on Czech and Slovak Diptera associated with fungi; review of published records; new records; summary: this limoniid species apparently prefers the soft polypore Bjerkandera adusta, but it was reared also from other species of fungi, as well as from decaying wood (Jakovlev 1994), probably due to the pupation of larvae there when the fungus has decomposed. Habitat. Stary et al., 2013b: 377 (on Limonidae from Vraz near Pisek, with general habitat description). Habitat. Stary and Vonicka, 2018: 67 (on Limoniidae and Pediciidae of two localities in northern Bohemia, with list of localities, altitudes and habitats). Denmark Habitat. Byriel and Rojas, 2017: 118 (among the 101 species in a study on cranefly hotspots in unmanaged and managed forests) (in Danish). Finland Phenology. Salmela, 2001a: 143, 144, 151 (obviously two generations per year in southern Finland). Habitat. Salmela and Stary, 2009: 271 (adults were obtained from trunk-emergence traps on Betula logs lying on the forest floor and decayed by polyporous fungi). Habitat. Halme et al., 2012: 507 (part of a study on saproxylic nematoceran communities occupying different parts of decaying fallen Aspen trunks in a boreal forest in central Finland). Habitat. Salmela, 2012c: 4 (strictly fungivorous). Habitat. Rasimus, 2018: 17 (on saproxylic craneflies in the Tampere region) (in Finnish). Habitat. Rasimus, 2018: 17 (on saproxylic craneflies in the Tampere region) (in Finnish). France Habitat. Kramer and Langlois, 2019a: 83 (on larval fungal habitats in the Ravin de Valbois; see paper for details). Biology. Quindroit, 2020a: 39, 45-48 (annotated list with information on abundancy and habitat of the Tipuloidea of the Pays de la Loire regoin, distr) (in French). Habitat. Quindroit, 2020b: 139 (loc(s) Nord (59) and Pas-de-Calais (62) with notes on habitats and distr) (in French). Habitat. Quindroit and Lemoine, 2021b: 7 (loc(s) Nord (59); see paper for habitat description) (in French). Habitat. Quindroit, 2021c: 424, 426 (loc(s) Mercantour National Park, Alpes-Maritimes (06), wide distribution through France; infrequent, usually from old forests; mycophagous). Habitat. Quindroit and Lemoine, 2022a: 25 (loc(s) Aisne (02) with notes on habitats and distr) (in French). Georgia Habitat. Stary and Obona, 2020: 137 (loc(s); with list of localities, habitats and altitudes). Germany Habitat. Dunk, 2018: 80, 96 (loc(s) Bayern with list of habtats). Great Britain Habitat. Alexander, 2002: 90 (broadleaved woodland; larvae in a wide range of bracket fungi on trees). Habitat. Skidmore, 2003: 23-24, 27 (larvae in Ganoderma on Fagus). Biology. Stubbs, 2003: 68, 77-78 (habitat: only known from forests where good stands of veteran Fagus occur; the larvae live in large bract fungi, apparently only Inonotus hispidus and Polyporus schweinitzii, growing on such trees; adults have been recorded in april-june and also in august suggesting two generations a year but this needs to be confirmed ; the paper includes further details on monitoring techniques and examples of good/bad practice). Habitat. Chandler, 2010c: 431 (overview of known fungus associations). Habitat. Chandler, 2017: 92 (one male in decayed hardwood of Beech [Fagus]; first reared fromWindsor by O.W. Richards in 1932, latest Windsor record in 2014; develops in bracket fungi, only otherwise known in Britain from Epping and New Forests). Habitat. Donisthorpe, 2017a: 92 (male in decayed heartwood of Beech [Fagus]; develops in bracket fungi). Biology. Brock, 2018: 5 (observations in the New Forest on the gathering of adults on Beech [Fagus] and Oak [Quercus] and association with Inotus bracket fungi; see paper for details). Italy Habitat. Stary, 2003a: 125. Japan Biology. Sueyoshi et al., 2007: 247-257 (loc(s) Honshu; on host abundance of limoniid flies of Agaricales (gill mushrooms) and Aphyllophorales (non-gilled fungi) and the importance of these flies as phoronts of mites inhabiting these fungal orders; see paper for details on fungal species and forest types). Lithuania Habitat. Podeniene, 2003a: 1-295 (larva, descr, figs, habitat) (in Lithuanian). Habitat. Podeniene, 2003b: 11, 23, 24 (larval habitat: uppermost layer of soil in old deciduous forest rich in epiphytes, mushrooms). Habitat. Podeniene et al., 2010: 234-237 (list of fungi in which larvae develop; habitat: deciduous swamp forest and mixed forest). Biology. Gorban and Podeniene, 2022: 5, 8-9 (loc(s); reared from Poplar [Populus tremula], mycetophagous species; see paper for details). Luxembourg Biology. Vogtenhuber, 2007: 355 (habitat: Larvenentwicklung in Basisiomyceten; phenology: die Phanologie der Imagines korreliert mit den Pilzen, die ersten Imagines in Juni, die letzten Anfang Oktober). Mongolia Habitat. Yadamsuren et al., 2015: 475 (fungi and soil). Habitat. Oliveira and Eck, 2025: Specieslist (collected at a riverside vegetation, and in forest steppe (?), see habitat photos). Norway Habitat. Olsen and Andersen, 2021: 216 (loc(s) Innlandet, in former Hedmark, with habitats specified separately, for habitats see also Jonassen and Andersen, 2020). Poland Habitat. Wiedenska, 2007a: 76 (mycophagous, feeding on fungi). Habitat. Wiedenska, 2017a: 24 (faunistic and habitat data of species from the Gorce NP) (in Polish). Romania Habitat. Ujvarosi, 2005a: 246 (fleshy fungi, citing various sources). Habitat. Ujvarosi et al., 2011b: 109 (loc(s) Dupa Lanca marshy area near Voslobeni) (in Hungarian, species list with habitat information in English). Russia Habitat. Krivosheina, N.P., 2008: 781 (larvae are believed to develop in the fruit bodies of some polypores and the saddle fungus Gyromitra (cting various authors); they typically occur in sap concretions on trunks and stumps, in the brown dust under the bark of stumps, and in the decomposed wood of fallen trees, they were also found in the humid decomposed fruit bodies of Laetiporus sulphureus and in the loose old carpophores of Fomes fomentarius). Habitat. Krivosheina, N.P., 2009a: 125-133 (review Russian xylophilous Limoniidae, mainly inhabiting bark and wood of decomposing trees, with new original data on larval biology) (in Russian). Habitat. Krivosheina and Krivosheina, 2010: 777-779 (from brown rot of a Birch [Betula] knot and from fungi; the paper present further details on larval and pupal development, literature review). Habitat. Paramonov, 2016: 131 (from area with old Salix caprea trees) (in Russian). Biology. Tschorsnig, 2017: 59 (host of Palaearctic Tachinidae; for species of Tachinidae, localities and references see paper). Habitat. Polevoi et al., 2018: supplementary species list: emerged from fallen Aspens [Populus tremula] in Kivach Nature Reserve, Kareliya). Habitat. Dvorak et al., 2020: 6 (on Diptera collected in beer traps in the RUC: Mordoviya Rep.; collected in Pine [Pinus], floodplain, deciduous and mixed forests). Habitat. Dvorakova et al., 2020: 14-15 (on Diptera collected in beer traps in the western Ukraine; collected in a deciduous forest). Habitat. Kolcsar et al., 2021b: 153 (Tatarstan Rep.: funnel karst origin). Habitat. Dvorak et al., 2022a: 134 (on Diptera collected in beer traps in the Volga Region, M. quadrimaculata being collected in Pine [Pinus], decidous and mixed forests). Slovakia Biology. Sevcik, 2010: 10 (on Czech and Slovak Diptera associated with fungi; review of published records; new records; summary: this limoniid species apparently prefers the soft polypore Bjerkandera adusta, but it was reared also from other species of fungi, as well as from decaying wood (Jakovlev 1994), probably due to the pupation of larvae there when the fungus has decomposed. Biology. Obona et al., 2025a: 30 (collected in a mosquito trap at the Kosice Zoo, perhaps because of being attracted to CO2). South Korea Habitat. Podenas and Byun, 2016: 340 (adults were collected in wet deciduous and mixed forests, usually not far from streams). on characters and taxonomy (mainly from 2000 onward): China Revision. Mao and Yang, 2010a: 2-3, 6-8, 10-11 (descr, comparison, key Chinese Metalimnobia, figs). Country not relevant Characters. Frantsevich, 2004: 133 (structure of coxa). Characters. Salmela and Stary, 2009: 269-273 (comparison, figs). Immatures. Krivosheina and Krivosheina, 2011c: 1-294 (larva: loc(s), descr, figs, biology, key) (in Russian, key also in English). Description. Podenas and Byun, 2016: 334-339, 343, 345 (short descr, key, comparison, figs). Characters. Jiang and Zhang, 2020: 103 (comparison). Characters. Kato, 2020: 39 (comparison). Great Britain Immatures. Skidmore, 2003: 23-24, 27 (fig pupa). Lithuania Immatures. Podeniene, 2003a: 1-295 (larva, descr, figs, habitat) (in Lithuanian). Immatures. Podeniene et al., 2010: 237-240 (key to crane fly larvae developing in mushrooms, figs). Mongolia Key. Podenas and Gelhaus, 2007: 73-75 (key, figs). Russia Immatures. Krivosheina and Krivosheina, 2010: 765, 777-781 (descr, key, figs, larva, pupa). Characters. Belyaev and Farisenkov, 2019: 8 (used in a study on allometry of wing shape and venation in Diptera, material was collected in RUC: Moskovskaya oblast). South Korea Description. Jo, 2012: 25-26 (descr, figs) (in Korean; more or less repeated, in English, in Jo and Kong, 2014). on distribution (mainly from 2000 onward): Albania First record. Kolcsar et al., 2021b: 152 (loc(s)). Armenia Misinterpretation. Obona et al., 2016: 140 (checklist but excluded for Armenia by Hakobyan and Jenderedjian, 2023). Austria Distribution. Reusch and Heiss, 2012: 323-332 (loc(s) Nat. Park Gesause, see PDF for months and altitudes). Distribution. Vogtenhuber and Kofler, 2017: 140 (loc(s) Osttirol). Azerbaijan Distribution. Snegovaya, 2021: 1140 (checklist with details on distr in Azerbaijan). Belarus First record. Gavryushin in litt., 2013d: (loc(s) Minsk region). Distribution. Paramonov, 2019a: 95 (loc(s) Berezinsky Biosphere Reserve, distr, habitats, citing various authors) (in Russian) (first published record for Belarus). Belgium Distribution. Tack et al., 2021: 149 (loc(s) Antwerpen, alle taxa biodiversiteit inventarisatie Bos t Ename, op paddestoelen op/in dood hout in nat bos (in Dutch), see paper for description). Bosnia-Herzegowina Distribution. Oosterbroek and Simova-Tosic, 2004: 448 (review literature). China First record. Mao and Yang, 2010a: 8 (loc(s) Hebei, Henan). Country not relevant Distribution. Krivosheina and Krivosheina, 2011c: 1-294 (larva: loc(s), descr, figs, biology, key) (in Russian, key also in English). Czech Rep. Distribution. Stary et al., 2005a: 28 (loc(s)). Distribution. Sevcik, 2006: 8 (loc(s)). Distribution. Sevcik, 2006: 8 (rearing records; possibly associated with Bjerkandera adusta, but it was reared also from other species of fungi, as well as from decaying wood (Jakovlev 1994), probably due to the pupation of larvae there when the fungus has decomposed). Distribution. Stary, 2009i: webpage (checklist). Distribution. Stary et al., 2013b: 377 (on Limonidae from Vraz near Pisek, with general habitat description). Distribution. Stary and Vonicka, 2018: 67 (on Limoniidae and Pediciidae of two localities in northern Bohemia, with list of localities, altitudes and habitats). Denmark Distribution. Petersen and Jong, 2001b: 150 (checklist). Finland Distribution. Salmela, 2001a: 143, 144, 151 (loc(s)). Distribution. Stary and Salmela, 2004: 47 (loc(s)). Distribution. Salmela et al., 2007b: 46 (loc(s)). Distribution. Halme et al., 2012: 507 (part of a study on saproxylic nematoceran communities occupying different parts of decaying fallen Aspen trunks in a boreal forest in central Finland). Distribution. Salmela, 2012b: 238 (annotated list of Finnish crane flies). Distribution. Salmela and Petrasiunas, 2014: 31 (checklist Finnish Tipulomorpha). Distribution. Rasimus, 2018: 17 (on saproxylic craneflies in the Tampere region) (in Finnish). Distribution. Rasimus, 2018: 17 (on saproxylic craneflies in the Tampere region) (in Finnish). France Distribution. Kramer and Langlois, 2019a: 76 (record(s) Ravin de Valbois, Doubs (25)). Distribution. Quindroit, 2020a: 39, 45-48 (annotated list with information on abundancy and habitat of the Tipuloidea of the Pays de la Loire regoin, distr) (in French). Distribution. Quindroit, 2020b: 139 (loc(s) Nord (59) and Pas-de-Calais (62) with notes on habitats and distr) (in French). Distribution. Quindroit and Lemoine, 2021b: 7 (loc(s) Nord (59); see paper for habitat description) (in French). Distribution. Quindroit, 2021c: 424, 426 (loc(s) Mercantour National Park, Alpes-Maritimes (06), wide distribution through France; infrequent, usually from old forests; mycophagous). Distribution. Quindroit and Lemoine, 2022a: 25 (loc(s) Aisne (02) with notes on habitats and distr) (in French). Distribution. Quindroit and Lemoine, 2022a: 37 (table specifying the presence in the five departments of the Hauts-de-France region). Distribution. Quindroit and Racine, 2025: 360 (table specifying the presence in the five departments of the Pays de la Loire region). Georgia Distribution. Stary and Obona, 2020: 137 (loc(s); with list of localities, habitats and altitudes). Germany Distribution. Schacht, 1999: 135 (checklist Bayern). Distribution. Reusch and Oosterbroek, 2000: 162 (checklist German Bundeslander). Distribution. Reusch and Bellstedt, 2001b: 75 (checklist Thuringen). Distribution. Reusch et al., 2004: 119 (loc(s) Brandenburg). Distribution. Schacht, 2005: 5 (checklist Bayern). Distribution. Hable et al., 2010b: 91 (loc(s) Bayern). Distribution. Schacht, 2010: 17 (checklist Diptera Bayern). Distribution. Dunk and Kraus, 2014: 169 (loc(s) Bayern). Distribution. Dunk, 2018: 80, 96 (loc(s) Bayern with list of habtats). Distribution. Stuke, 2019: 144 (checklist Niedersachsen and Bremen with reference to original sources). Distribution. Stuke et al., 2020: 283 (loc(s) Niedersachsen). Great Britain Distribution. Skidmore, 2003: 23-24, 27 (loc(s) Windsor Park). Distribution. Stubbs, 2003: 77 (distr in GB). Distribution. Chandler, 2017: 92 (one male in decayed hardwood of Beech [Fagus]; first reared fromWindsor by O.W. Richards in 1932, latest Windsor record in 2014; develops in bracket fungi, only otherwise known in Britain from Epping and New Forests). Distribution. Donisthorpe, 2017a: 92 (loc(s) Berkshire: Windsor). Distribution. Brock, 2018: 5 (observations in the New Forest on the gathering of adults on Beech [Fagus] and Oak [Quercus] and association with Inotus bracket fungi; see paper for details). Italy Distribution. Stary, 2003a: 125 (loc(s)). Distribution. Podenas and Podeniene, 2008: 351-352 (annotated checklist South Tyrol [Prov. Bolzano]). Japan Distribution. Kinota, 2006: 183 (loc(s) Sapporo) (as subspecies truncata) (in Japanese). Distribution. Nakamura, 2002: 1, 169 (loc(s) Honshu). Distribution. Sueyoshi et al., 2007: 247-257 (loc(s) Honshu; on host abundance of limoniid flies of Agaricales (gill mushrooms) and Aphyllophorales (non-gilled fungi) and the importance of these flies as phoronts of mites inhabiting these fungal orders; see paper for details on fungal species and forest types). Distribution. Kato, 2023a: 30 (loc(s) Honshu) (in Japanese). Distribution. Kato and Yamauchi, 2024: 28 (loc(s) Toyama pref., Honshu). Distribution. Kato, 2024b: 11 (records from Tokamachi city, Honshu) (in Japanese). Kazakhstan First record. Devyatkov, 2020a: 53 (loc(s) East Kazakhstan, distr) (in Russian). Latvia First record. Stary, 2004f: (Fauna Europaea: added Latvia). Distribution. Salmela and Vartija, 2007: 11 (loc(s)). Lithuania Distribution. Pakalniskis et al., 2006: 22 (checklist). Distribution. Podeniene et al., 2010: 234 (loc(s), distr). Distribution. Gorban and Podeniene, 2022: 5, 8-9 (loc(s); reared from Poplar [Populus tremula], mycetophagous species; see paper for details). Luxembourg First record. Vogtenhuber, 2007: 354 (loc(s)). Mongolia Distribution. Yadamsuren et al., 2015: 475 (loc(s) Mongolia). Distribution. Oliveira and Eck, 2025: Specieslist (loc(s) Ovorhangay and Khentii aimags). Montenegro Distribution. Oosterbroek and Simova-Tosic, 2004: 448 (review literature). Netherlands First record. Theowald, 1954b: 30 (XXX). Distribution. Jong and Oosterbroek, 2002b: 34 (checklist). North Korea First record. Podenas and Byun, 2016: 340, 344 (loc(s), map). Norway Distribution. Olsen et al., 2018: 162-163 (annotated checklist Nordic countries and for Norway according to the Strand regions). Distribution. Olsen and Andersen, 2021: 216 (loc(s) Innlandet, in former Hedmark, with habitats specified separately, for habitats see also Jonassen and Andersen, 2020). Poland Distribution. Wiedenska, 2007a: 76 (checklist). Distribution. Wiedenska, 2008: 46-47 (loc(s) Parowy Janinowski nature reserve near Lodz) (in Polish). Distribution. Wiedenska, 2010: 134 (loc(s) Parku Krajobrazowego Wzniesien Lodzkich, Lodz). Distribution. Wiedenska, 2015b: 56 (loc(s) near Lodz). Distribution. Wiedenska, 2017a: 24 (faunistic and habitat data of species from the Gorce NP) (in Polish). Distribution. Obona et al., 2019a: 124 (records of Diptera from the Babia Gora Massif). Distribution. Palaczyk and Klasa, 2021: 173 (list of Diptera in Catalogue of the fauna of the Ojcow National Park). Romania Distribution. Parvu, 2004: 186 (loc(s)). Distribution. Ujvarosi, 2007: 228-231 (checklist Limoniidae). Distribution. Parvu, 2009: 448 (on some Diptera from Bucuresti and its surroundings). Distribution. Ujvarosi et al., 2011b: 109 (loc(s) Dupa Lanca marshy area near Voslobeni) (in Hungarian, species list with habitat information in English). Russia Distribution. Lantsov, 2003c: 104 (loc(s) Caucasus region, distr). Distribution. Paramonov, 2006a: 152 (loc(s) RUC: Ulyanovskaya oblast, distr). Distribution. Pilipenko and Sidorenko, 2006b: 150 (checklist Kedrovaya Pad, distr) (in Russian). Distribution. Pilipenko, 2009c: 335 (loc(s) Lazovsky Nature Reserve, Primorskiy kray). Distribution. Krivosheina and Krivosheina, 2010: 777 (loc(s) RUN: Vologda and RUC: Moscow provs., Primorsky kray). Distribution. Humala and Polevoi, 2012: 143 (loc(s) RUN: Kareliya) (in Russian). Distribution. Jakovlev et al., 2014: 310 (loc(s) RUN: Kareliya). Distribution. Paramonov and Klepikov, 2014: 36 (loc(s) Yaroslavskaya oblast). Distribution. Paramonov, 2014: 39-43 (revision coll. Eversmann, loc(s) RUE: Orenburgskaya oblast) (in Russian). Distribution. Paramonov, 2015: 69 (loc(s) RUS: Saratovskaya oblast) (in Russian). Distribution. Ruchin and Pilipenko, 2015: 58 (loc(s), checklist RUC: Mordoviya Rep.). Distribution. Paramonov and Pilipenko, 2016: 100 (loc(s) RUC: Tverskaya oblast) (in Russian). Distribution. Paramonov, 2016: 130-131 (loc(s) RUC: Chuvash Rep.) (in Russian). Distribution. Paramonov, 2017a: 172 (loc(s) RUC: Chuvash Rep.) (in Russian). Distribution. Tschorsnig, 2017: 59 (host of Palaearctic Tachinidae; for species of Tachinidae, localities and references see paper). Distribution. Paramonov, 2018a: 184 (loc(s) RUC: Chuvash Rep., distr) (in Russian). Distribution. Polevoi et al., 2018: supplementary species list: emerged from fallen Aspens [Populus tremula] in Kivach Nature Reserve, Kareliya). Distribution. Belyaev and Farisenkov, 2019: 8 (used in a study on allometry of wing shape and venation in Diptera, material was collected in RUC: Moskovskaya oblast). Distribution. Dvorak et al., 2020: 6 (on Diptera collected in beer traps in the RUC: Mordoviya Rep.; collected in Pine [Pinus], floodplain, deciduous and mixed forests). Distribution. Dvorakova et al., 2020: 14-15 (on Diptera collected in beer traps in the western Ukraine; collected in a deciduous forest). Distribution. Pilipenko et al., 2020: 362 (loc(s) RUC: Mordoviya Rep., distr). Distribution. Kolcsar et al., 2021b: 152-154 (loc(s) RUE: Bashkortostan Rep. and Tatarstan Rep.). Distribution. Dvorak et al., 2022a: 134 (on Diptera collected in beer traps in the Volga Region, M. quadrimaculata being collected in Pine [Pinus], decidous and mixed forests). Distribution. Solodkii and Sergeev, 2023: 172 (loc(s) Primorskiy kray) (in Russian). Distribution. Devyatkov and Dubatolov, 2025: 36 (loc(s) Khabarovskiy kray, Bolshekhekhtsirsky Nature Reserve, distr) (in Russian). Slovakia Distribution. Stary, 2009d: 37 (loc(s) Polana area). Distribution. Stary, 2009i: webpage (checklist). Distribution. Obona et al., 2025a: 30 (collected in a mosquito trap at the Kosice Zoo, perhaps because of being attracted to CO2). South Korea First record. Jo, 2012: 26 (loc(s)) (in Korean; more or less repeated, in English, in Jo and Kong, 2014). Distribution. Podenas, 2013: 20 (literature overview of North and South Korean records). Distribution. Podenas and Byun, 2016: 340, 344 (loc(s), map). Spain First record. Eiroa and Carles-Tolra, 2019: 132 (loc(s) Navarra, distr) (in Spanish). Sweden Distribution. Fritz and Lindstrom, 2013: 62 (loc(s) Hallands County) (in Schwedish). Switzerland Distribution. Podenas et al., 2006: 175, 285 (map, vertical distr per thermic level). Turkey First record. Koc et al., 2005b: 616-618 (loc(s), map, checklist). Distribution. Koc et al., 2016: 12 (provinces Marmara region). on flight period (mainly from 2000 onward): Albania Kolcsar et al., 2021b: month(s): 7. Austria Vogtenhuber and Kofler, 2017: month(s): 9. Belarus Gavryushin in litt., 2013d: month(s): 7. Paramonov, 2019a: month(s): 6. China Mao and Yang, 2010a: month(s): 7-8. Czech Rep. Stary et al., 2005a: month(s): 5-9. Sevcik, 2006: month(s): 6-7. Sevcik, 2010: month(s): 5-10. Stary et al., 2013b: month(s): 6-9. Stary and Vonicka, 2018: month(s): 6-8(10). Finland Salmela, 2001a: month(s): 6-8. France Quindroit, 2020a: month(s): 5-6. Quindroit, 2020b: month(s): 6-7, 9-10. Quindroit and Lemoine, 2021b: month(s): 5, 9. Quindroit and Lemoine, 2022a: month(s): 9. Georgia Stary and Obona, 2020: month(s): 5. Germany Reusch et al., 2004: month(s): 5-8. Dunk, 2018: month(s): 5-7. Great Britain Skidmore, 2003: month(s): 7. Stubbs, 2003: month(s): 4-6, 8. Chandler, 2017: month(s): 7. Donisthorpe, 2017a: month(s): 7. Brock, 2018: month(s): 5-7. Japan Nakamura, 2002: month(s): 5-6, 9-10. Kato, 2023a: month(s): 6, 8. Kato and Yamauchi, 2024: month(s): 8-9. Kato, 2024b: month(s): 6. Kazakhstan Devyatkov, 2020a: month(s): 6-8. Latvia Salmela and Vartija, 2007: month(s): 5. Lithuania Podeniene et al., 2010: month(s): 5-10. Luxembourg Vogtenhuber, 2007: month(s): 6-10. Mongolia Oliveira and Eck, 2025: month(s): 7-8. North Korea Podenas and Byun, 2016: month(s): 7. Norway Olsen and Andersen, 2021: month(s): (6-9). Poland Wiedenska, 2008: month(s): 6-7. Wiedenska, 2017a: month(s): 8. Obona et al., 2019a: month(s): 6. Romania Parvu, 2004: month(s): 6, 8. Parvu, 2009: month(s): 9. Ujvarosi et al., 2011b: month(s): 6. Russia Lantsov, 2003c: month(s): 8. Paramonov, 2006a: month(s): 5-6. Pilipenko, 2009c: month(s): 7-9. Paramonov and Klepikov, 2014: month(s): 5-6, 8. Paramonov, 2015: month(s): 6-7. Ruchin and Pilipenko, 2015: month(s): 5-6, 8. Paramonov and Pilipenko, 2016: month(s): 5-7. Paramonov, 2016: month(s): 5, 7-9. Paramonov, 2017a: month(s): 5-6, 8. Paramonov, 2018a: month(s): 5-9. Dvorak et al., 2020: month(s): 8-10. Dvorakova et al., 2020: month(s): 7. Pilipenko et al., 2020: month(s): 5-9. Kolcsar et al., 2021b: month(s): 4, 6-7. Dvorak et al., 2022a: month(s): 5-8. Solodkii and Sergeev, 2023: month(s): 7. Devyatkov and Dubatolov, 2025: month(s): 6, 8-9. Slovakia Stary, 2009d: month(s): 5-10. Sevcik, 2010: month(s): 5-10. South Korea Jo, 2012: month(s): 6, 9. Podenas and Byun, 2016: month(s): 4, 6-9. Spain Eiroa and Carles-Tolra, 2019: month(s): 8. Switzerland Podenas et al., 2006: month(s): 5-9. Turkey Koc et al., 2005b: month(s): 6-7. on altitude (mainly from 2000 onward): Albania Kolcsar et al., 2021b: altitude: 690 m. Austria Vogtenhuber and Kofler, 2017: altitude: 700-1630 m. China Mao and Yang, 2010a: altitude: 1200-1400 m. France Quindroit, 2021c: altitude: 1400 m. Georgia Stary and Obona, 2020: altitude: 810 m. Germany Reusch et al., 2004: altitude: 45-100 m. Italy Stary, 2003a: altitude: 1500-1600 m. Japan Kato and Yamauchi, 2024: altitude: 330-664 m. Kazakhstan Devyatkov, 2020a: altitude: 465-1030 m. Mongolia Oliveira and Eck, 2025: altitude: 1000-2280 m. North Korea Podenas and Byun, 2016: altitude: 1311 m. Norway Olsen and Andersen, 2021: altitude: 290 m. Poland Wiedenska, 2017a: altitude: 800 m. Obona et al., 2019a: altitude: 790 m. Russia Lantsov, 2003c: altitude: 650-1350 m. Paramonov, 2015: altitude: 302 m. Kolcsar et al., 2021b: altitude: 75-533 m. Devyatkov and Dubatolov, 2025: altitude: 70-80 m. South Korea Podenas and Byun, 2016: altitude: 439-885 m. Spain Eiroa and Carles-Tolra, 2019: altitude: 610-650 m. Turkey Koc et al., 2005b: altitude: 380-840 m.