Details for Atypophthalmus (Atypophthalmus) umbratus

Name: Atypophthalmus (Atypophthalmus) umbratus (de Meijere, 1911) Publication: Tijdschr. Ent. 54: 25 (as Dicranomyia) Status: Recognized taxon, Synonym(s): holopticus Brunetti, 1911; subumbratus (Alexander, 1923); fissilis (Alexander, 1926). Classification: Family Limoniidae Subfamily Limoniinae Descriptive note: Type-material: de Jong, 2000. Short description in Alexander, 1964i. Short description based on type-material of holopticus Brunetti in Joseph, 1976a. Monotypy: Hynes, 1993. Synonymy: Edwards, 1924e; Alexander, 1939f, 1956c, 1971f. Keys: Stary and Freidberg, 2007 (Limoniidae Israel); Kato, 2022a (Atypophthalmus Japan) Variant spelling(s): umbrata, umbrosa habitus male habitus male habitus male habitus male habitus female plus 26 more images of body part(s), body part(s), body part(s), body part(s), hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, ovipositor, ovipositor, wing, wing, wing, wing, wing, wing, wing and wing. Distribution: Belgium, Denmark, Great Britain, Netherlands, Spain (Barcelona, Canary Is [La Palma]), Sweden; Egypt; Israel;; Japan (Honshu, Kyushu);; Mexico; Cuba; Brazil, Peru;; Kenya, Madagascar, Mauritius, Nigeria, South Africa;; India (W Bengal), Indonesia (Sumatra, Java), Malaysia (Penins.), Philippines (Luzon), China (Taiwan), Taiwan, Thailand;; Hawaiian Is. Region(s): Westpalaearctic;; Eastpalaearctic;; Neotropic;; Afrotropic;; Oriental;; Australian/Oceanian General note: Added: Malaysia (Penins.) (Edwards, 1928a), Thailand (Alexander, 1953r), Madagascar (Alexander, 1964i). Presumably accidentally transported by commerce, shipping, etc. (Alexander, 1939f, 1942h, 1956c, 1967m; Byers, 1966), this also applies for the European countries where the species is found mostly in greenhouses and indoors. For Great Britain check Stubbs, 2021, and citations below. The species is not known from Tenerife, as was given in earlier versions of this catalogue (CCW) and in publications based upon it. Map: Kato, 2022a (Japan) Biology: Alexander, 1920r (habitat and short description immature stages). Byers, 1966 (habitat). Figure references: Habitus: Kato, 2020 (male); Kato, 2022a (male) Head: Mederos et al., 2019a Wing: Alexander, 1926g (as fissilis); Alexander, 1972g; Mederos, 2009; Mederos et al., 2019a; Kato, 2022a; Oosterbroek et al., 2024a Hypopygium: Alexander, 1926g (as fissilis); Byers, 1966; Gelhaus, 2009; Mederos, 2009; Mederos et al., 2019a; Kato, 2020; Kato, 2022a Other figures: Mederos et al., 2019a (thorax) Citations on biology (mainly from 2000 onward): Belgium Habitat. Kolcsar et al., 2021b: 18 (on a male collected by J. Soors in a greenhouse [in Vlaams-Brabant], 15.XI.2019, det. N.-J. Dek, see photo(s) above]). Country not relevant Biology. Mederos et al., 2019a: 157-158 (frequently collected indoors and apparently rather tolerant to pollution, citing various authors) (in Spanish). Sweden Biology. Ennerfelt, 2020: 10-11 (collected in a greenhouse at the Botanical Garden of Gothenburg) (in Schwedish) (see Artdatabanken for additional records from Sweden). on characters and taxonomy (mainly from 2000 onward): Country not relevant Characters. Mederos et al., 2023: 8 (comparison). Great Britain Characters. Stubbs, 2021: 350 (short description). Figures. Kramer, 2022c: 6 (figs (from CCW); remark, as British, based on Stubbs, 2021, distr). Israel Key. Stary and Freidberg, 2007: 303-304, 308, 330-331 (key). Japan Description. Kato, 2020: 18 (diagnosis, comparison, figs). Description. Kato, 2022a: 124, 126, 132-133 (material examined not mentioned, descr, key, comparison, figs). Spain Figures. Mederos, 2009: 179-181 (comparison, fig(s)). Figures. Mederos et al., 2019a: 157-158 (loc(s) Barcelona, first record for mainland Spain, distr) (in Spanish). on distribution (mainly from 2000 onward): Belgium First record. Kolcsar et al., 2021b: 18 (on a male collected by J. Soors in a greenhouse [in Vlaams-Brabant], 15.XI.2019, det. N.-J. Dek, see photo(s) above]). Brazil Distribution. Kato, 2020: 18 (loc(s) Minas Gerais). Canary Is First record. Mederos, 2009: 179-181 (new for Spain: Canary Is: La Palma). Cuba Distribution. Oosterbroek et al., 2024: 26 (overview craneflies West Indies with relevant reference(s) per island). Denmark First record. Gritsch in litt., 2024: (recorded on Denmarks Naturbasen from Hovedstaden in december 2021 and october 2023, including photos (see above)). Egypt First record. Stary and Freidberg, 2007: 331 (only Egypt, no loc(s)). Great Britain First record. Stubbs, 2021: 350 (introduced but meanwhile established, first recorded in hothouses at Kew Gardens in 1987 but not seen after 2002 because of the overhand of spiders and their webs in the hothouses). Distribution. Kramer, 2022c: 6 (figs (from CCW); remark, as British, based on Stubbs, 2021, distr). Hawaiian Is Distribution. Evenhuis, 2004: 31 (checklist, adventive). Israel Distribution. Stary and Freidberg, 2007: 303-304, 308, 330-331 (loc(s), distr). Japan First record. Kato, 2020: 18 (loc(s) Honshu, Kyushu, distr). Distribution. Kato, 2022a: 132, 140 (loc(s) not mentioned but on map indicated for Honshu and Kyushu, distr, map). Mexico Distribution. Contreras-Ramos and Gelhaus, 2002: 593 (checklist Mexico, distr specified at state level) (as Limonia). Netherlands First record. Oosterbroek, 2009c: 180 (found in two greenhouses in 2005 [not 1905 as given for one of the records], presumably introduced by accidental transport). Nigeria First record. Kato, 2020: 18 (loc(s) W. state). Philippines First record. Kato, 2020: 18 (loc(s) Luzon). South Africa Distribution. Kato, 2020: 18 (loc(s) Natal). Spain Distribution. Mederos et al., 2019a: 157-158 (loc(s) Barcelona, first record for mainland Spain, distr) (in Spanish). Sweden First record. Ennerfelt, 2020: 10-11 (collected in a greenhouse at the Botanical Garden of Gothenburg) (in Schwedish) (see Artdatabanken for additional records from Sweden). on flight period (mainly from 2000 onward): Belgium Kolcsar et al., 2021b: month(s): 11. Brazil Kato, 2020: month(s): 4. Canary Is Mederos, 2009: month(s): 8. Denmark Gritsch in litt., 2024: month(s): 11-12. Israel Stary and Freidberg, 2007: month(s): 7. Japan Kato, 2020: month(s): 5, 9. Netherlands Oosterbroek, 2009c: month(s): 1, 3. Nigeria Kato, 2020: month(s): 4. Philippines Kato, 2020: month(s): 1. South Africa Kato, 2020: month(s): 11. Spain Mederos et al., 2019a: month(s): 4, 8. Sweden Ennerfelt, 2020: month(s): 11. on altitude (mainly from 2000 onward): Canary Is Mederos, 2009: altitude: 750 m. Japan Kato, 2020: altitude: 3-6 m.