Details for Helius (Helius) longirostris

Name: Helius (Helius) longirostris (Meigen, 1818) Publication: Syst. Beschr. 1: 146 (as Limnobia) Status: Recognized taxon, Synonym(s): inornatus (Meigen, 1830); dubius Edwards, 1921. Classification: Family Limoniidae Subfamily Limoniinae Keys: Stary and Freidberg, 2007 (Limoniidae Israel); Boardman, 2016 (craneflies Shropshire); Peeters en Oosterbroek, 2016c (craneflies Netherlands, Belgium and Luxembourg) (in Dutch); Stubbs and Kramer, 2016h; Stubbs, 2021 (British craneflies); Quindroit, 2022c (European Helius, in French and English); Solodkii and Pilipenko, 2025 (Helius Russia) Variant spelling(s): lognirostris habitus male habitus male habitus male habitus male habitus female plus 43 more images of habitus, habitus, body part(s), body part(s), body part(s), body part(s), body part(s), body part(s), body part(s), hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, ovipositor, ovipositor, wing, wing, wing, wing, wing, wing, wing, wing, habitat, habitat, habitat, habitat, habitat, habitat, habitat, habitat and habitat. Distribution: Austria, Belarus, Belgium, Bulgaria, Czech Rep., Denmark, Finland, France (incl. Corsica), Germany, Great Britain, Hungary, Ireland, Italy (Trentino-Alto Adige), Latvia, Lithuania, Netherlands, Norway, Poland, Romania, Serbia, Slovakia, Sweden, Switzerland, Turkey (European part: Edirne, Kirklareli), Ukraine; Russia: RUN, RUW, RUC, RUE, RUS, North Caucasus; Turkey (Asiatic part: see citations for provinces); Israel;; Russia: WS (Altay); Kazakhstan (east). Note: see the manual for abbreviations: present-day Russia. Region(s): Westpalaearctic;; Eastpalaearctic General note: Added: Italy (Trentino-Alto Adige) (Stary and Oosterbroek, 1996). For Finland check Salmela, 2012b, 2012c, and citations below. For Norway check Olsen et al., 2018, and citations below. For Great Britain and Ireland check Stubbs, 2021, and citations below. For France check Quindroit, 2022c, and citations below. To be confirmed for Egypt (in Palaearctic Catalogue (Savchenko et al., 1992)). Map: Podenas et al., 2006 (Switzerland); Boardman, 2007 (Shropshire); Quindroit, 2022c (France) Biology: Alexander, 1919e, 1920r (habitat). Figure references: Habitus: Quindroit, 2022c (copula) Head: Boardman, 2007; Quindroit, 2022c Wing: Slipka and Stary, 1977b; Podenas et al., 2006; Podenas and Gelhaus, 2007; Quindroit, 2022c Hypopygium: Stary and Rozkosny, 1970a; Mendl, 1979a; Podenas et al., 2006; Podenas and Gelhaus, 2007; Kramer and Morris, 2022b; Quindroit, 2022c; Solodkii and Pilipenko, 2025 Ovipositor: Stary and Rozkosny, 1970a; Quindroit, 2022c Other figures: Frantsevich, 2004 (mid coxa) Immature stages Larva: Podeniene, 2003a; Krivosheina and Krivosheina, 2011c Citations on biology (mainly from 2000 onward): Belarus Habitat. Paramonov, 2019a: 95 (loc(s) Berezinsky Biosphere Reserve, distr, habitats, citing various authors) (in Russian). Country not relevant Habitat. Krivosheina and Krivosheina, 2011c: 87-88 (note on larva, figs, biology) (in Russian). Biology. Olsen et al., 2018: 148 (habitats of adults and larvae more or less as in Helius flavus; larvae occur at the waterline of slowly flowing or small standing waters, breathing by holding the stigma field between leaves of shore plants (Bangerter 1929, Cramer 1968); adults were reared from a wet decorticated log, probably from the mud adhering to the woody debris (Godfrey 2003); details on rearing and lifecycle in Brinkmann (1991), Cramer (1968) and Reusch (1988); the period of flight is may-september; see table 1 for habitat information of the Norwegian locality(ties)). Czech Rep. Habitat. Stary et al., 2013b: 377 (on Limonidae from Vraz near Pisek, with general habitat description). Habitat. Stary and Vonicka, 2018: 65 (on Limoniidae and Pediciidae of two localities in northern Bohemia, with list of localities, altitudes and habitats). Denmark Habitat. Byriel and Rojas, 2017: 118 (among the 101 species in a study on cranefly hotspots in unmanaged and managed forests) (in Danish). Finland Habitat. Salmela, 2001a: 144, 146, 151 (inhabits marshes and swamps, citing Brinkmann, 1991). Biology. Salmela, 2008: 11, 24-26, 32, 51 (49) (ecology or habitat: swamp; phenology:abundant later in the season, weeks 26-29). France Habitat. Kramer and Withers, 2007: 159-160 (marshland species with larvae in the more or less saturated organic mud of lake margins and marshes). Biology. Quindroit, 2020a: 36, 45-48 (annotated list with information on abundancy and habitat of the Tipuloidea of the Pays de la Loire regoin, distr) (in French). Habitat. Quindroit, 2020b: 137 (loc(s) Nord (59) and Pas-de-Calais (62) with notes on habitats and distr) (in French). Habitat. Herbrecht et al., 2021: 57 (loc(s) Loire-Atlantique (44) based on malaise-trapping with notes on habitats) (in French). Habitat. Quindroit and Lemoine, 2021a: 23, 25 (loc(s), resultats de deux campagnes de piegeage dans le Pas-de-Calais (62); see paper for details on habitat). Habitat. Quindroit et al., 2021: 256-258 (collected during an entomological inventory in the canopy of Oak [Quercus] stands, see paper for habitat and forest details; marais). Habitat. Quindroit and Lemoine, 2022a: 23 (loc(s) Oise (60) and Somme (80) with notes on habitats and distr) (in French). Habitat. Quindroit, 2022c: 343-350 (review of the five French species of Helius, loc(s) 20 departements, map) (in French). Germany Habitat. Lehmann and Reusch, 2009: 122 (from around lowland springs). Habitat. Reiff et al., 2015: 234 (loc(s) Baden-Wurttemberg with list of habitats). Great Britain Habitat. Godfrey, 2003a: 22, 43 (this non-xylophagous species was reared from a wet decorticated log, probably from the mud adhering to the woody debris; review literature). Biology. Stubbs, 2003: 20, 22-23 (larvae of Helius are aquatic, living among (and hiding under) submerged parts of tall emergent vegetation; larvae are assumed to be either grazers of decaying leaf surfaces or less specialised detritivores; adults have a long proboscis ideal for nectar feeding but there are no records at flowers (such activity may have gone un-noticed at night)). Habitat. Boardman, 2007: 81 (found at ditch, pond or lake margins over much of the lowlands, also in carr or on marshy ground). Habitat. Cranston and Drake, 2010: 175 (Helius larvae can be found in marginal situations along flowing and standing waters [no species mentioned]). Habitat. Kramer, 2011j: 22, 25 (by ponds and marshlands). Biology. Featherstone and Chandler, 2013a: 43, 2013b: 180 (first european record of a forcipomyiine midge (Diptera, Ceratopoginidae) feeding on a cranefly wing (Diptera, Limoniidae) and first record of an Atrichopogon species exhibiting this behaviour; review of published observations of ceratopogonid midges feeding on craneflies). Habitat. Brighton, 2017c: 28, 39 (overall regional checklist Lancashire and Cheshire (VC58, 59 and 60) with notes on habitats and/or regional occurrence and/or status; see paper for details). Habitat. Stubbs, 2018b: 2 (prefers wet scrub carr). Habitat. Kramer and Morris, 2022b: 21 (info on the species, on the distr and map of loc(s) Leicestershire and Rutland (VC55); found near waterbodies). Ireland Habitat. Ashe et al., 2008: 11 (specis associated with weedy ponds and bogs). Lithuania Habitat. Podeniene, 2003a: 1-295 (larva, descr, figs, habitat) (in Lithuanian). Habitat. Podeniene, 2003b: 11, 21, 22, 28 (larval habitat: sand and silt in riparian zone of small rivulets, swamps at the bottom of karst pits in old deciduous forest, mud (in bogs of low lying woody swamps, on swampy riversides, in marshy meadows)). Norway Habitat. Olsen and Andersen, 2021: 215 (loc(s) Innlandet, in former Hedmark, with habitats specified separately, for habitats see also Jonassen and Andersen, 2020). Romania Habitat. Parvu, 2003: 233. Habitat. Ujvarosi, 2005a: 245 (swamps, moist soil, standing water, citing various sources). Habitat. Kolcsar et al., 2013: 72 (loc(s) Cluj-Napoca and surroundings; list of habitats with altitudes). Russia Habitat. Przhiboro, 2003: 363 (most numerous limoniid species in the water margin zone of two eutrophic lakes; Helius larvae are the only Limoniidae inhabiting the surface water layer within the lakes littoral zone, they occur only in sites with very dense emergent vegetation; rearing). Habitat. Przhiboro, 2004: 103, 111, 117 (in Russian). Biology. Przhiboro, 2012: 260 (on the abundance and biomass values of Tipuloidea immatures at five sites of the lake water margin zone, Rakovye lakes, Leningradskaya oblast) (in Russian). Habitat. Lantsov in litt., 2014: (in buckbean (Menyanthes trifoliata L.) sedge (Carex vesicaria L., Carex acuta L.) grass bog community (Dead Lake) and in fir forest with wood sorrel). Habitat. Lantsov, 2014b: 153 (loc(s) North Caucasus: Karachay-Cherkess Rep.; found in same habitats as Prionocera turcica). Biology. Lantsov, 2015a: 367 (loc(s) North Caucasus: Krasnodarskiy kray, Karachay-Cherkess Rep., Kabardino-Balkar. Rep., Dagestan, with data on habitats, ecology and chorology) (in Russian). Habitat. Lantsov, 2017b: 107 (on dominant species of the crane flies (Diptera, Tipuloidea) of the North Caucasus as biological indicators of habitats) (in Russian). Habitat. Kolcsar et al., 2021b: 126 (RUE: wetland, dry meadow). on characters and taxonomy (mainly from 2000 onward): Country not relevant Characters. Krzeminski, 2002: 318-319 (comparison with fossil species). Characters. Podeniene, 2002: 299-301 (comparison). Characters. Frantsevich, 2004: 133-134 (structure of coxa, figs). Immatures. Krivosheina and Krivosheina, 2011c: 87-88 (note on larva, figs, biology) (in Russian). Characters. Olsen et al., 2018: 148 (belongs to the Westpalaearctic species of Helius with a dark rostrum and a faint pterostigma; among these species characterized by the brown to grey dorsal thorax with three dark brown median stripes; fig). Characters. Solodkii and Pilipenko, 2025: 400-401, 404-406 (notes on the morphology of Helius, comparison). France Key. Quindroit, 2022c: 343-350 (review of the five French species of Helius, key, figs) (in French, key also in English). Great Britain Characters. Kramer and Morris, 2022b: 21 (diagnosis, figs). Israel Key. Stary and Freidberg, 2007: 303, 305, 345 (key). Lithuania Immatures. Podeniene, 2003a: 1-295 (larva, descr, figs, habitat) (in Lithuanian). on distribution (mainly from 2000 onward): Belarus First record. Gavryushin in litt., 2013d: (loc(s) Minsk region). Distribution. Paramonov, 2019a: 95 (loc(s) Berezinsky Biosphere Reserve, distr, habitats, citing various authors) (in Russian). Belgium Distribution. Martens et al., 2014: 114, 124 (loc(s) Oost-Vlaanderen). Bulgaria Distribution. Hubenov, 2021a: 36 (checklist with summary of distr in Bulgaria, references). Czech Rep. Distribution. Stary et al., 2005a: 27 (loc(s)). Distribution. Stary, 2009i: webpage (checklist). Distribution. Stary et al., 2013b: 377 (on Limonidae from Vraz near Pisek, with general habitat description). Distribution. Stary and Vonicka, 2018: 65 (on Limoniidae and Pediciidae of two localities in northern Bohemia, with list of localities, altitudes and habitats). Denmark Distribution. Petersen and Jong, 2001b: 150 (checklist). Finland Distribution. Salmela, 2001a: 144, 146, 151 (loc(s)). Distribution. Salmela, 2002: 30-32 (loc(s) lake Iidesjarvi) (in Finnish). Distribution. Salmela et al., 2007b: 46 (loc(s)). Distribution. Salmela, 2008: 11, 51 (49) (loc(s), known from all Finnish ecoregions). Distribution. Salmela, 2012b: 237 (annotated list of Finnish crane flies). Distribution. Salmela and Petrasiunas, 2014: 30 (checklist Finnish Tipulomorpha). Distribution. Salmela et al., 2015: 102 (list of localities in extensive report on malaise-trapping of insects in conservation areas in Lapland in 2012–2014) (in Finnish). France Distribution. Kramer and Withers, 2007: 159-160, 163 (loc(s) Ain (01)). Distribution. Kramer, 2007a: 33 (loc(s) Ain (01)). Distribution. Withers, 2008: 137 (loc(s) Ain (01)). Distribution. Quindroit, 2020a: 36, 45-48 (annotated list with information on abundancy and habitat of the Tipuloidea of the Pays de la Loire regoin, distr) (in French). Distribution. Quindroit, 2020b: 137 (loc(s) Nord (59) and Pas-de-Calais (62) with notes on habitats and distr) (in French). Distribution. Herbrecht et al., 2021: 57 (loc(s) Loire-Atlantique (44) based on malaise-trapping with notes on habitats) (in French). Distribution. Quindroit and Lemoine, 2021a: 23, 25 (loc(s), resultats de deux campagnes de piegeage dans le Pas-de-Calais (62); see paper for details on habitat). Distribution. Quindroit et al., 2021: 257 (recorded for Loiret (45)). Distribution. Quindroit and Lemoine, 2022a: 23 (loc(s) Oise (60) and Somme (80) with notes on habitats and distr) (in French). Distribution. Quindroit and Lemoine, 2022a: 37 (table specifying the presence in the five departments of the Hauts-de-France region). Distribution. Quindroit, 2022a: 25-28 (additions to the annotated list of the Tipuloidea of Pays de la Loire as given in Quindroit, 2020a). Distribution. Quindroit, 2022c: 343-350 (review of the five French species of Helius, loc(s) 20 departements, map) (in French). Distribution. Quindroit and Racine, 2025: 360 (table specifying the presence in the five departments of the Pays de la Loire region). Germany Distribution. Schacht, 1999: 135 (checklist Bayern). Distribution. Reusch and Oosterbroek, 2000: 162 (checklist German Bundeslander). Distribution. Reusch and Bellstedt, 2001b: 75 (checklist Thuringen). Distribution. Schacht, 2005: 5 (checklist Bayern). Distribution. Lehmann and Reusch, 2009: 122 (loc(s) Niedersachsen). Distribution. Hable et al., 2010b: 91 (loc(s) Bayern). Distribution. Schacht, 2010: 17 (checklist Diptera Bayern). Distribution. Reiff et al., 2015: 234 (loc(s) Baden-Wurttemberg with list of habitats). Distribution. Stuke, 2019: 143 (checklist Niedersachsen and Bremen with reference to original sources). Distribution. Stuke et al., 2020: 283 (loc(s) Niedersachsen). Great Britain Distribution. Kramer, 1999a: 31 (loc(s) Dorset). Distribution. Godfrey, 2003a: 22, 43 (loc(s), distr, review literature). Distribution. Boardman, 2007: 81 (loc(s), map). Distribution. Skidmore, 2009: 129 (review of occurrence on the Western Isles of Scotland, distr). Distribution. Hancock, 2011a: 2 (loc(s) Coll, Southern Inner Hebrides). Distribution. Kramer, 2011j: 22 (loc(s) Leicestershire and Rutland (VC55)). Distribution. Featherstone and Chandler, 2013a: 43 (loc(s) Inverness-shire). Distribution. Hewitt, 2014: (provisional checklist Cumbrian Diptera). Distribution. Drake, 2016: 20 (study on the relative importance to Diptera of pasture and ditch margins on Catcott Reserve, Somerset). Distribution. Brighton, 2017c: 28, 39 (overall regional checklist Lancashire and Cheshire (VC58, 59 and 60) with notes on habitats and/or regional occurrence and/or status; see paper for details). Distribution. Kramer, 2018f: 1 (loc(s) Staffordshire). Distribution. Stubbs, 2018b: 2 (loc(s) Staffordshire). Distribution. Kramer and Morris, 2022b: 21 (info on the species, on the distr and map of loc(s) Leicestershire and Rutland (VC55); found near waterbodies). Hungary Distribution. Toth, 2005: 169 (loc(s)). Ireland Distribution. Ashe et al., 2008: 11 (additional Irish record(s), distr). Distribution. Chandler et al., 2008: 13 (checklist). Israel Distribution. Stary and Freidberg, 2007: 303, 305, 345 (only known from older literature, confirmation required, distr). Kazakhstan First record. Devyatkov, 2019b: 247 (loc(s) Pavlodarskaya oblast, distr) (in Russian). Distribution. Devyatkov, 2020a: 52 (loc(s) East Kazakhstan, distr) (in Russian). Latvia Distribution. Stary, 2004f: (Fauna Europaea: added Latvia). Lithuania Distribution. Pakalniskis et al., 2006: 20 (checklist). Netherlands Distribution. Jong and Oosterbroek, 2002b: 34 (checklist). Distribution. Hoven, 2015: 10 (loc(s) Limburg). Distribution. Hoven, 2018: 16 (loc(s) Zeeland). Distribution. Stary, 2020a: 45 (Limoniidae Kaaistoep, Noord-Brabant). Norway First record. Olsen et al., 2018: 148, 162-163 (loc(s) in seven regions; distr; annotated checklist Nordic countries and for Norway according to the Strand regions). Distribution. Olsen and Andersen, 2021: 215 (loc(s) Innlandet, in former Hedmark, with habitats specified separately, for habitats see also Jonassen and Andersen, 2020). Poland Distribution. Wiedenska, 2007a: 75 (checklist). Distribution. Wiedenska, 2008: 46 (loc(s) Parowy Janinowski nature reserve near Lodz) (in Polish). Distribution. Wiedenska, 2010: 133 (loc(s) Parku Krajobrazowego Wzniesien Lodzkich, Lodz). Distribution. Wiedenska, 2015b: 56 (loc(s) near Lodz). Distribution. Wiedenska and Syratt, 2023: 144 (loc(s) Bieszczady Mountains, distr and remarks). Romania Distribution. Parvu, 2003: 233 (loc(s)). Distribution. Parvu, 2006: 261 (loc(s)). Distribution. Ujvarosi, 2007: 228-231 (checklist Limoniidae). Distribution. Kolcsar et al., 2013: 72 (loc(s) Cluj-Napoca and surroundings; list of habitats with altitudes). Russia Distribution. Polevoi and Humala, 2003: 70 (loc(s) RUN: Kareliya). Distribution. Przhiboro, 2003: 363 (loc(s) NW Russia). Distribution. Przhiboro, 2004: 103, 111, 117 (loc(s) RUW: Leningradskaya oblast) (in Russian). Distribution. Humala and Polevoi, 2009: 66 (loc(s) RUN: Kareliya) (in Russian). Distribution. Gavryushin in litt., 2011f: (loc(s) RUN: Arkhangelskaya oblast). Distribution. Przhiboro, 2012: 260 (on the abundance and biomass values of Tipuloidea immatures at five sites of the lake water margin zone, Rakovye lakes, Leningradskaya oblast) (in Russian). Distribution. Jakovlev et al., 2014: 309 (loc(s) RUN: Kareliya). Distribution. Lantsov in litt., 2014: (new for the North Caucasus, no further locality details presented). Distribution. Lantsov, 2014b: 153 (loc(s) North Caucasus: Karachay-Cherkess Rep.; found in same habitats as Prionocera turcica). Distribution. Paramonov and Klepikov, 2014: 36 (loc(s) Yaroslavskaya oblast). Distribution. Lantsov, 2015a: 367 (loc(s) North Caucasus: Krasnodarskiy kray, Karachay-Cherkess Rep., Kabardino-Balkar. Rep., Dagestan, with data on habitats, ecology and chorology) (in Russian). Distribution. Lantsov, 2016: 165-170 (on craneflies from Arkhyz, Karachay-Cherkessia Rep., North Caucasus) (in Russian). Distribution. Paramonov and Pilipenko, 2016: 99 (loc(s) RUC: Tverskaya oblast) (in Russian). Distribution. Lantsov, 2017b: 107 (on dominant species of the crane flies (Diptera, Tipuloidea) of the North Caucasus as biological indicators of habitats) (in Russian). Distribution. Pilipenko, 2017a: 389 (investigation of Tipuloidea of the Altay, first record for Altay and Eastpalaearctic) (in Russian). Distribution. Paramonov, 2018b: 61 (loc(s) RUS: Saratovskaya oblast) (in Russian). Distribution. Pilipenko et al., 2020: 361 (loc(s) RUC: Mordoviya Rep., distr). Distribution. Kolcsar et al., 2021b: 125-127 (loc(s) various regions in RUE). Distribution. Solodkii and Pilipenko, 2025: 400 (studied from the Moscovskaya oblast and Mordoviya Rep.). Serbia First record. Kolcsar et al., 2021b: 127 (loc(s)). Slovakia Distribution. Stary, 2009d: 36 (loc(s) Polana area). Distribution. Stary, 2009i: webpage (checklist). Sweden Distribution. Salmela, 2010b: 129, 133 (loc(s) Malmo, distr). Distribution. Fritz and Lindstrom, 2013: 62 (loc(s) Hallands County) (in Schwedish). Switzerland Distribution. Podenas et al., 2006: 169, 263 (map, vertical distr per thermic level). Turkey First record. Koc et al., 2016: 9 (loc(s) European and Asiatic part, Marmara region). Distribution. Provinces Asiatic part, 2025: (known from Balikesir, Canakkale, Kocaeli, Sakarya). on flight period (mainly from 2000 onward): Belarus Gavryushin in litt., 2013d: month(s): 7. Belgium Martens et al., 2014: month(s): 5. Czech Rep. Stary et al., 2005a: month(s): 5-6. Stary et al., 2013b: month(s): 5-6. Stary and Vonicka, 2018: month(s): 5-8. Finland Salmela, 2001a: month(s): 6-7. Salmela, 2008: month(s): 5-7. France Kramer and Withers, 2007: month(s): 5, 7. Kramer, 2007a: month(s): 6. Quindroit, 2020a: month(s): 5-6. Quindroit, 2020b: month(s): 4-6, 8. Quindroit and Lemoine, 2022a: month(s): 4-5, 7. Quindroit, 2022c: month(s): 4-9. Germany Reiff et al., 2015: month(s): 6. Great Britain Kramer, 1999a: month(s): 6. Hancock, 2011a: month(s): 6. Kramer, 2011j: month(s): 6-7. Featherstone and Chandler, 2013a: month(s): 6. Kramer, 2018f: month(s): 6. Stubbs, 2018b: month(s): 6. Kramer and Morris, 2022b: month(s): 6-7. Ireland Ashe et al., 2008: month(s): 5. Kazakhstan Devyatkov, 2019b: month(s): 5. Devyatkov, 2020a: month(s): 5-6. Netherlands Hoven, 2015: month(s): 5. Hoven, 2018: month(s): 5. Norway Olsen et al., 2018: month(s): 5-7. Olsen and Andersen, 2021: month(s): 6. Poland Wiedenska, 2008: month(s): 7. Wiedenska and Syratt, 2023: month(s): 6, 8. Romania Parvu, 2003: month(s): 7. Parvu, 2006: month(s): 7. Kolcsar et al., 2013: month(s): 8. Russia Gavryushin in litt., 2011f: month(s): 8. Paramonov and Klepikov, 2014: month(s): 6-7. Lantsov, 2015a: month(s): 5-7. Paramonov and Pilipenko, 2016: month(s): 6-7. Paramonov, 2018b: month(s): 6-7. Pilipenko et al., 2020: month(s): 6, 8-9. Kolcsar et al., 2021b: month(s): 6-8. Serbia Kolcsar et al., 2021b: month(s): 4-5. Slovakia Stary, 2009d: month(s): 6-7. Switzerland Podenas et al., 2006: month(s): 3, 5-10. Turkey Koc et al., 2016: month(s): 4-5, 9. on altitude (mainly from 2000 onward): Bulgaria Hubenov, 2021a: altitude: 0-700 m. France Quindroit, 2022c: altitude: 1100 m. Germany Reiff et al., 2015: altitude: 650 m. Kazakhstan Devyatkov, 2019b: altitude: 126 m. Devyatkov, 2020a: altitude: 300-982 m. Norway Olsen and Andersen, 2021: altitude: 251 m. Poland Wiedenska and Syratt, 2023: altitude: 470-700 m. Romania Parvu, 2003: altitude: 800 m. Russia Gavryushin in litt., 2011f: altitude: 20 m. Lantsov in litt., 2014: altitude: 1490 m. Lantsov, 2015a: altitude: 20-1506 m. Paramonov, 2018b: altitude: 302 m. Kolcsar et al., 2021b: altitude: 50-494 m. Serbia Kolcsar et al., 2021b: altitude: 66-67 m. Slovakia Stary, 2009d: altitude: 850-870 m. Turkey Koc et al., 2016: altitude: 10-215 m.