Details for Nephrotoma guestfalica guestfalicaName: | Nephrotoma guestfalica guestfalica (Westhoff, 1879) | Publication: | Jber. Westf. ProvVer. Wiss. Kunst 8: 51 (as Pachyrhina) | Status: | Recognized taxon, Synonym(s): escorialensis (Strobl, 1909). | Classification: | Family Tipulidae Subfamily Tipulinae
| Revision: | de Jong et al., 2021. | Descriptive note: | Phylogeny: Jong, 1993. | Keys: | Peeters en Oosterbroek, 2013d (craneflies Netherlands, Belgium and Luxembourg) (in Dutch); Boardman, 2016 (craneflies Shropshire); Stubbs and Kramer, 2016c; Stubbs, 2021 (British craneflies); de Jong et al., 2021 (subspecies of guestfalica) | Variant spelling(s): | guestphalica |  |  |  |  |  |
habitus male | habitus male | habitus male | habitus male | habitus male |
plus 34 more images of habitus, habitus, habitus, habitus, habitus, habitus, habitus, habitus, habitus, habitus, body part(s), body part(s), body part(s), body part(s), body part(s), body part(s), body part(s), body part(s), hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, ovipositor, ovipositor, ovipositor, wing, wing, habitat, habitat, habitat, habitat, habitat and habitat. | | | Distribution: | Belarus, Belgium, Bosnia-Herzegovina, Bulgaria, Croatia, Czech Rep., Denmark, France (incl. Corsica), Germany, Great Britain, Greece (incl. Kerkyra [Corfu], Lefkada [Lefkas], Lesvos), Hungary, Italy (incl. Sicily), Luxembourg, Netherlands, North Macedonia, Poland, Portugal, Romania, Slovakia, Spain (incl. Balearic Is [Mallorca]), Sweden (south), Switzerland, Turkey (European part: Istanbul); Turkey (Asiatic part: see citations for provinces). | Region(s): | Westpalaearctic | General note: | Added: Turkey (European part: Istanbul), Balikesir (Jong, 1998). For Spain check Eiroa and Carles-Tolra, 2019, and citations below. For Portugal check Oosterbroek et al., 2020, and citations below. For Great Britain check Stubbs, 2021, and citations below. To be confirmed for Austria (in Palaearctic Catalogue (Oosterbroek and Theowald, 1992; not listed for Austria by Franz, 1990, or Vogtenhuber, 1994) and European Russia (in Palaearctic Catalogue (Oosterbroek and Theowald, 1992; see Pilipenko, 2009a). | Map: | Jong, 1998 (general); Koc, 2004 (Mugla); Koc, 2006 (Aydin); Boardman, 2007 (Shropshire) | Figure references: | Habitus: Jong et al., 2021 (copula) Head: Jong et al., 2021 Wing: Jong et al., 2021 Hypopygium: Parvu, 2005; Koc, 2006; Jong et al., 2021; Kramer and Morris, 2021b Ovipositor: Koc, 2006; Jong et al., 2021 Other figures: de Jong et al., 2021 (abdomen tergite 1)
| | | Citations | | | Belgium Biology. Peeters, 2023: 229 (loc(s) Botanic Garden Jean Massart, Brussels-Capital Region, common spring and summer species from a variety of habitats, frequently found in gardens). | | France Habitat. Dufour, 2003b: 85 (loc(s) with information on habitat(s)) (in French). Habitat. Nicolas, 2006: 67 (on findings in the Picardie or Ile-de-France) (in French). Habitat. Kramer, 2007a: 33 (adults were collected at an open marsh). Habitat. Tillier and Dehalleux, 2019: 152 (loc(s) with habitat info, summary habitat) (in French). Habitat. Tillier and Oosterbroek, 2019: 228 (loc(s) and distr Mercantour; habitat: pentes seches avec feuillus [dry slopes with deciduous forest]). Habitat. Quindroit, 2020a: 16, 45-48 (annotated list with information on abundancy and habitat of the Tipuloidea of the Pays de la Loire regoin, distr) (in French). Habitat. Quindroit, 2020b: 125 (loc(s) Nord (59) and Pas-de-Calais (62) with notes on habitats and distr) (in French). Habitat. Herbrecht et al., 2021: 57 (loc(s) Loire-Atlantique (44) based on malaise-trapping with notes on habitats) (in French). Habitat. Quindroit and Lemoine, 2021a: 24, 26 (loc(s), resultats de deux campagnes de piegeage dans le Pas-de-Calais (62); see paper for details on habitat). Habitat. Quindroit and Lemoine, 2022a: 13 (loc(s) Aisne (02) and Oise (60) with notes on habitats and distr) (in French). | | Germany Habitat. Heiss et al., 2017: 162 (loc(s) Thuringen, review distr; habitats, citing various authors). | | Great Britain Habitat. Godfrey, 1999: 77 (terrestrial and river edge sand). Habitat. Hewitt et al., 2005: 37, 46, 52 (indicator species of exposed riverine sediment habitats: species of sandy river banks). Habitat. Boardman, 2007: 17 (apparently prefers sandy soils). Habitat. Kramer, 2007b: 35 (a woodland species). Habitat. Kramer, 2008d: 2 (sandy river margins). Habitat. Kramer, 2011j: 24 (larvae thrive in the rich humus on the forest floor). Habitat. Stubbs, 2013: 27 (on sand along river). Habitat. Brighton, 2017c: 30, 42 (overall regional checklist Lancashire and Cheshire (VC58, 59 and 60) with notes on habitats and/or regional occurrence and/or status; see paper for details). Habitat. Stubbs, 2024c: 34 (loc(s) Lancashire, see paper for habitat information). | | Netherlands Biology. Peeters, 2023: 229 (common spring and summer species from a variety of habitats, frequently found in gardens). | | Turkey Biology. Koc, 2004: 32-33, 115-117, 141 (habitat; phenology) (in Turkish). Phenology. Serce and Koc, 2005: 41, 46-48. Biology. Koc, 2006: 26, 28, 40-42, 120-121, 148, 166 (ecology; phenology) (in Turkish). Phenology. Koc et al., 2013: 6 (on the period of flight in the southwest Anatolia region). | | | | Country not relevant Revision. Jong et al., 2021: 33-34 (descr, key, comparison, figs). Taxonomy. Jong et al., 2021: 43-44 (discussion on subspecies concept in Tipulidae systematics). | | Netherlands Characters. Oosterbroek and Jong, 2001: 103 (records of intersexes from The Netherlands). | | Portugal Characters. Ferreira et al., 2021: 12 (barcoding based on material mentioned in Oosterbroek et al. 2020). | | Spain Figures. Mederos, 2018: 141-142 (figs and loc(s) Barcelona) (in Spanish). | | Turkey Description. Koc, 2006: 26, 28, 40-42, 120-121, 148, 166 (descr, key, figs) (in Turkish). | | | | Belgium Distribution. Martens et al., 2013: 125 (loc(s) Oost-Vlaanderen). Distribution. Peeters, 2023: 229 (loc(s) Botanic Garden Jean Massart, Brussels-Capital Region, common spring and summer species from a variety of habitats, frequently found in gardens). | | Bosnia-Herzegowina Distribution. Simova-Tosic and Oosterbroek, 2003: 45 (review literature). | | Bulgaria Distribution. Hubenov, 2021a: 26 (listed for Bulgaria, without data, references). | | Channel Is Distribution. Stubbs, 2022b: 4 (on a collection of craneflies from Jersey, collected between 1985 and 2002 by Tony Warne). | | Croatia First record. Jong et al., 2021: 34 (loc(s), distr). | | Czech Rep. Distribution. Stary, 2009k: webpage (checklist). | | Denmark Distribution. Petersen et al., 2001b: 162 (checklist). | | France Distribution. Dufour, 2003b: 85 (loc(s) with information on habitat(s)) (in French). Distribution. Nicolas, 2006: 67 (on findings in the Picardie or Ile-de-France) (in French). Distribution. Kramer, 2007a: 33 (loc(s) Ain (01)). Distribution. Garrin and Herbrecht, 2016: Annexe 14 (loc(s) Notre-Dame-des-Landes, Loire-Atlantique (44)). Distribution. Tillier and Dehalleux, 2019: 152 (new and published records Ile-de-France, loc(s) Paris (75), Seine-et-Marne (77), Yvelines (78), Seine-Saint-Denis (93) and Val-d’Oise (95)). Distribution. Tillier and Oosterbroek, 2019: 228 (loc(s) and distr Mercantour; habitat: pentes seches avec feuillus [dry slopes with deciduous forest]). Distribution. Tillier, 2019: 256 (loc(s) Var (81)). Distribution. Quindroit, 2020a: 16, 45-48 (annotated list with information on abundancy and habitat of the Tipuloidea of the Pays de la Loire regoin, distr) (in French). Distribution. Quindroit, 2020b: 125 (loc(s) Nord (59) and Pas-de-Calais (62) with notes on habitats and distr) (in French). Distribution. Herbrecht et al., 2021: 57 (loc(s) Loire-Atlantique (44) based on malaise-trapping with notes on habitats) (in French). Distribution. Labat, 2021: 93 (on Diptera taxa recorded along the over-all length of the Dordogne River and some of its tributaries; see paper for the locality details) (in French). Distribution. Quindroit and Lemoine, 2021a: 24, 26 (loc(s), resultats de deux campagnes de piegeage dans le Pas-de-Calais (62); see paper for details on habitat). Distribution. Quindroit and Lemoine, 2022a: 13 (loc(s) Aisne (02) and Oise (60) with notes on habitats and distr) (in French). Distribution. Quindroit and Lemoine, 2022a: 34 (table specifying the presence in the five departments of the Hauts-de-France region). Distribution. Lemoine and Quindroit, 2023: 5 (loc(s) Somme (80)). Distribution. Tillier et al., 2023: 172 (loc(s) Jura (39), with list of altitude(s) and habitat(s)) (in French). Distribution. Tillier, 2023b: 487 (loc(s) Corsica). Distribution. Quindroit and Racine, 2025: 356 (table specifying the presence in the five departments of the Pays de la Loire region). | | Germany Distribution. Heiss and Bellstedt, 2003: 224 (loc(s) Thuringen). Distribution. Heiss, 2003: 12 (checklist Thuringen). Distribution. Heiss et al., 2017: 162 (loc(s) Thuringen, review distr; habitats, citing various authors). Distribution. Heiss, 2017b: 21 (checklist Tipulidae Thuringen). Distribution. Heiss et al., 2019: 248 (loc(s); annotated checklist of the Tipulidae of Niedersachsen and Bremen, including the presence in the surrounding Bundeslandern and the Netherlands) (in German). Distribution. Heiss, 2019: 35, 39, 44 (loc(s) and checklist Mecklenburg-Vorpommern, see paper for further details). Distribution. Stuke, 2019: 239 (checklist Niedersachsen and Bremen with reference to original sources). Distribution. Stuke et al., 2020: 290 (loc(s) Niedersachsen, review records Niedersachsen). Distribution. Heiss and Malec, 2023: 328 (annotated checklist Hessen). Distribution. Kopetz et al., 2024: 210 (loc(s) Thuringen). | | Great Britain Distribution. Denton, 2000: 36 (loc(s) Cumbria). Distribution. Hewitt et al., 2005: 37, 46, 52 (loc(s) Cumbria). Distribution. Boardman, 2007: 17 (loc(s), map). Distribution. Kramer, 2007b: 35 (loc(s) Leicestershire). Distribution. Kramer, 2008d: 2 (loc(s) Scotland). Distribution. Kramer, 2011j: 8 (loc(s) Leicestershire and Rutland (VC55)). Distribution. Kramer, 2013i: 5, 8 (checklist of species from Leigh Woods NNR, Bristol district). Distribution. Stubbs, 2013: 27 (loc(s) Lancaster). Distribution. Hewitt, 2014: (provisional checklist Cumbrian Diptera). Distribution. Brighton, 2017c: 30, 42 (overall regional checklist Lancashire and Cheshire (VC58, 59 and 60) with notes on habitats and/or regional occurrence and/or status; see paper for details). Distribution. Kramer and Morris, 2021b: 8 (info on the species, on the distr and map of loc(s) Leicestershire and Rutland (VC55)). Distribution. Stubbs, 2024c: 34 (loc(s) Lancashire, see paper for habitat information). | | Greece Distribution. Oosterbroek, 2009c: 188 (new for Lesvos and Lefkada [Lefkas]). | | Netherlands Distribution. Oosterbroek and Jong, 2001: 103 (records of intersexes from The Netherlands). Distribution. Jong and Oosterbroek, 2002d: 40 (checklist). Distribution. Oosterbroek and Dek, 2020: 17 (Limoniidae and Tipulidae Nationaal Park Hollandse Duinen in 2018, Zuid-Holland). Distribution. Oosterbroek and Jong, 2020: 46 (Tipulidae Kaaistoep, Noord-Brabant). | | North Macedonia Distribution. Simova-Tosic and Oosterbroek, 2003: 45 (review literature). | | Poland Distribution. Skibinska and Chudzicke, 2007b: 77 (checklist). | | Portugal Distribution. Eiroa and Baez, 2002b: 79 (checklist). Distribution. Hancock et al., 2015b: 152 (loc(s) Faro). Distribution. Oosterbroek et al., 2020: 339 (recorded from five provinces, distr). | | Romania First record. Parvu, 2005: 149-150 (loc(s)). Distribution. Torok et al., 2014: 17 (checklist Romania). | | Russia Distribution. Pilipenko, 2009a: 219 (CET (west) in Palaearctic Catalogue (Oosterbroek and Theowald, 1992): omitted from Pilipenko's list for the Central European territory. | | Slovakia Distribution. Stary, 2009k: webpage (checklist). | | Spain Distribution. Eiroa and Baez, 2002b: 79 (checklist). Distribution. Carles-Tolra, 2006a: 317 (loc(s)). Distribution. Carles-Tolra, 2011: 37 (loc(s) Guipuzcoa). Distribution. Hancock et al., 2015a: 242 (loc(s) Lugo). Distribution. Mederos and Eiroa, 2018: 27 (loc(s) Lerida, distr). Distribution. Mederos, 2018: 141-142 (figs and loc(s) Barcelona) (in Spanish). Distribution. Eiroa and Carles-Tolra, 2019: 133 (loc(s) Barcelona, list of Spanish provinces, distr) (in Spanish). Distribution. Mederos et al., 2019b: 213 (Barcelona: checklist Parc Natural Serra de Collserola). Distribution. Hancock, 2020: 92 (loc(s) Jaen, Cazorla National Park). | | Switzerland Distribution. Dufour and Merz, 2012: 355 (annotated checklist canton Geneva). | | Turkey Distribution. Koc and Oosterbroek, 2001: 464 (checklist). Distribution. Koc, 2004: 32-33, 115-117, 141 (loc(s), distr, map) (in Turkish). Distribution. Serce and Koc, 2005: 41, 46-48 (loc(s)). Distribution. Koc, 2006: 26, 28, 40-42, 120-121, 148, 166 (loc(s) and map Aydin, distr) (in Turkish). Distribution. Oosterbroek, 2009c: 188 (new for Sakarya). Distribution. Koc et al., 2012: 117 (checklist of occurrence in the six provinces of southwestern Turkey). Distribution. Koc et al., 2015: 49 (results of Tipulidae investigations in the 11 provinces of the Marmara region which forms the passage between European and Asiatic Turkey). Distribution. Provinces Asiatic part, 2025: (known from Aydin, Balikesir, Bilecik, Bursa, Canakkale, Denizli, Izmir, Mugla, Sakarya, Yalova). | | | | Belgium Martens et al., 2013: month(s): 7-8. | | Channel Is Stubbs, 2022b: month(s): 6. | | Croatia Jong et al., 2021: month(s): 5. | | France Dufour, 2003b: month(s): 6. Kramer, 2007a: month(s): 6. Tillier and Dehalleux, 2019: month(s): 6-7. Tillier and Oosterbroek, 2019: month(s): 7. Tillier, 2019: month(s): 5-6. Quindroit, 2020a: month(s): 5-7. Quindroit, 2020b: month(s): 4, 6-7. Quindroit and Lemoine, 2022a: month(s): 4, 6. Lemoine and Quindroit, 2023: month(s): 5. Tillier et al., 2023: month(s): 6. Tillier, 2023b: month(s): 5. | | Germany Heiss and Bellstedt, 2003: month(s): 6. Heiss et al., 2017: month(s): 8. Heiss et al., 2019: month(s): 6. Heiss, 2019: month(s): 6-8. | | Great Britain Denton, 2000: month(s): 6-7. Kramer, 2011j: month(s): 6-7. Stubbs, 2013: month(s): 7. Kramer and Morris, 2021b: month(s): 7. Stubbs, 2024c: month(s): 6. | | Netherlands Oosterbroek and Jong, 2001: month(s): 7. Oosterbroek and Dek, 2020: month(s): 6-7. | | Portugal Hancock et al., 2015b: month(s): 4. Oosterbroek et al., 2020: month(s): 4-5. | | Romania Parvu, 2005: month(s): 5-6. | | Spain Carles-Tolra, 2006a: month(s): 5. Carles-Tolra, 2011: month(s): 5. Hancock et al., 2015a: month(s): 5. Mederos and Eiroa, 2018: month(s): 6. Mederos, 2018: month(s): 5. Eiroa and Carles-Tolra, 2019: month(s): 5. Hancock, 2020: month(s): 6. | | Turkey Koc, 2004: month(s): 4-5. Serce and Koc, 2005: month(s): 6. Koc, 2006: month(s): 4-5, 9. Koc et al., 2013: month(s): 4-6, 9-10. | | | | France Dufour, 2003b: altitude: 35-580 m. Tillier and Oosterbroek, 2019: altitude: 360-500 m. Labat, 2021: altitude: 13 m. Tillier, 2023b: altitude: 2-18 m. | | Spain Hancock et al., 2015a: altitude: 610 m. Mederos and Eiroa, 2018: altitude: 960 m. | | Turkey Koc, 2004: altitude: 23-455 m. Serce and Koc, 2005: altitude: 833 m. Koc, 2006: altitude: 12-350 m. |
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