Details for Phylidorea (Phylidorea) squalens

Name: Phylidorea (Phylidorea) squalens (Zetterstedt, 1838) Publication: Insecta Lapp.: 834 (as Limnobia) Status: Recognized taxon, Synonym(s): denudata (Zetterstedt, 1838); tarda (Walker, 1848). Classification: Family Limoniidae Subfamily Limnophilinae Descriptive note: Monotypy: Stary and Oosterbroek, 2008. Keys: Krivosheina, M.G., 2010 (larvae 6 Phylidorea species); Peeters en Oosterbroek, 2014c (craneflies Netherlands, Belgium and Luxembourg) (in Dutch); Boardman, 2016 (craneflies Shropshire); Stubbs and Kramer, 2016g; Stubbs, 2021 (British craneflies) habitus male habitus male habitus male habitus female body part(s) antenna plus 15 more images of hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, wing, wing, wing, habitat, habitat, habitat, habitat, habitat and habitat. Distribution: Austria, Belarus, Belgium, Czech Rep., Denmark, Estonia, Finland, France, Germany, Great Britain, Ireland, Italy (Piemonte), Latvia, Lithuania, Netherlands, Norway, Poland, Romania, Slovakia, Sweden, Switzerland, Ukraine; Russia: RUN, RUW, RUC;; Russia: ES (south side Baikal lake); Kazakhstan (east); Mongolia. Note: see the manual for abbreviations: present-day Russia. Region(s): Westpalaearctic;; Eastpalaearctic General note: Added: Italy (Piemonte) (Stary and Oosterbroek, 1996). For Finland check Salmela, 2012b, 2012c, and citations below. For Norway check Olsen et al., 2018, and citations below. For Great Britain and Ireland check Stubbs, 2021, and citations below. Map: Ashe et al., 2005a (Ireland); Podenas et al., 2006 (Switzerland); Boardman, 2007 (Shropshire) Figure references: Habitus: Olsen and Andersen, 2021 (female); Quindroit, 2022a (male) Wing: Podenas et al., 2006; Boardman, 2007 Hypopygium: Stary, 1968b; Podenas et al., 2006 Immature stages Larva: Podeniene, 2003a Citations on biology (mainly from 2000 onward): Austria Habitat. Schultz et al., 2006: 351 (Hochmoor). Belarus Habitat. Paramonov and Sushko, 2010: 44-45 (loc(s), distr, about fauna of raised bogs, for details see paper) (in Russian). Habitat. Paramonov, 2019a: 96 (loc(s) Berezinsky Biosphere Reserve, distr, habitats, citing various authors) (in Russian). Czech Rep. Habitat. Stary and Vonicka, 2018: 52 (on Limoniidae and Pediciidae of two localities in northern Bohemia, with list of localities, altitudes and habitats). Finland Habitat. Salmela, 2004: 3, 4, 9 (species probably typical of open or semi-open mire types, very rarely recorded from springs). Habitat. Salmela and Ilmonen, 2005: 88-91 (a common species; the occurrence in relation to the trophic status of mires within an open boreal mire system is investigated; identified as indicator for mesotrophic sites). Habitat. Salmela and Autio, 2007b: 49 (common and widespread mire species). Biology. Salmela, 2008: 10, 20-26, 50 (49) (ecology or habitat: peatland; phenology:abundant later in the season, weeks 26-29). Habitat. Autio and Salmela, 2010: 48, 50 (habitats Aland Is, abundant open mire specialist to be found mostly from minerotrophc mires, being very abundant on oligotrophic and mesotrophic habitats, also found in wooded mires). Habitat. Salmela, 2011a: 1-28 (study on three wetland habitats: aapa mires, springs and headwater streams; among the most common species). Habitat. Salmela, 2012c: 13 (classified as a mire-dwelling species). Habitat. Autio et al., 2013: 1128, 1130 (species of fens and bogs, one of the species found in a study on species richness and rarity in a boreal mire). France Habitat. Quindroit, 2020b: 141 (loc(s) Nord (59) with notes on habitats and distr) (in French). Habitat. Quindroit, 2022a: 21, 27 (annotated list with information on abundancy and habitat of the Tipuloidea of the Pays de la Loire regoin, distr) (in French). Great Britain Habitat. Stubbs, 2003: 18 (given in Coulson, 1959 as confined to peat moorland, but from wider experience mainly a bog pool species or in other very wet boggy ground) (as Euphylidorea). Habitat. Boyce, 2004: 32, 61, 104 (a species of upland blanket mires; larvae are found in in bog pools). Habitat. Boardman, 2007: 72 (Sphagnum bogmoss, schwingmoor, boggy ditches on peat bogs, and wet boggy moorland). Habitat. Boardman, 2008: 8 (found at a number of wet flushes). Habitat. Kramer, 2008b: 7 (at boggy groud with Bog Asphodel). Habitat. Kramer, 2009f: 1 (Sphagnum bog). Habitat. Kramer, 2009g: 2 (along boggy margin of stream). Habitat. Cranston and Drake, 2010: 175 (Phylidorea larvae can be found in marginal situations along standing waters [no species mentioned]). Habitat. Stubbs, 2010a: 2 (normally in open bog habitat but can occur on bog woodlands). Habitat. Leftwich, 2011: 4 (heathland species). Habitat. Brighton, 2017c: 28, 37 (overall regional checklist Lancashire and Cheshire (VC58, 59 and 60) with notes on habitats and/or regional occurrence and/or status; see paper for details). Habitat. Stubbs, 2018b: 2 (an acid specialist, most characteristic of open habitats such as central raised moss but can occur in scub). Habitat. Boardman, 2020a: 1 (loc(s) Staffordshire, Sphagnum associate in area of valley mire). Habitat. Boardman, 2020b: 2 (on a visit to a bog close to Plas-y-brenin, Capel Curig, North Wales on 3 june 2019). Latvia Habitat. Spungis, 2008: 36 (raised bogs). Lithuania Habitat. Podeniene, 2002: 306 (larvae were found in riparian zones of small rivulet). Habitat. Podeniene, 2003a: 1-295 (larva, descr, figs, habitat) (in Lithuanian). Habitat. Podeniene, 2003b: 10, 17, 21-23, 29 (larval habitat: sand and silt in riparian zone of small rivulets, mud (in bogs of low lying woody swamps, on swampy riversides), fallings in swampy deciduous forests). Mongolia Habitat. Gelhaus and Podenas, 2006: 279, 290, 296-298 (habitat preference: 50% lotic, 50% Hovsgol lakeshore). Habitat. Yadamsuren et al., 2015: 475 (riparian zone, marsh). Norway Habitat. Olsen and Andersen, 2021: 212-213 (loc(s) Innlandet, in former Hedmark, with habitats specified separately, for habitats see also Jonassen and Andersen, 2020). Poland Habitat. Chaniecka and Wiedenska, 2006: 145 (occurrence in spring-, springbrook-, and/or bogspring plant communities) (in Polish). Habitat. Wiedenska, 2007a: 76 (tyrphobiont, species living inhabiting peatland). Habitat. Wiedenska, 2017a: 14 (faunistic and habitat data of species from the Gorce NP) (in Polish). Romania Habitat. Ujvarosi, 2005a: 247 (peat bog, brooks, small river, swamps, citing various sources). Habitat. Ujvarosi and Poti, 2006: 256 (habitat in Dupa Lunca marshland: predominantly at damp meadows). Habitat. Ujvarosi et al., 2011b: 108 (loc(s) Dupa Lanca marshy area near Voslobeni) (in Hungarian, species list with habitat information in English). Sweden Habitat. Lindstrom and Fritz, 2015: 44, 48 (distributin and ecology in five fens in Hallands County) (in Schwedish). on characters and taxonomy (mainly from 2000 onward): Country not relevant Immatures. Podeniene, 2004: 40 (comparison larva). Phylogeny. Ahonen, 2007: 1-28 (included in phylogenetic analysis of Tipuloidea subfamilies relationships based on molecular data). Taxonomy. Stary and Oosterbroek, 2008: 11 (most probably monotypic, subspecies elevated to species). Immatures. Krivosheina, M.G., 2010a: 19 (key to larvae of 6 Phylidorea species) (for key in English see Krivosheina, M.G., 2010b). Immatures. Podeniene and Gelhaus, 2010: 200-202 (larvae of the genus Phylidorea can be divided into the abdominalis, ferruginea and squalens groups based on the sclerotization of the spiracular field, the squalens group consists of longicornis and squalens). Lithuania Immatures. Podeniene, 2003a: 1-295 (larva, descr, figs, habitat) (in Lithuanian). Immatures. Podeniene, 2003b: 10, 17, 21-23, 29 (discriminating larval characters). on distribution (mainly from 2000 onward): Austria Distribution. Schultz et al., 2006: 351 (loc(s)). Distribution. Reusch and Heiss, 2012: 333-342 (loc(s) Nat. Park Gesause, see PDF for months and altitudes). Distribution. Vogtenhuber and Kofler, 2017: 141 (loc(s) Osttirol). Belarus First record. Paramonov and Sushko, 2010: 44-45 (loc(s), distr, about fauna of raised bogs, for details see paper) (in Russian). Distribution. Paramonov, 2019a: 96 (loc(s) Berezinsky Biosphere Reserve, distr, habitats, citing various authors) (in Russian). Czech Rep. Distribution. Stary, 2009i: webpage (checklist). Distribution. Stary and Vonicka, 2018: 52 (on Limoniidae and Pediciidae of two localities in northern Bohemia, with list of localities, altitudes and habitats). Denmark Distribution. Petersen and Jong, 2001b: 151 (checklist). Estonia Distribution. Stary, 2004f: (Fauna Europaea: added Estonia). Finland Distribution. Salmela, 2001a: 144, 150 (loc(s)). Distribution. Salmela, 2004: 3, 4, 9 (loc(s)). Distribution. Salmela and Ilmonen, 2005: 88-91 (loc(s)). Distribution. Salmela and Autio, 2007b: 54 (loc(s)). Distribution. Salmela et al., 2007b: 45 (loc(s)). Distribution. Salmela, 2008: 10, 50 (49) (loc(s), known from all Finnish ecoregions). Distribution. Autio and Salmela, 2010: 48 (loc(s) Aland Is). Distribution. Salmela, 2011a: 24 (loc(s) subalpine northern boreal Finland). Distribution. Salmela, 2012b: 239 (annotated list of Finnish crane flies). Distribution. Salmela, 2013: 18 (belongs to the most abundant and frequent cranefly species of Finland, see table above under Habitat). Distribution. Salmela and Petrasiunas, 2014: 28 (checklist Finnish Tipulomorpha). Distribution. Salmela et al., 2015: 106 (list of localities in extensive report on malaise-trapping of insects in conservation areas in Lapland in 2012–2014) (in Finnish). France Distribution. Quindroit, 2020b: 141 (loc(s) Nord (59) with notes on habitats and distr) (in French). Distribution. Labat, 2021: 92 (on Diptera taxa recorded along the over-all length of the Dordogne River and some of its tributaries; see paper for the locality details) (in French). Distribution. Quindroit and Lemoine, 2022a: 38 (table specifying the presence in the five departments of the Hauts-de-France region). Distribution. Quindroit, 2022a: 20 (loc(s) Orne (61) (subscription fig 5)). Distribution. Quindroit, 2022a: 21, 27 (annotated list with information on abundancy and habitat of the Tipuloidea of the Pays de la Loire regoin, distr) (in French). Distribution. Quindroit and Racine, 2025: 359 (table specifying the presence in the five departments of the Pays de la Loire region). Germany Distribution. Schacht, 1999: 134 (checklist Bayern). Distribution. Reusch and Bellstedt, 2000: 261 (loc(s) Thuringen). Distribution. Reusch and Oosterbroek, 2000: 156 (checklist German Bundeslander). Distribution. Reusch and Bellstedt, 2001b: 72 (checklist Thuringen). Distribution. Schacht, 2005: 6 (checklist Bayern). Distribution. Schacht, 2010: 18 (checklist Diptera Bayern). Distribution. Stuke, 2019: 146 (checklist Niedersachsen and Bremen with reference to original sources). Great Britain Distribution. Boyce, 2004: 32, 61, 104 (distr). Distribution. Boardman, 2007: 72 (loc(s), map). Distribution. Phillips, 2007: 150 (loc(s) Isle of Rum) (as Limnophila). Distribution. Anonymous, 2008b: 11 (loc(s) Shropshire: Preston Montford). Distribution. Boardman, 2008: 8 (loc(s) Shropshire). Distribution. Kramer, 2008b: 7 (loc(s) Wales). Distribution. Kramer, 2009f: 1 (loc(s) Yorkshire). Distribution. Kramer, 2009g: 2 (loc(s) Swansea). Distribution. Skidmore, 2009: 128 (review of occurrence on the Western Isles of Scotland, distr). Distribution. Leftwich, 2011: 4 (loc(s) Dorset). Distribution. Boardman, 2012a: 1 (loc(s) Shropshire). Distribution. Hewitt, 2014: (provisional checklist Cumbrian Diptera). Distribution. Boardman, 2017: 3 (loc(s) Dorset). Distribution. Brighton, 2017c: 28, 37 (overall regional checklist Lancashire and Cheshire (VC58, 59 and 60) with notes on habitats and/or regional occurrence and/or status; see paper for details). Distribution. Boardman, 2018: 3 (loc(s) Surrey). Distribution. Stubbs, 2018b: 2 (loc(s) Staffordshire). Distribution. Boardman, 2020a: 1 (loc(s) Staffordshire, Sphagnum associate in area of valley mire). Distribution. Boardman, 2020b: 2 (on a visit to a bog close to Plas-y-brenin, Capel Curig, North Wales on 3 june 2019). Ireland Distribution. Ashe et al., 2005a: 317, 333 (review Irish records). Distribution. Chandler et al., 2008: 12 (checklist). Kazakhstan First record. Devyatkov, 2020d: 271 (loc(s) East Kazakhstan, distr) (in Russian). Latvia Distribution. Spungis, 2008: 36 (loc(s)). Lithuania Distribution. Podeniene, 2002: 306 (loc(s)). Distribution. Pakalniskis et al., 2006: 23 (checklist). Mongolia First record. Gelhaus and Podenas, 2006: 279, 290, 296-298 (loc(s)). Distribution. Yadamsuren et al., 2015: 475 (loc(s) Mongolia). Netherlands Distribution. Jong and Oosterbroek, 2002b: 32 (checklist). Norway Distribution. Olsen et al., 2018: 160-161 (annotated checklist Nordic countries and for Norway according to the Strand regions). Distribution. Olsen and Andersen, 2021: 212-213 (loc(s) Innlandet, in former Hedmark, with habitats specified separately, for habitats see also Jonassen and Andersen, 2020). Poland Distribution. Chaniecka and Wiedenska, 2006: 145 (loc(s)). Distribution. Wiedenska, 2007a: 76 (checklist). Distribution. Wiedenska, 2017a: 14 (faunistic and habitat data of species from the Gorce NP) (in Polish). Romania Distribution. Ujvarosi and Poti, 2006: 256 (loc(s) eastern Carpathians). Distribution. Parvu, 2007: 234 (loc(s) Maramures Depression). Distribution. Ujvarosi, 2007: 228-231 (checklist Limoniidae). Distribution. Ujvarosi et al., 2011b: 108 (loc(s) Dupa Lanca marshy area near Voslobeni) (in Hungarian, species list with habitat information in English). Russia Distribution. Stary and Oosterbroek, 2008: 11 (loc(s) East Siberia, distr). Distribution. Humala and Polevoi, 2009: 66 (loc(s) RUN: Kareliya) (in Russian). Distribution. Gavryushin in litt., 2011e: (loc(s) RUN: Murmanskaya oblast). Distribution. Jakovlev et al., 2014: 309 (loc(s) RUN: Kareliya). Distribution. Paramonov and Pilipenko, 2016: 98 (loc(s) RUC: Tverskaya oblast, distr) (in Russian). Distribution. Pilipenko et al., 2020: 358 (loc(s) RUC: Mordoviya Rep., distr). Slovakia Distribution. Stary, 2009d: 26 (loc(s) Polana area). Distribution. Stary, 2009i: webpage (checklist). Sweden Distribution. Lindstrom and Fritz, 2015: 44, 48 (distributin and ecology in five fens in Hallands County) (in Schwedish). Switzerland Distribution. Podenas et al., 2006: 148, 323 (map, vertical distr per thermic level). on flight period (mainly from 2000 onward): Austria Vogtenhuber and Kofler, 2017: month(s): 7. Belarus Paramonov and Sushko, 2010: month(s): 6-9. Paramonov, 2019a: month(s): 6. Czech Rep. Stary and Vonicka, 2018: month(s): (4)5-7(8). Finland Salmela, 2001a: month(s): 6-7. Salmela, 2008: month(s): 6-8. France Quindroit, 2020b: month(s): 6. Quindroit, 2022a: month(s): 6. Quindroit, 2022a: month(s): 6. Germany Reusch and Bellstedt, 2000: month(s): 5, 7. Great Britain Phillips, 2007: month(s): 8. Kramer, 2008b: month(s): 7. Kramer, 2009f: month(s): 5. Kramer, 2009g: month(s): 7. Boardman, 2017: month(s): 6. Stubbs, 2018b: month(s): 6. Ireland Ashe et al., 2005a: month(s): 5-7. Kazakhstan Devyatkov, 2020d: month(s): 6, 8. Mongolia Gelhaus and Podenas, 2006: month(s): 7. Norway Olsen and Andersen, 2021: month(s): 6-7. Romania Parvu, 2007: month(s): 6. Ujvarosi et al., 2011b: month(s): 5-6. Russia Stary and Oosterbroek, 2008: month(s): 6. Gavryushin in litt., 2011e: month(s): 7. Paramonov and Pilipenko, 2016: month(s): 6. Pilipenko et al., 2020: month(s): 6. Slovakia Stary, 2009d: month(s): 5-7. Switzerland Podenas et al., 2006: month(s): 4-9. on altitude (mainly from 2000 onward): Austria Vogtenhuber and Kofler, 2017: altitude: 1650 m. France Labat, 2021: altitude: 1358 m. Kazakhstan Devyatkov, 2020d: altitude: 1654-2032 m. Mongolia Gelhaus and Podenas, 2006: altitude: 1645-1679 m. Romania Ujvarosi and Poti, 2006: altitude: 670 m. Slovakia Stary, 2009d: altitude: 850-870 m.