Details for Lipsothrix remota
Name:Lipsothrix remota (Walker, 1848)
Publication:List Dipt. Colln Brit. Mus. 1: 43. (as Limnobia)
Status:Recognized taxon, Synonym(s): ignota (Walker, 1848); clara (Tonnoir, 1921).
Classification:Family Limoniidae
Subfamily Limoniinae
Revision:Petersen, 2015.
Descriptive note:See tokunagai.
Keys:Godfrey, 2001a (Lipsothrix Great Britain, pupa); 2002a (Lipsothrix Great Britain, larva, pupa); Hancock et al., 2009 (Lipsothrix Great Britain, pupa based on thoracic spiracular gill [pupal horn]); Hancock et al., 2015a (adults Westpalaearctic Lipsothrix); Boardman, 2016 (craneflies Shropshire); Peeters en Oosterbroek, 2016c (craneflies Netherlands, Belgium and Luxembourg) (in Dutch); Stubbs and Kramer, 2016i; Stubbs, 2021 (British craneflies)
Lipsothrix remota : habitus - maleLipsothrix remota : habitus - maleLipsothrix remota : habitus - maleLipsothrix remota : habitus - maleLipsothrix remota : habitus - male
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plus 30 more images of habitus, habitus, habitus, habitus, body part(s), body part(s), body part(s), hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, ovipositor, ovipositor, ovipositor, wing, wing, wing, habitat, habitat, habitat, habitat, habitat, habitat, habitat, habitat and habitat.
Distribution:Albania, Austria, Belgium, Bosnia-Herzegovina, Bulgaria, Croatia, Czech Rep., Denmark, Estonia, France (incl. Corsica), Germany, Great Britain, Greece (incl. Euboea [Evia]), Hungary, Ireland, Italy, Latvia, Lithuania, Luxembourg, Netherlands, North Macedonia, Norway, Poland, Romania, Slovakia, Slovenia, Spain, Sweden, Switzerland, Ukraine.
Region(s):Westpalaearctic
General note:Added: Latvia (Savchenko, 1989), Ireland (Ashe et al., 1991), Spain (Geiger et al., 1994: Gerona). For Italy check Stary and Oosterbroek, 1996, and citations below. For Norway check Olsen et al., 2018, and citations below. For Great Britain and Ireland check Stubbs, 2021, and citations below.
Map:Hewitt and Parker, 2006 (Great Britain: Cumbria); Podenas et al., 2006 (Switzerland); Boardman, 2007 (Shropshire); Reusch and Weber, 2013 (Luxembourg)
Figure
references:
Habitus: Quindroit et al., 2023 (male)
Wing: Podenas et al., 2006; Krivosheina and Krivosheina, 2011c; Petersen, 2015
Hypopygium: Stary and Rozkosny, 1970a; Podenas et al., 2006; Petersen, 2015

Immature stages
Larva: Podeniene, 2003a
Pupa: Godfrey, 2001a; Hancock et al., 2009 (thoracic spiracular gill [pupal horn])

Miscellaneous
Godfrey, 2001a (oviposition); Quindroit et al., 2023 (habitat); Mitchell, 2025 (habitat)
Citationson biology (mainly from 2000 onward):
Belgium
Habitat. Reusch and Weber, 2013: 272 (found in caves, with references for Belgium).
Bulgaria
Habitat. Hubenov, 2025a: suppl: 3 (annotated list Diptera Stara Planina Mts, local distr, habitats and zoogeography).
Habitat. Hubenov, 2025b: suppl: 6 (annotated list Diptera high Bulgarian mts, local distr, habitats and zoogeography).
Country not relevant
Biology. Olsen et al., 2018: 149 (habitats of adults and larvae more or less as in Lipsothrix errans (for details see especially Hancock 2002); in several countries also recorded from caves (Reusch & Weber 2013); the period of flight is may-september; see table 1 for habitat information of the Norwegian locality(ties)).
Croatia
Habitat. Kolcsar et al., 2015a: 34 (associated with woodland streams and springs).
Czech Rep.
Habitat. Stary and Vonicka, 2018: 67 (on Limoniidae and Pediciidae of two localities in northern Bohemia, with list of localities, altitudes and habitats).
Denmark
Habitat. Byriel and Rojas, 2017: 118 (among the 101 species in a study on cranefly hotspots in unmanaged and managed forests) (in Danish).
Finland
Biology. Zaenker and Fahldieck, 2022: 295-300 (during the investigation of the cave fauna in Hesse, specimens were found in two mine tunnels; these seem to represent real cave populations; therefore, the species can be classified as eutroglophilic from a biospeleological point of view; see paper for details) (in German with English abstract).
France
Habitat. Kramer, 2008c: 12 (collected at a limestone wooded stream-filled gorge, the stream can be torrential with moss covered boulders, and there are seepages from the steep sides of the limstone gorge).
Habitat. Reusch and Weber, 2013: 272 (found in caves, with references for France).
Biology. Quindroit, 2020a: 39, 45-48 (annotated list with information on abundancy and habitat of the Tipuloidea of the Pays de la Loire regoin, distr) (in French).
Habitat. Quindroit, 2020b: 138-139 (loc(s) Nord (59) and Pas-de-Calais (62) with notes on habitats and distr) (in French).
Habitat. Quindroit and Lemoine, 2022a: 24 (loc(s) Aisne (02) with notes on habitats and distr) (in French).
Habitat. Quindroit et al., 2023: 503 (locality on Corsica as in Kolcsar et al., 2021b but here with habitat information and photo: collected on a gravelly muddy seep in deciduous forest at 920 m (habitat photo)).
Germany
Biology. Warmke and Hering, 2000: 67-78 (investigation on the macroinvertebrate community inhabiting woody debris in low-order mountain streams in Hessen; L. remota was among the specimens studied but observations and conclusions are presented as Lipsothrix spec).
Habitat. Ruckert, 2005: 47, 53 (found in a study on the association with the type of source: limnokrene, rheokrene, or helokrene in Schlesig-Holstein; see paper for details, as well as Anhang A2-A10).
Habitat. Wittrock, 2005: 84 (summer emergence; rheokrene and limnokrene).
Habitat. Lehmann and Reusch, 2009: 122 (from around lowland springs).
Habitat. Reusch and Hohmann, 2009: 129 (from spring brooks).
Habitat. Reusch and Weber, 2013: 272 (found in caves, with references for Germany).
Habitat. Reiff et al., 2015: 234 (loc(s) Baden-Wurttemberg with list of habitats).
Great Britain
Habitat. Godfrey, 2000a: 7-16, 20, 23 (associated with semi-submerged wood in streams).
Biology. Godfrey, 2001a: 5-37 (habitat: associated with semi-submerged wood in streams; oviposition; biology).
Biology. Rotheray, 2001: 1, 13, 15 (in Scotland, the Malloch Society have reared L. remota from larvae found in wet, soft, decayed Fraxinus and Betula logs lying in seepages in woodlands; during this study the species was reared from Alder [Alnus], Elm [Ulmus], Birch [Betula], Oak [Quercus], Pine [Pinus] and other trees; see for biology also Rotheray, 2001 under L. ecucullata).
Habitat. Alexander, 2002: 90 (seepages and carr; larvae develop in wet dead wood and in wet ground by streams).
Biology. Hancock, 2002: 1-14 (report on the autecology in Scotland of Lipsothrix ecucullata and L. remota with as main findings: the preference for certain sizes of wood for larval development were identified; the timing of adult emergence was established; a methodology for surveying the species can be suggested; recommendations for habitat conservation are possible; see paper for details).
Habitat. Godfrey, 2003a: 24, 44 (larvae develop in semi-submerged decorticated wood in stream; review literature).
Habitat. Hewitt and Parker, 2006: 151-157 (immature stages in wet wood of Alnus, Fraxinus, Betula, Quercus and Fagus, lying in woodland seepages and streams).
Habitat. Boardman, 2007: 86 (woodland streams and seepages where saturated dead wood occurs).
Biology. Hancock et al., 2009: 79-80 (ubiquitous in wet wood lying in the water in stream margins; observations on behaviour of pupa in relation to immersion and emergence).
Habitat. Kramer, 2009f: 1 (in a wooded river valley swept from around the many natural dams formed by fallen branches).
Habitat. Cranston and Drake, 2010: 175 (Lipsothrix larvae can be found in marginal situations along flowing and standing waters [no species mentioned]).
Habitat. Drake, 2011a: 24 (among the Diptera from a wet Alder [Alnus] wood in Devon).
Habitat. Kramer, 2011j: 23 (a local but not uncommon species; the larvae feed in wet wood of tree branches in streams).
Habitat. Kramer, 2013i: 7 (associated with calcareous habitats).
Habitat. Kramer, 2016a: 1 (habitat photo from Gillfield Wood, see above).
Habitat. Brighton, 2017c: 29, 39 (overall regional checklist Lancashire and Cheshire (VC58, 59 and 60) with notes on habitats and/or regional occurrence and/or status; see paper for details).
Habitat. Showers, 2017c: 26 (collected in a long neglected Birch [Betula] woodland).
Habitat. Drake, 2022: 3 (collected in Devon: Knapp Copse, SY156953, 11 october 2021 in a small patch of possibly slightly acidic hillside seepage under sparse Sallow [Salix cinerea] woodland).
Habitat. Kramer and Morris, 2022b: 25 (info on the species, on the distr and map of loc(s) Leicestershire and Rutland (VC55); larvae feed on rotting wood in tree branches in streams).
Ireland
Habitat. Ashe et al., 2008: 11 (develops in wet dead wood in seepages and carr but also in wet ground along streams).
Habitat. Mitchell, 2025: 48 (loc(s) Co Sligo; from Union Wood, mainly comprising old oak woodland, see paper for details and habitat photo).
Lithuania
Habitat. Podeniene, 2003a: 1-295 (larva, descr, figs, habitat) (in Lithuanian).
Luxembourg
Habitat. Reusch and Weber, 2013: 272 (found in two caves, summary of biology).
Netherlands
Habitat. Cuppen, 2009: 7 (found locally in springs and upper reaches of brooks; larvae can be collected under bark of logs and thick branches that are lying in the water).
Poland
Habitat. Wiedenska, 2014: 12 (on the occurence of Limoniidae in six selected plant communities in the Kamienica Valley, Gorce National Park, Western Carpathians) (in Polish, habitat tables also in English).
Phenology. Wiedenska, 2015a: 52 (on the flight activity of Limoniidae in the Kamienica Valley, Gorce National Park, Western Carpathians) (in Polish, tables also in English).
Habitat. Wiedenska, 2017a: 20-21 (faunistic and habitat data of species from the Gorce NP) (in Polish).
Romania
Habitat. Ujvarosi, 2005a: 246 (springs, moist soil, citing various sources).
Habitat. Kolcsar et al., 2013: 72 (loc(s) Cluj-Napoca and surroundings; list of habitats with altitudes).
Sweden
Habitat. Fritz and Lindstrom, 2013: 13 (on distribution, habitat preference and habitat conservation in nature reserved forest and tree habitats in Hallands County) (in Schwedish).
on characters and taxonomy (mainly from 2000 onward):
Country not relevant
Key. Hancock et al., 2015a: 236 (adults Westpalaearctic species).
Characters. Olsen et al., 2018: 149 (Among the Lipsothrix species without a pterostigma, recognized by tip of femora not darkened, tip of tibiae darkened; fig).
Characters. Podenas et al., 2019b: 567 (comparison).
Great Britain
Immatures. Hancock et al., 2009: 79-87 (thoracic spiracular gill [pupal horn]: morphology, function, scanning electron figs, key).
Revision. Petersen, 2015: 277-278 (redescription, comparison, phylogeny, figs).
Characters. Kramer and Morris, 2022b: 25 (diagnosis).
Lithuania
Immatures. Podeniene, 2003a: 1-295 (larva, descr, figs, habitat) (in Lithuanian).
on distribution (mainly from 2000 onward):
Austria
Distribution. Vogtenhuber and Kofler, 2017: 140 (loc(s) Osttirol).
Bosnia-Herzegowina
Distribution. Oosterbroek and Simova-Tosic, 2004: 448 (review literature).
Bulgaria
Distribution. Hubenov, 2015: 221 (annotated list Diptera Pirin Mt, loc(s), faunistics and zoogeography).
Distribution. Hubenov, 2016: 49 (annotated checklist Diptera of the Rila Mountains based on data from the literature).
Distribution. Hubenov, 2017: 70 (on vertical distribution in the Pirin and Rila Mts; see paper for details on forests and subalpine-alpine zones, distr).
Distribution. Hubenov, 2021a: 35 (checklist with summary of distr in Bulgaria, references).
Distribution. Hubenov, 2025a: suppl: 3 (annotated list Diptera Stara Planina Mts, local distr, habitats and zoogeography).
Distribution. Hubenov, 2025b: suppl: 6 (annotated list Diptera high Bulgarian mts, local distr, habitats and zoogeography).
Croatia
First record. Kolcsar et al., 2015a: 34 (loc(s), distr).
Distribution. Kolcsar et al., 2023b: 25 (loc(s)).
Czech Rep.
Distribution. Stary, 2009i: webpage (checklist).
Distribution. Petersen, 2015: 277 (loc(s)).
Distribution. Stary and Vonicka, 2018: 67 (on Limoniidae and Pediciidae of two localities in northern Bohemia, with list of localities, altitudes and habitats).
Denmark
Distribution. Petersen and Jong, 2001b: 150 (checklist).
Estonia
Distribution. Stary, 2004f: (Fauna Europaea: added Estonia).
Finland
Distribution. Zaenker and Fahldieck, 2022: 295-300 (during the investigation of the cave fauna in Hesse, specimens were found in two mine tunnels; these seem to represent real cave populations; therefore, the species can be classified as eutroglophilic from a biospeleological point of view; see paper for details) (in German with English abstract).
France
Distribution. Kramer, 2008c: 12 (loc(s) Savoie (73)).
Distribution. Petersen, 2015: 277 (loc(s) B. Pyrenees (?)).
Distribution. Kramer and Langlois, 2019a: 76 (record(s) Ravin de Valbois, Doubs (25)).
Distribution. Kramer and Langlois, 2019b: 228 (list of craneflies captured 25-27 june 2019 in Ravin de Valbois, Doubs (25)).
Distribution. Quindroit, 2020a: 39, 45-48 (annotated list with information on abundancy and habitat of the Tipuloidea of the Pays de la Loire regoin, distr) (in French).
Distribution. Quindroit, 2020b: 138-139 (loc(s) Nord (59) and Pas-de-Calais (62) with notes on habitats and distr) (in French).
Distribution. Kolcsar et al., 2021b: 151 (loc(s) Corsica).
Distribution. Tissot et al., 2021: 209 (the Diptera of the Reserves Naturelles Nationales du Doubs (25), results of Malaise trapping since 2009) (in French).
Distribution. Quindroit and Lemoine, 2022a: 24 (loc(s) Aisne (02) with notes on habitats and distr) (in French).
Distribution. Quindroit and Lemoine, 2022a: 37 (table specifying the presence in the five departments of the Hauts-de-France region).
Distribution. Quindroit, 2022a: 25-28 (additions to the annotated list of the Tipuloidea of Pays de la Loire as given in Quindroit, 2020a).
Distribution. Lemoine and Quindroit, 2023: 9 (loc(s) Haute-Marne (52), Puy-de-Dome (63)).
Distribution. Quindroit and Racine, 2025: 360 (table specifying the presence in the five departments of the Pays de la Loire region).
Germany
Distribution. Schacht, 1999: 135 (checklist Bayern).
Distribution. Reusch and Oosterbroek, 2000: 162 (checklist German Bundeslander).
Distribution. Warmke and Hering, 2000: 67-78 (investigation on the macroinvertebrate community inhabiting woody debris in low-order mountain streams in Hessen; L. remota was among the specimens studied but observations and conclusions are presented as Lipsothrix spec).
Distribution. Reusch and Bellstedt, 2001b: 75 (checklist Thuringen).
Distribution. Ruckert, 2005: 47, 53 (found in a study on the association with the type of source: limnokrene, rheokrene, or helokrene in Schlesig-Holstein; see paper for details, as well as Anhang A2-A10).
Distribution. Schacht, 2005: 5 (checklist Bayern).
Distribution. Wittrock, 2005: 84 (loc(s) Schleswig-Holstein).
Distribution. Ruckert et al., 2007: 56 (loc(s) Schleswig-Holstein).
Distribution. Lehmann and Reusch, 2009: 122 (loc(s) Niedersachsen).
Distribution. Reusch and Hohmann, 2009: 129 (loc(s) Sachsen-Anhalt).
Distribution. Schacht, 2010: 17 (checklist Diptera Bayern).
Distribution. Petersen, 2015: 277 (loc(s) Baden-Wurttemberg, Bayern, Nordrhein-Westfalen, Rheinland-Pfalz).
Distribution. Reiff et al., 2015: 234 (loc(s) Baden-Wurttemberg with list of habitats).
Distribution. Stuke, 2019: 144 (checklist Niedersachsen and Bremen with reference to original sources).
Great Britain
Distribution. Godfrey, 2000a: 7-16, 20, 23 (see paper for details of sites surveyed in Shropshire and into Wales).
Distribution. Godfrey, 2001a: 5-37 (recorded from sites in Shropshire and just over the country border in Montgmeryshire; see paper for details).
Distribution. Rotheray, 2001: 1, 14 (loc(s) Scotland).
Distribution. Hancock, 2002: 4-5 (loc(s) Scotland).
Distribution. Godfrey, 2003a: 24, 44 (distr, review literature).
Distribution. Hewitt et al., 2005: 38 (loc(s) Cumbria).
Distribution. Hewitt and Parker, 2006: 151-157 (loc(s), distr, map).
Distribution. Boardman, 2007: 86 (distr, map).
Distribution. Kramer, 2009f: 1 (loc(s) Yorkshire).
Distribution. Skidmore, 2009: 129 (review of occurrence on the Western Isles of Scotland, distr).
Distribution. Drake, 2011a: 24 (loc(s) Devon).
Distribution. Kramer, 2011a: 3 (loc(s) Pembrokeshire).
Distribution. Kramer, 2011j: 23 (loc(s) Leicestershire and Rutland (VC55)).
Distribution. Kramer, 2013i: 8 (checklist of species from Leigh Woods NNR, Bristol district).
Distribution. Hewitt, 2014: (provisional checklist Cumbrian Diptera).
Distribution. Petersen, 2015: 277 (loc(s) Arran Is, Avon, Buckinghamshire, Denbighshire, Derbyshire, Devon, Hampshire, Isle of Wight, Peak, Pertshire, Powys, Staffordshire, Westmorland, Wiltshire, Yorkshire [written out here because the paper mentiones mostly localities]).
Distribution. Kramer, 2016a: 1 (loc(s) Sheffield).
Distribution. Brighton, 2017c: 29, 39 (overall regional checklist Lancashire and Cheshire (VC58, 59 and 60) with notes on habitats and/or regional occurrence and/or status; see paper for details).
Distribution. Showers, 2017c: 26 (loc(s) Northamptonshire).
Distribution. Cunningham, 2022: 28 (loc(s) Woodah Farm, Devon).
Distribution. Drake, 2022: 3 (collected in Devon: Knapp Copse, SY156953, 11 october 2021 in a small patch of possibly slightly acidic hillside seepage under sparse Sallow [Salix cinerea] woodland).
Distribution. Kramer and Morris, 2022b: 25 (info on the species, on the distr and map of loc(s) Leicestershire and Rutland (VC55); larvae feed on rotting wood in tree branches in streams).
Greece
First record. Petersen, 2015: 277 (loc(s) mainland and Euboea [Evia]).
Ireland
Distribution. Ashe et al., 2008: 11 (additional Irish record(s), distr).
Distribution. Chandler et al., 2008: 14 (checklist).
Distribution. Mitchell, 2025: 48 (loc(s) Co Sligo; from Union Wood, mainly comprising old oak woodland, see paper for details and habitat photo).
Italy
Distribution. Petersen, 2015: 277 (loc(s)).
Lithuania
Distribution. Pakalniskis et al., 2006: 22 (checklist).
Distribution. Petersen, 2015: 277 (loc(s)).
Luxembourg
First record. Reusch and Weber, 2013: 272-273 (loc(s) cantons Clervaux and Esch-sur-Alzette, map, distr).
Netherlands
Distribution. Jong and Oosterbroek, 2002b: 34 (checklist).
Distribution. Cuppen, 2009: 7 (loc(s) larval records).
Distribution. Hoven, 2024: 13 (loc(s) Limburg).
North Macedonia
Distribution. Oosterbroek and Simova-Tosic, 2004: 448 (review literature).
Distribution. Petersen, 2015: 277 (loc(s)).
Norway
First record. Olsen et al., 2018: 149, 162-163 (loc(s) TEI; distr; annotated checklist Nordic countries and for Norway according to the Strand regions).
Poland
Distribution. Wiedenska, 2007a: 75 (checklist).
Distribution. Wiedenska, 2010: 134 (loc(s) Parku Krajobrazowego Wzniesien Lodzkich, Lodz).
Distribution. Wiedenska, 2014: 12 (on the occurence of Limoniidae in six selected plant communities in the Kamienica Valley, Gorce National Park, Western Carpathians) (in Polish, habitat tables also in English).
Distribution. Wiedenska, 2017a: 20-21 (faunistic and habitat data of species from the Gorce NP) (in Polish).
Distribution. Wiedenska and Syratt, 2023: 144 (loc(s) Bieszczady Mountains, distr and remarks).
Romania
Distribution. Parvu, 2006: 261 (loc(s)).
Distribution. Ujvarosi, 2007: 228-231 (checklist Limoniidae).
Distribution. Keresztes, 2012: 31 (loc(s) Rimetea area [Alba County]).
Distribution. Kolcsar et al., 2013: 72 (loc(s) Cluj-Napoca and surroundings; list of habitats with altitudes).
Distribution. Petersen, 2015: 277 (loc(s)).
Slovakia
Distribution. Stary, 2009d: 37 (loc(s) Polana area).
Distribution. Stary, 2009i: webpage (checklist).
Distribution. Petersen, 2015: 277 (loc(s)).
Slovenia
Distribution. Oosterbroek and Simova-Tosic, 2004: 448 (review literature).
Distribution. Petersen, 2015: 277 (loc(s)).
Spain
Distribution. Eiroa and Baez, 2002a: 55 (checklist).
Distribution. Mederos and Eiroa, 2018: 27 (loc(s) Lerida, distr).
Sweden
Distribution. Salmela, 2011b: 117 (loc(s) Skane, distr).
Distribution. Fritz and Lindstrom, 2013: 13 (on distribution, habitat preference and habitat conservation in nature reserved forest and tree habitats in Hallands County) (in Schwedish).
Switzerland
Distribution. Podenas et al., 2006: 175, 284 (map, vertical distr per thermic level).
Distribution. Dufour and Merz, 2012: 355 (annotated checklist canton Geneva).
on flight period (mainly from 2000 onward):
Austria
Vogtenhuber and Kofler, 2017: month(s): 5.
Bulgaria
Hubenov, 2015: month(s): 6.
Hubenov, 2016: month(s): 6.
Croatia
Kolcsar et al., 2015a: month(s): 4-5.
Kolcsar et al., 2023b: month(s): 6, 9.
Czech Rep.
Petersen, 2015: month(s): 6-7.
Stary and Vonicka, 2018: month(s): 6-7.
France
Kramer, 2008c: month(s): 5.
Petersen, 2015: month(s): 9.
Kramer and Langlois, 2019b: month(s): 6.
Quindroit, 2020a: month(s): 5.
Quindroit, 2020b: month(s): 4-5.
Kolcsar et al., 2021b: month(s): 6.
Quindroit and Lemoine, 2022a: month(s): 6.
Lemoine and Quindroit, 2023: month(s): 5, 7.
Germany
Petersen, 2015: month(s): 5-8.
Reiff et al., 2015: month(s): 6-7.
Great Britain
Godfrey, 2000a: month(s): 6.
Godfrey, 2001a: month(s): 5.
Hancock, 2002: month(s): 5-8.
Kramer, 2009f: month(s): 5.
Kramer, 2011j: month(s): 5-7.
Petersen, 2015: month(s): 5-7.
Kramer, 2016a: month(s): 9.
Showers, 2017c: month(s): 5.
Cunningham, 2022: month(s): 6.
Kramer and Morris, 2022b: month(s): 5-7.
Greece
Petersen, 2015: month(s): 5-6.
Ireland
Ashe et al., 2008: month(s): 5-6.
Italy
Petersen, 2015: month(s): 8.
Lithuania
Petersen, 2015: month(s): 6.
Netherlands
Hoven, 2024: month(s): 5.
North Macedonia
Petersen, 2015: month(s): 8.
Norway
Olsen et al., 2018: month(s): 6-7.
Poland
Wiedenska, 2014: month(s): 6-7.
Wiedenska, 2015a: month(s): 6-7.
Wiedenska, 2017a: month(s): 6-8.
Wiedenska and Syratt, 2023: month(s): 5-8.
Romania
Parvu, 2006: month(s): 7.
Kolcsar et al., 2013: month(s): 6.
Petersen, 2015: month(s): 6.
Slovakia
Stary, 2009d: month(s): 6-7.
Petersen, 2015: month(s): 6.
Spain
Mederos and Eiroa, 2018: month(s): 6.
Switzerland
Podenas et al., 2006: month(s): 5-9.
on altitude (mainly from 2000 onward):
Austria
Vogtenhuber and Kofler, 2017: altitude: 1070 m.
Bulgaria
Hubenov, 2015: altitude: 1230-1350 m.
Hubenov, 2016: altitude: 1147 m.
Hubenov, 2017: altitude: 1147-1350 m.
Hubenov, 2021a: altitude: 1147-1350 m.
Hubenov, 2025a: altitude: 1147-1350 m.
Hubenov, 2025b: altitude: 1147-1350 m.
Croatia
Kolcsar et al., 2015a: altitude: 55 m.
Kolcsar et al., 2023b: altitude: 630-720 m.
Czech Rep.
Petersen, 2015: altitude: 900 m.
France
Kolcsar et al., 2021b: altitude: 920 m.
Germany
Reusch and Hohmann, 2009: altitude: 550 m.
Petersen, 2015: altitude: 1200 m.
Reiff et al., 2015: altitude: 666 m.
Great Britain
Hewitt and Parker, 2006: altitude: 10-350 m.
Greece
Petersen, 2015: altitude: 720-1700 m.
North Macedonia
Petersen, 2015: altitude: 427 m.
Poland
Wiedenska, 2017a: altitude: 800-100 m.
Wiedenska and Syratt, 2023: altitude: 500-950 m.
Romania
Petersen, 2015: altitude: 762 m.
Slovakia
Petersen, 2015: altitude: 914 m.
Spain
Mederos and Eiroa, 2018: altitude: 1650 m.

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