Details for Tipula (Pterelachisus) submarmorata
Name:Tipula (Pterelachisus) submarmorata Schummel, 1833
Publication:Beitr. Ent., Breslau 3: 36 (as Tipula)
Status:Recognized taxon, Synonym(s): meigeni Mannheims, 1966.
Classification:Family Tipulidae
Subfamily Tipulinae
Keys:Peeters en Oosterbroek, 2016a (craneflies Netherlands, Belgium and Luxembourg) (in Dutch); Boardman, 2016 (craneflies Shropshire); Stubbs and Kramer, 2016d; Stubbs, 2021 (British craneflies)
Tipula (Pterelachisus) submarmorata : habitus - maleTipula (Pterelachisus) submarmorata : habitus - maleTipula (Pterelachisus) submarmorata : habitus - maleTipula (Pterelachisus) submarmorata : habitus - maleTipula (Pterelachisus) submarmorata : habitus - male
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plus 23 more images of habitus, body part(s), body part(s), body part(s), body part(s), body part(s), hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, hypopygium, ovipositor, ovipositor, wing, wing, wing, habitat, habitat, habitat, habitat and habitat.
Distribution:Austria, Belarus, Belgium, Bosnia-Herzegovina, Bulgaria, Czech Rep., Denmark, Finland, France, Germany, Great Britain, Hungary, Ireland, Italy, Latvia, Liechtenstein, Lithuania, Luxembourg, Netherlands, Norway, Poland, Romania, Slovakia, Slovenia, Spain, Sweden, Switzerland, Ukraine; Russia: RUN, RUW, RUC.
Note: see the manual for abbreviations: present-day Russia.
Region(s):Westpalaearctic
General note:For Finland check Salmela, 2012b, 2012c, and citations below. For Norway check Hofsvang et al., 2019, and citations below. For Great Britain and Ireland check Stubbs, 2021, and citations below.
Map:Ashe et al., 2007a (Ireland); Boardman, 2007 (Shropshire)
Figure
references:
Ovipositor: Vogtenhuber, 2004b

Immature stages
Larva: Podeniene, 2003a

Miscellaneous
Quindroit, 2023 (habitat); Heiss and Malec, 2023 (habitat)
Citationson biology (mainly from 2000 onward):
Belgium
Habitat. Tack et al., 2021: 126 (loc(s) Antwerpen, alle taxa biodiversiteit inventarisatie Bos t Ename, combinaties van nat en vochtig bos (in Dutch), see paper for description).
Denmark
Habitat. Nielsen and Nielsen, 2009: 124 (emerging from a forest floor in a Beech [Fagus] stand)(in Danish).
Habitat. Byriel and Rojas, 2017: 118 (among the 101 species in a study on cranefly hotspots in unmanaged and managed forests) (in Danish).
Finland
Habitat. Salmela and Vartija, 2007: 10 (occurs in moist forests and around brooks and lake shores, citing Salmela, 2006).
Habitat. Salmela, 2009b: 249, 257 (main habitat: riparian forests; the paper includes further habitat details).
France
Habitat. Dufour, 2003b: 89 (loc(s) with information on habitat(s)) (in French).
Habitat. Tillier and Dehalleux, 2019: 164 (loc(s) with habitat info, summary habitat).
Habitat. Tillier and Oosterbroek, 2019: 235 (loc(s) and distr Mercantour; habitat: collected in various forest types; see paper for details).
Habitat. Quindroit, 2020b: 129 (loc(s) Nord (59) with notes on habitats and distr) (in French).
Habitat. Quindroit, 2023: 62 (loc(s) Hautes-Alpes (05)) [locality and habitat are not given in the text but with the habitat figure].
Germany
Habitat. Heiss and Merkel-Wallner, 2013: 21-24 (loc(s) Bayern, table with Malaise trap habitats and species; see paper for details).
Biology. Heiss and Flugel, 2015: 271, 278 (loc(s) Hessen; review local distribution and biology, citing various authors
Habitat. Heiss et al., 2017: 180-182 (overview of collecting methods and habitats).
Habitat. Heiss, 2017a: 563, 568, 572 (loc(s) Dinkelberg, Baden-Wurttemberg and table with details on presence in the various biotop-groups).
Habitat. Dunk, 2018: 79, 96 (loc(s) Bayern with list of habtats).
Great Britain
Habitat. Godfrey, 2001a: 21, 23 (also as meigeni).
Habitat. Boardman, 2007: 25 (a cranefly of broad-leaved woodlands).
Habitat. Kramer, 2011j: 10, 25 (occasional spring species of woodland).
Habitat. Kramer and Morris, 2021b: 25 (info on the species, on the distr and map of loc(s) Leicestershire and Rutland (VC55); in woodlands during spring with the larvae in the humus of rich woodland soils).
Latvia
Habitat. Salmela and Vartija, 2007: 10 (lakeshore with meso-mesoeutrophic mire vegetation).
Lithuania
Habitat. Podeniene, 2003a: 1-295 (larva, descr, figs, habitat) (in Lithuanian).
Habitat. Podeniene, 2003b: 12, 28 (larval habitat: uppermost layer of soil in forests and in fallings).
Luxembourg
Habitat. Vogtenhuber, 2007: 353 (Larve wurde nur in Waldern unter Moos, sowohl in der Streuschicht als auch in stark vermodernten Holz gefunden).
Poland
Biology. Szujecki, 1987: 390 (one of a series of forest saprophages that play an important role in soil-forming processes).
on characters and taxonomy (mainly from 2000 onward):
Country not relevant
Characters. Vogtenhuber, 2004b: 349 (comparison, figs).
Phylogeny. Pilipenko et al., 2012a: 54 (barcoding, phylogeny).
Characters. Oosterbroek et al., 2013: 45-46 (comparison).
Phylogeny. Men et al., 2017a: 67, 74 (barcoding, phylogeny).
Immatures. Cai et al., 2023: 8-11 (larval comparison).
Lithuania
Immatures. Podeniene, 2003a: 1-295 (larva, descr, figs, habitat) (in Lithuanian).
on distribution (mainly from 2000 onward):
Austria
Distribution. Vogtenhuber, 2011: 50 (checklist Austria, list of provinces).
Distribution. Reusch and Heiss, 2012: 343-351 (loc(s) Nat. Park Gesause, see PDF for months and altitudes).
Distribution. Aistleitner, 2015: 96 (loc(s) Vorarlberg).
Distribution. Vogtenhuber and Kofler, 2017: 128 (loc(s) Osttirol, distr in Austria).
Belarus
Distribution. Palaczyk, 2001: 270 (loc(s)).
Belgium
Distribution. Tack et al., 2021: 126 (loc(s) Antwerpen, alle taxa biodiversiteit inventarisatie Bos t Ename, combinaties van nat en vochtig bos (in Dutch), see paper for description).
Bosnia-Herzegowina
Distribution. Simova-Tosic and Oosterbroek, 2003: 53 (review literature).
Bulgaria
First record. Kolcsar et al., 2023a: 22 (loc(s) Vitosha Mts).
Czech Rep.
Distribution. Stary, 2009k: webpage (checklist).
Denmark
Distribution. Petersen et al., 2001b: 163 (checklist).
Distribution. Nielsen and Nielsen, 2009: 124 (loc(s)).
Distribution. Salmela, 2012d: 122 (all Danish 89 specimens of hortulana belong to submarmorata; see paper for details).
Finland
Distribution. Polevoi, 2001: 191 (loc(s)) (as meigeni).
Distribution. Salmela, 2009b: 249, 253, 257 (common in the southern boreal zone, much less so towards the north and rare or absent in most of northern boreal Finland, red-list status, distr, map).
Distribution. Pilipenko et al., 2012a: 54 (loc(s)).
Distribution. Salmela, 2012b: 241 (annotated list of Finnish crane flies).
Distribution. Salmela and Petrasiunas, 2014: 34 (checklist Finnish Tipulomorpha).
France
Distribution. Dufour, 2003b: 89 (loc(s) with information on habitat(s)) (in French).
Distribution. Kramer and Langlois, 2019a: 76 (record(s) Ravin de Valbois, Doubs (25)).
Distribution. Tillier and Dehalleux, 2019: 164 (first records for Ile-de-France).
Distribution. Tillier and Oosterbroek, 2019: 235 (loc(s) and distr Mercantour; habitat: collected in various forest types; see paper for details).
Distribution. Quindroit, 2020b: 129 (loc(s) Nord (59) with notes on habitats and distr) (in French).
Distribution. Tissot et al., 2021: 217 (the Diptera of the Reserves Naturelles Nationales du Doubs (25), results of Malaise trapping since 2009) (in French).
Distribution. Quindroit, 2023: 62 (loc(s) Hautes-Alpes (05)) [locality and habitat are not given in the text but with the habitat figure].
Germany
Distribution. Schacht et al., 2001: 427 (loc(s) Bayern).
Distribution. Heiss, 2003: 14 (checklist Thuringen).
Distribution. Schacht, 2005: 4 (checklist Bayern).
Distribution. Hable et al., 2010b: 90 (loc(s) Bayern).
Distribution. Schacht, 2010: 38 (checklist Diptera Bayern).
Distribution. Merkel-Wallner et al., 2011: 214 (loc(s) Bayern).
Distribution. Heiss and Merkel-Wallner, 2013: 21-24 (loc(s) Bayern, table with Malaise trap habitats and species; see paper for details).
Distribution. Dunk and Kraus, 2014: 168 (loc(s) Bayern).
Distribution. Merkel-Wallner, 2014: 49 (loc(s) Bayern).
Distribution. Heiss and Flugel, 2015: 271, 278 (loc(s) Hessen; review local distribution and biology, citing various authors
Distribution. Heiss et al., 2017: 168 (loc(s) Thuringen).
Distribution. Heiss, 2017a: 563, 568, 572 (loc(s) Dinkelberg, Baden-Wurttemberg and table with details on presence in the various biotop-groups).
Distribution. Heiss, 2017b: 23 (checklist Tipulidae Thuringen).
Distribution. Dunk, 2018: 79, 96 (loc(s) Bayern with list of habtats).
Distribution. Heiss et al., 2019: 252 (loc(s); annotated checklist of the Tipulidae of Niedersachsen and Bremen, including the presence in the surrounding Bundeslandern and the Netherlands) (in German).
Distribution. Heiss, 2019: 37, 44 (loc(s) and checklist Mecklenburg-Vorpommern, see paper for further details).
Distribution. Stuke, 2019: 241 (checklist Niedersachsen and Bremen with reference to original sources).
Distribution. Heiss et al., 2021: 241 (found in caves in Hessen and Bayern, see paper on details of which part of the caves) (in German).
Distribution. Heiss and Malec, 2023: 318 (loc(s) Hessen).
Distribution. Heiss and Malec, 2023: 329 (annotated checklist Hessen).
Great Britain
Distribution. Godfrey, 2001a: 21, 23 (loc(s)) (also as meigeni).
Distribution. Merrifield and Merrifield, 2004: 171 (loc(s) Cawdor Woods (NH8449), Inverness, 31.v.1982).
Distribution. Boardman, 2007: 25 (loc(s), map).
Distribution. Skidmore, 2009: 124 (review of occurrence on the Western Isles of Scotland, distr).
Distribution. Kramer, 2011j: 10 (loc(s) Leicestershire and Rutland (VC55)).
Distribution. Kramer, 2013f: 1 (loc(s) Northamptonshire).
Distribution. Kramer, 2013i: 8 (checklist of species from Leigh Woods NNR, Bristol district).
Distribution. Hewitt, 2014: (provisional checklist Cumbrian Diptera).
Distribution. Showers, 2016: 2 (loc(s) Northamptonshire).
Distribution. Brighton, 2017b: 30 (overall regional checklist Lancashire and Cheshire (VC58, 59 and 60) with notes on habitats and/or regional occurrence and/or status; see paper for details).
Distribution. Kramer and Morris, 2021b: 25 (info on the species, on the distr and map of loc(s) Leicestershire and Rutland (VC55); in woodlands during spring with the larvae in the humus of rich woodland soils).
Ireland
Distribution. Ashe et al., 2007a: 302, 323-324, 353 (review Irish records, map).
Distribution. Chandler et al., 2008: 6 (checklist).
Italy
Distribution. Oosterbroek, 2008c: 339, 341 (loc(s), annotated checklist South Tyrol [Prov. Bolzano]).
Distribution. Terzani et al., 2012: 306 (known from Riserva naturale Camaldoli (Arezzo), citing Oosterbroek, 2005).
Latvia
First record. Salmela and Vartija, 2007: 10 (loc(s)).
Liechtenstein
First record. Aistleitner, 2015: 97 (loc(s)).
Lithuania
Distribution. Pakalniskis et al., 2006: 28 (checklist).
Luxembourg
Distribution. Vogtenhuber, 2007: 351, 353 (loc(s), distr).
Netherlands
Distribution. de Jong and Oosterbroek, 2002d: 41 (checklist).
Distribution. Oosterbroek et al., 2013: 43 (number of Dutch records, distr).
Distribution. Oosterbroek and de Jong, 2020: 46 (Tipulidae Kaaistoep, Noord-Brabant).
Norway
Distribution. Hofsvang, 2010: 13 (loc(s)).
Distribution. Hofsvang et al., 2019: 117-118, 142-143 (review Norwegian records, annotated checklist Nordic countries and for Norway according to the Strand regions).
Poland
Distribution. Skibinska and Chudzicke, 2007b: 78 (checklist).
Romania
Distribution. Ujvarosi, 2007: 232-233 (checklist Tipulidae).
Distribution. Keresztes, 2012: 31 (loc(s) Rimetea area [Alba County]).
Distribution. Torok et al., 2014: 19 (checklist Romania).
Russia
Distribution. Pilipenko, 2009a: 215 (annotated checklist Tipulidae Central European Territory, loc(s) RUC: Tverskaya oblast, Moskovskaya oblast, distr).
Distribution. Ruchin and Pilipenko, 2015: 59 (loc(s), checklist RUC: Mordoviya Rep.).
Distribution. Paramonov and Pilipenko, 2016: 101 (loc(s) RUC: Tverskaya oblast) (in Russian).
Distribution. Pilipenko et al., 2020: 365-366 (loc(s) RUC: Mordoviya Rep., distr).
Slovakia
Distribution. Stary, 2009k: webpage (checklist).
Slovenia
Distribution. Simova-Tosic and Oosterbroek, 2003: 53 (review literature).
Spain
Distribution. Eiroa and Baez, 2002b: 80 (checklist).
Sweden
Distribution. Salmela, 2012d: 122 (Swedish specimens of pseudovariipennis belong to submarmorata).
Distribution. Fritz and Lindstrom, 2013: 63 (loc(s) Hallands County) (in Schwedish).
Switzerland
Distribution. Freitag, 2009: 176 (loc(s) Vaud).
on flight period (mainly from 2000 onward):
Austria
Aistleitner, 2015: month(s): 5-6.
Vogtenhuber and Kofler, 2017: month(s): 5-6.
Bulgaria
Kolcsar et al., 2023a: month(s): 6.
France
Dufour, 2003b: month(s): 6.
Tillier and Dehalleux, 2019: month(s): 4-5.
Tillier and Oosterbroek, 2019: month(s): 5-7.
Quindroit, 2020b: month(s): 4.
Quindroit, 2023: month(s): 6.
Germany
Schacht et al., 2001: month(s): 4-6.
Heiss and Merkel-Wallner, 2013: month(s): 4-6.
Heiss et al., 2017: month(s): 5-7.
Heiss, 2017a: month(s): 4-5.
Dunk, 2018: month(s): 6-7.
Heiss, 2019: month(s): 5.
Heiss et al., 2021: month(s): 5.
Heiss and Malec, 2023: month(s): 4-5.
Great Britain
Godfrey, 2001a: month(s): 5.
Merrifield and Merrifield, 2004: month(s): 5.
Boardman, 2007: month(s): 4-6.
Kramer, 2011j: month(s): 5.
Kramer, 2013f: month(s): 5.
Kramer and Morris, 2021b: month(s): 4-5.
Ireland
Ashe et al., 2007a: month(s): 5, 7.
Italy
Oosterbroek, 2008c: month(s): 6.
Latvia
Salmela and Vartija, 2007: month(s): 5.
Liechtenstein
Aistleitner, 2015: month(s): 5-6.
Norway
Hofsvang et al., 2019: month(s): 5-7.
Russia
Pilipenko, 2009a: month(s): 5-6.
Ruchin and Pilipenko, 2015: month(s): 5.
Pilipenko et al., 2020: month(s): 5-6.
on altitude (mainly from 2000 onward):
Austria
Aistleitner, 2015: altitude: 425 m.
Vogtenhuber and Kofler, 2017: altitude: 950-1940 m.
France
Dufour, 2003b: altitude: 1200-1780 m.
Tillier and Oosterbroek, 2019: altitude: 1290-1850 m.
Quindroit, 2023: altitude: 1900 m.
Germany
Merkel-Wallner et al., 2011: altitude: <700-1399 m.
Heiss et al., 2021: altitude: 332-440 m.
Italy
Oosterbroek, 2008c: altitude: 1630 m.
Liechtenstein
Aistleitner, 2015: altitude: 480-1350 m.

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